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Collaboration, License, and Other Agreements - Intellia Narrative (Details) - USD ($)
$ in Millions
3 Months Ended 6 Months Ended
Jun. 30, 2021
Jun. 30, 2020
Jun. 30, 2021
Jun. 30, 2020
May 31, 2020
Research Collaboration and Licensing Arrangements [Line Items]          
Research and development $ 714.2 $ 722.0 $ 1,457.1 $ 1,305.9  
Intellia          
Research Collaboration and Licensing Arrangements [Line Items]          
Upfront payment made for collaboration agreement   70.0      
Research and development   $ 15.0      
Intellia | Common Stock          
Research Collaboration and Licensing Arrangements [Line Items]          
Shares purchased (in shares)         925,218
Shares purchase price         $ 30.0