License Agreements and Grants - National Institute of Health Grant (Details) - National Institute of Health Grant - USD ($) $ in Thousands	1 Months Ended	3 Months Ended		24 Months Ended	26 Months Ended
	Feb. 28, 2015	Mar. 31, 2017	Mar. 31, 2016	Jan. 31, 2017	Mar. 31, 2017
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Length of project (years)	5 years
Total grant funding	$ 5,000	$ 3	$ 9	$ 818	$ 560