Revenue (Details) - USD ($) $ in Thousands	3 Months Ended	12 Months Ended
	Mar. 31, 2020	Dec. 31, 2021	Dec. 31, 2020	Dec. 31, 2019
Revenue
Revenue, contract liabilities		$ 3,400
Revenues		4,651	$ 7,418	$ 3,573
Contract assets		0	0
Contract liability		433	3,386
Product development and licensing agreements
Revenue
Revenues		31	2,301	473
Contracts and grants
Revenue
Revenues		4,620	5,117	3,100
Rockefeller | Product development and licensing agreements
Revenue
Revenues	$ 1,800
Rockefeller | Contracts and grants
Revenue
Revenues		4,100	4,500	$ 2,600
IAVI & Frontier | Contracts and grants
Revenue
Revenues		$ 400	$ 400