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<SEC-DOCUMENT>0000077551-07-000124.txt : 20070504
<SEC-HEADER>0000077551-07-000124.hdr.sgml : 20070504
<ACCEPTANCE-DATETIME>20070504135014
ACCESSION NUMBER:		0000077551-07-000124
CONFORMED SUBMISSION TYPE:	4
PUBLIC DOCUMENT COUNT:		1
CONFORMED PERIOD OF REPORT:	20070502
FILED AS OF DATE:		20070504
DATE AS OF CHANGE:		20070504

REPORTING-OWNER:	

	OWNER DATA:	
		COMPANY CONFORMED NAME:			AYERS RICHARD H
		CENTRAL INDEX KEY:			0001200844

	FILING VALUES:
		FORM TYPE:		4
		SEC ACT:		1934 Act
		SEC FILE NUMBER:	001-04389
		FILM NUMBER:		07819459

	BUSINESS ADDRESS:	
		STREET 1:		C/O APPLERA CORP
		STREET 2:		301 MERRITT 7
		CITY:			NORWALK
		STATE:			CT
		ZIP:			06856-5435
		BUSINESS PHONE:		2038402000

ISSUER:		

	COMPANY DATA:	
		COMPANY CONFORMED NAME:			APPLERA CORP
		CENTRAL INDEX KEY:			0000077551
		STANDARD INDUSTRIAL CLASSIFICATION:	LABORATORY ANALYTICAL INSTRUMENTS [3826]
		IRS NUMBER:				061534213
		STATE OF INCORPORATION:			DE
		FISCAL YEAR END:			0630

	BUSINESS ADDRESS:	
		STREET 1:		301 MERRITT 7
		CITY:			NORWALK
		STATE:			CT
		ZIP:			06851
		BUSINESS PHONE:		2038402000

	MAIL ADDRESS:	
		STREET 1:		301 MERRITT 7
		CITY:			NORWALK
		STATE:			CT
		ZIP:			06851

	FORMER COMPANY:	
		FORMER CONFORMED NAME:	PE CORP
		DATE OF NAME CHANGE:	19990129

	FORMER COMPANY:	
		FORMER CONFORMED NAME:	PERKIN ELMER CORP
		DATE OF NAME CHANGE:	19930601
</SEC-HEADER>
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        <nonDerivativeTransaction>
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                <transactionShares>
                    <value>6</value>
                </transactionShares>
                <transactionPricePerShare>
                    <value>30.75</value>
                </transactionPricePerShare>
                <transactionAcquiredDisposedCode>
                    <value>D</value>
                </transactionAcquiredDisposedCode>
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            <postTransactionAmounts>
                <sharesOwnedFollowingTransaction>
                    <value>6875</value>
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                <directOrIndirectOwnership>
                    <value>D</value>
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                <sharesOwnedFollowingTransaction>
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            <ownershipNature>
                <directOrIndirectOwnership>
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                <natureOfOwnership>
                    <value>By Suzanne L. Ayers Living Trust</value>
                    <footnoteId id="F2"/>
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            <transactionAmounts>
                <transactionShares>
                    <value>150</value>
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                <transactionPricePerShare>
                    <value>6.5525</value>
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                    <value>A</value>
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                <sharesOwnedFollowingTransaction>
                    <value>2865</value>
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                    <value>9</value>
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                <transactionPricePerShare>
                    <value>14.37</value>
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                <transactionAcquiredDisposedCode>
                    <value>D</value>
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                <sharesOwnedFollowingTransaction>
                    <value>2856</value>
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        <nonDerivativeTransaction>
            <securityTitle>
                <value>Celera Group Common Stock</value>
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                <directOrIndirectOwnership>
                    <value>D</value>
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                    <value></value>
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        </nonDerivativeTransaction>
    </nonDerivativeTable>

    <derivativeTable>
        <derivativeTransaction>
            <securityTitle>
                <value>Applied Biosystems Group Director Stock Options-Right to Buy</value>
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            <conversionOrExercisePrice>
                <value>16.0728</value>
            </conversionOrExercisePrice>
            <transactionDate>
                <value>2007-05-02</value>
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            <transactionCoding>
                <transactionFormType>4</transactionFormType>
                <transactionCode>M</transactionCode>
                <equitySwapInvolved>0</equitySwapInvolved>
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            <transactionAmounts>
                <transactionShares>
                    <value>600</value>
                </transactionShares>
                <transactionPricePerShare>
                    <value>0</value>
                </transactionPricePerShare>
                <transactionAcquiredDisposedCode>
                    <value>D</value>
                </transactionAcquiredDisposedCode>
            </transactionAmounts>
            <exerciseDate>
                <value>1999-10-20</value>
            </exerciseDate>
            <expirationDate>
                <value>2007-10-16</value>
            </expirationDate>
            <underlyingSecurity>
                <underlyingSecurityTitle>
                    <value>Applied Biosystems Group Common Stock</value>
                </underlyingSecurityTitle>
                <underlyingSecurityShares>
                    <value>600</value>
                </underlyingSecurityShares>
            </underlyingSecurity>
            <postTransactionAmounts>
                <sharesOwnedFollowingTransaction>
                    <value>1800</value>
                </sharesOwnedFollowingTransaction>
            </postTransactionAmounts>
            <ownershipNature>
                <directOrIndirectOwnership>
                    <value>D</value>
                </directOrIndirectOwnership>
                <natureOfOwnership>
                    <value></value>
                </natureOfOwnership>
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        </derivativeTransaction>
        <derivativeTransaction>
            <securityTitle>
                <value>Celera Group Director Stock Options-Right to Buy</value>
            </securityTitle>
            <conversionOrExercisePrice>
                <value>6.5525</value>
            </conversionOrExercisePrice>
            <transactionDate>
                <value>2007-05-02</value>
            </transactionDate>
            <transactionCoding>
                <transactionFormType>4</transactionFormType>
                <transactionCode>M</transactionCode>
                <equitySwapInvolved>0</equitySwapInvolved>
            </transactionCoding>
            <transactionAmounts>
                <transactionShares>
                    <value>150</value>
                </transactionShares>
                <transactionPricePerShare>
                    <value>0</value>
                </transactionPricePerShare>
                <transactionAcquiredDisposedCode>
                    <value>D</value>
                </transactionAcquiredDisposedCode>
            </transactionAmounts>
            <exerciseDate>
                <value>1999-10-20</value>
            </exerciseDate>
            <expirationDate>
                <value>2007-10-16</value>
            </expirationDate>
            <underlyingSecurity>
                <underlyingSecurityTitle>
                    <value>Celera Group Common Stock</value>
                </underlyingSecurityTitle>
                <underlyingSecurityShares>
                    <value>150</value>
                </underlyingSecurityShares>
            </underlyingSecurity>
            <postTransactionAmounts>
                <sharesOwnedFollowingTransaction>
                    <value>450</value>
                </sharesOwnedFollowingTransaction>
            </postTransactionAmounts>
            <ownershipNature>
                <directOrIndirectOwnership>
                    <value>D</value>
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                <natureOfOwnership>
                    <value></value>
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            </ownershipNature>
        </derivativeTransaction>
    </derivativeTable>

    <footnotes>
        <footnote id="F1">Reflects sale by independent third-party administrator under the issuer's Insider Diversification Program.</footnote>
        <footnote id="F2">The reporting person is a co-trustee of the Suzanne L. Ayers Living Trust, a trust for the benefit of the reporting person's wife.</footnote>
    </footnotes>

    <remarks>This is the first of two forms being filed by the reporting person on May 4, 2007.</remarks>

    <ownerSignature>
        <signatureName>/s/ Thomas P. Livingston, Attorney-in-Fact for Richard H. Ayers</signatureName>
        <signatureDate>2007-05-04</signatureDate>
    </ownerSignature>
</ownershipDocument>
</XML>
</TEXT>
</DOCUMENT>
</SEC-DOCUMENT>
-----END PRIVACY-ENHANCED MESSAGE-----