Acquisitions - Assets Acquired and Liabilities Assumed (Details) - USD ($)	Sep. 30, 2021	Dec. 31, 2020	Oct. 01, 2020
Business Acquisition [Line Items]
Goodwill	$ 190,183,000	$ 189,662,000
Pfenex
Business Acquisition [Line Items]
Cash			$ 51,407,000
Restricted cash			200,000
Accounts and unbilled receivables			1,359,000
Property and equipment, net			7,823,000
Right-of-use asset			3,070,000
Other assets			1,338,000
Intangibles acquired			385,000,000
Goodwill			91,271,000
Accounts payable			(6,814,000)
Accrued liabilities			(8,466,000)
Deferred revenue			(3,908,000)
Lease liabilities			(3,070,000)
Other liabilities			(1,382,000)
Deferred tax liabilities, net			(52,730,000)
Total consideration			465,098,000
Goodwill acquired that is expected to be deductible for tax purposes			$ 0