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Basis of Presentation and Significant Accounting Policies - Segment Information (Details) - segment
3 Months Ended 9 Months Ended
Sep. 30, 2021
Sep. 30, 2020
Sep. 30, 2021
Sep. 30, 2020
Segment Information        
Number of operating segments 1 1 1 1
Other customers | Revenue | Customer concentration        
Segment Information        
Percentages of revenue recognized 0.00% 0.00% 0.00% 0.00%
Viridian | Revenue | Customer concentration        
Segment Information        
Percentages of revenue recognized 28.00%      
Royalty revenue | Roche | Revenue | Customer concentration        
Segment Information        
Percentages of revenue recognized 71.00% 99.00% 93.00% 97.00%