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Related Party Transactions (Details Textual) - USD ($)
$ / shares in Units, $ in Thousands
12 Months Ended
Dec. 31, 2012
Dec. 31, 2015
Aug. 07, 2015
Dec. 31, 2014
Aug. 31, 2012
Nov. 30, 2011
Related Party Transaction [Line Items]            
Common Stock, Shares, Issued   90,908,234   72,594,626    
Common Stock, Value, Issued   $ 91   $ 72    
Common Stock, Par or Stated Value Per Share   $ 0.001   $ 0.001    
Intrexon [Member]            
Related Party Transaction [Line Items]            
Common Stock, Shares, Issued     937,500   3,552,210 3,123,558
Common Stock, Value, Issued     $ 3,000   $ 7,800 $ 1,700
Common Stock, Par or Stated Value Per Share         $ 2.20 $ 0.54
Costs and Expenses, Related Party $ 2,900          
Intrexon [Member] | Research and Development Expense [Member]            
Related Party Transaction [Line Items]            
Costs and Expenses, Related Party $ 2,500