SEC FORM 4

SEC Form 4

FORM 4

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940

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Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

1. Name and Address of Reporting Person^*

WHITE TONY L

(Last)

(First)

(Middle)

APPLERA CORPORATION

301 MERRITT 7

(Street)

NORWALK

06851-1070

(City)

(State)

(Zip)

2. Issuer Name and Ticker or Trading Symbol
APPLERA CORP [ ABI/CRA ]

5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

X	Director	10% Owner
X	Officer (give title below)	Other (specify below)
	Chairman, President and CEO

3. Date of Earliest Transaction (Month/Day/Year)
02/04/2008

4. If Amendment, Date of Original Filed (Month/Day/Year)

6. Individual or Joint/Group Filing (Check Applicable Line)

X	Form filed by One Reporting Person
	Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	3. Transaction Code (Instr. 8)		4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5)			5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4)	6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	Code	V	Amount	(A) or (D)	Price		6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
Celera Group Common Stock	02/04/2008		S⁽¹⁾		438	D	$15.29	99,124.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		36	D	$15.28	99,088.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		15	D	$15.27	99,073.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		313	D	$15.26	98,760.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		36	D	$15.24	98,724.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		53	D	$15.22	98,671.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		458	D	$15.21	98,213.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		182	D	$15.2	98,031.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		340	D	$15.19	97,691.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		535	D	$15.18	97,156.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		109	D	$15.17	97,047.0998	D
Celera Group Common Stock	02/04/2008		S⁽¹⁾		37	D	$15.16	97,010.0998	D
Celera Group Common Stock								25,000	I	By Elizabeth Ann White Business Interests, LLLP⁽²⁾
Celera Group Common Stock								9,900	I	By Tony Lee White 2006 Two Year Grantor Retained Annuity Trust⁽³⁾
Celera Group Common Stock								89,000	I	By Tony Lee White 2007 Grantor Retained Annuity Trust⁽⁴⁾

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	4. Transaction Code (Instr. 8)		5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5)		6. Date Exercisable and Expiration Date (Month/Day/Year)		7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4)		8. Price of Derivative Security (Instr. 5)	9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)	10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	Code	V	(A)	(D)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares	8. Price of Derivative Security (Instr. 5)		10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)

Explanation of Responses:

1. Reflects sale by independent third-party administrator under the issuer's Insider Diversification Program.

2. Elizabeth Ann White Business Interests, LLLP, is a family limited partnership (the "FLP"). The reporting person's adult daughter is the sole general partner of the FLP, and the reporting person's wife and a grantor retained annuity trust established by the reporting person are currently the sole limited partners of the FLP.

3. The reporting person's wife is the sole trustee of the Tony Lee White 2006 Two Year Grantor Retained Annuity Trust.

4. The reporting person's wife is the sole trustee of the Tony Lee White 2007 Grantor Retained Annuity Trust.

Remarks:

This is the third of three forms being filed by the reporting person on February 6, 2008.

	/s/ Thomas P. Livingston, Attorney-in-Fact for Tony L. White	02/06/2008
	** Signature of Reporting Person	Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.