| Wanger Acorn | First Quarter Report 2022 | 1 |
| 2 | Wanger Acorn | First Quarter Report 2022 |
| Money Market Funds 1.4% | ||
| Shares | Value ($) | |
| Columbia Short-Term Cash Fund, 0.308%(f),(h) | 7,542,006 | 7,538,989 |
| Total Money Market Funds (Cost $7,538,932) | 7,538,989 | |
| Total Investments in Securities (Cost: $535,755,364) | 566,726,875 | |
| Obligation to Return Collateral for Securities Loaned | (5,873,600) | |
| Other Assets & Liabilities, Net | 1,030,970 | |
| Net Assets | 561,884,245 | |
| (a) | Non-income producing investment. |
| (b) | All or a portion of this security was on loan at March 31, 2022. The total market value of securities on loan at March 31, 2022 was $5,741,444. |
| (c) | Represents fair value as determined in good faith under procedures approved by the Board of Trustees. At March 31, 2022, the total value of these securities amounted to $1,351,201, which represents 0.24% of total net assets. |
| (d) | Denotes a restricted security, which is subject to legal or contractual restrictions on resale under federal securities laws. Disposal of a restricted investment may involve time-consuming negotiations and expenses, and prompt sale at an acceptable price may be difficult to achieve. Private placement securities are generally considered to be restricted, although certain of those securities may be traded between qualified institutional investors under the provisions of Section 4(a)(2) and Rule 144A. The Fund will not incur any registration costs upon such a trade. These securities are valued at fair value determined in good faith under consistently applied procedures established by the Fund’s Board of Trustees. At March 31, 2022, the total market value of these securities amounted to $1,351,201, which represents 0.24% of total net assets. Additional information on these securities is as follows: |
| Security | Acquisition Dates | Shares | Cost ($) | Value ($) |
| DNA Script | 10/01/2021 | 2,550 | 2,223,535 | 1,351,201 |
| (e) | Valuation based on significant unobservable inputs. |
| (f) | The rate shown is the seven-day current annualized yield at March 31, 2022. |
| (g) | Investment made with cash collateral received from securities lending activity. |
| (h) | As defined in the Investment Company Act of 1940, as amended, an affiliated company is one in which the Fund owns 5% or more of the company’s outstanding voting securities, or a company which is under common ownership or control with the Fund. The value of the holdings and transactions in these affiliated companies during the period ended March 31, 2022 are as follows: |
| Affiliated issuers | Beginning of period($) | Purchases($) | Sales($) | Net change in unrealized appreciation (depreciation)($) | End of period($) | Realized gain (loss)($) | Dividends($) | End of period shares |
| Columbia Short-Term Cash Fund, 0.308% | ||||||||
| 28,253,542 | 79,387,890 | (100,103,747) | 1,304 | 7,538,989 | (3,095) | 6,336 | 7,542,006 | |
| ADR | American Depositary Receipt |
| Wanger Acorn | First Quarter Report 2022 | 3 |