N-Q 1 main.htm

UNITED STATES
SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

FORM N-Q

QUARTERLY SCHEDULE OF PORTFOLIO HOLDINGS OF REGISTERED MANAGEMENT INVESTMENT COMPANY

Investment Company Act file number 811-3583

Fidelity Mt. Vernon Street Trust
(Exact name of registrant as specified in charter)

82 Devonshire St., Boston, Massachusetts 02109
(Address of principal executive offices) (Zip code)

Eric D. Roiter, Secretary

82 Devonshire St.

Boston, Massachusetts 02109
(Name and address of agent for service)

Registrant's telephone number, including area code: 617-563-7000

Date of fiscal year end:

November 30

Date of reporting period:

February 29, 2008

Item 1. Schedule of Investments

Quarterly Holdings Report

for

Fidelity ® Aggressive Growth Fund

February 29, 2008

1.797938.104

FEG-QTLY-0408

Investments February 29, 2008 (Unaudited)

Showing Percentage of Net Assets

Common Stocks - 97.7%

Shares

Value (000s)

CONSUMER DISCRETIONARY - 9.4%

Internet & Catalog Retail - 1.0%

Priceline.com, Inc. (a)

243,900

$ 27,809

Specialty Retail - 6.8%

Abercrombie & Fitch Co. Class A

1,000,200

77,546

Tween Brands, Inc. (a)(d)

1,063,857

31,490

Urban Outfitters, Inc. (a)(d)

2,202,200

63,379

Zumiez, Inc. (a)(d)(e)

1,640,579

28,841

201,256

Textiles, Apparel & Luxury Goods - 1.6%

Polo Ralph Lauren Corp. Class A

780,520

48,541

TOTAL CONSUMER DISCRETIONARY

277,606

ENERGY - 15.5%

Energy Equipment & Services - 1.1%

FMC Technologies, Inc. (a)

602,300

34,126

Oil, Gas & Consumable Fuels - 14.4%

Chesapeake Energy Corp.

682,900

30,881

CONSOL Energy, Inc.

819,712

62,282

Denbury Resources, Inc. (a)

1,240,804

39,569

Gulfport Energy Corp. (a)

2,058,101

29,719

Hess Corp.

687,600

64,071

OPTI Canada, Inc. (a)

1,951,900

34,315

Peabody Energy Corp.

658,602

37,290

Quicksilver Resources, Inc. (a)

894,776

30,780

Range Resources Corp.

506,356

30,979

Southwestern Energy Co. (a)

487,807

31,820

Suncor Energy, Inc.

341,300

35,203

426,909

TOTAL ENERGY

461,035

FINANCIALS - 13.0%

Capital Markets - 2.4%

Greenhill & Co., Inc. (d)

486,019

31,596

Indiabulls Securities Ltd. (a)(f)

1,803,517

9,126

T. Rowe Price Group, Inc.

593,679

29,999

70,721

Diversified Financial Services - 4.7%

Bolsa de Mercadorias & Futuros - BM&F SA

1,940,800

21,014

Bovespa Holding SA

1,018,000

16,218

Common Stocks - continued

Shares

Value (000s)

FINANCIALS - continued

Diversified Financial Services - continued

Indiabulls Financial Services Ltd.

1,803,517

$ 27,130

JSE Ltd.

3,555,298

32,917

MarketAxess Holdings, Inc. (a)(e)

1,545,120

14,478

MSCI, Inc. Class A

971,579

28,885

140,642

Real Estate Investment Trusts - 1.1%

Annaly Capital Management, Inc.

1,560,900

32,295

Real Estate Management & Development - 4.8%

Dev Property Development PLC (e)

13,800,000

24,262

Indiabulls Real Estate Ltd. (a)

7,560,167

118,339

142,601

TOTAL FINANCIALS

386,259

HEALTH CARE - 28.9%

Biotechnology - 7.1%

Alnylam Pharmaceuticals, Inc. (a)(d)(e)

3,134,541

89,021

Amylin Pharmaceuticals, Inc. (a)(d)

1,341,032

35,497

BioMarin Pharmaceutical, Inc. (a)

829,212

31,543

CytRx Corp. (a)(d)(e)

9,152,702

16,475

Isis Pharmaceuticals, Inc. (a)(d)

2,701,049

38,895

211,431

Health Care Equipment & Supplies - 17.1%

ArthroCare Corp. (a)(d)(e)

2,789,235

111,988

Conceptus, Inc. (a)(d)

856,608

14,614

Cyberonics, Inc. (a)(d)(e)

2,691,800

34,724

Masimo Corp.

487,800

15,639

NeuroMetrix, Inc. (a)(d)

11,869

29

NuVasive, Inc. (a)(d)(e)

3,434,150

132,352

St. Jude Medical, Inc. (a)

4,080,016

175,358

TranS1, Inc. (d)(e)

1,643,765

21,747

506,451

Health Care Technology - 1.8%

Eclipsys Corp. (a)

1,009,838

21,560

MedAssets, Inc.

1,794,786

32,593

54,153

Common Stocks - continued

Shares

Value (000s)

HEALTH CARE - continued

Life Sciences Tools & Services - 2.9%

AMAG Pharmaceuticals, Inc. (d)(e)

1,072,364

$ 46,937

QIAGEN NV (a)

1,706,400

37,524

84,461

TOTAL HEALTH CARE

856,496

INDUSTRIALS - 6.6%

Airlines - 0.6%

AirTran Holdings, Inc. (a)(d)

2,415,142

17,437

Commercial Services & Supplies - 2.4%

CoStar Group, Inc. (a)(d)

512,996

21,300

EnergySolutions, Inc.

724,400

15,792

IHS, Inc. Class A (a)

561,573

34,621

71,713

Construction & Engineering - 2.0%

URS Corp. (a)

1,466,527

59,072

Marine - 0.5%

Ultrapetrol (Bahamas) Ltd. (a)

1,172,228

16,411

Road & Rail - 1.1%

Landstar System, Inc.

682,979

31,677

TOTAL INDUSTRIALS

196,310

INFORMATION TECHNOLOGY - 17.7%

Communications Equipment - 5.2%

Infinera Corp.

739,909

8,642

Juniper Networks, Inc. (a)

3,631,011

97,384

Starent Networks Corp. (d)

2,998,119

47,160

153,186

Electronic Equipment & Instruments - 1.1%

Itron, Inc. (a)(d)

329,432

31,405

Internet Software & Services - 7.4%

Omniture, Inc. (a)(d)(e)

4,778,328

109,806

ValueClick, Inc. (a)(d)

4,381,200

84,601

WebMD Health Corp. Class A (a)(d)

935,206

26,055

220,462

Semiconductors & Semiconductor Equipment - 3.0%

Cavium Networks, Inc.

1,642,864

23,362

Common Stocks - continued

Shares

Value (000s)

INFORMATION TECHNOLOGY - continued

Semiconductors & Semiconductor Equipment - continued

Hittite Microwave Corp. (a)

1,029,016

$ 34,071

KLA-Tencor Corp.

731,700

30,739

88,172

Software - 1.0%

Concur Technologies, Inc. (a)(d)

1,051,000

30,731

TOTAL INFORMATION TECHNOLOGY

523,956

MATERIALS - 3.4%

Chemicals - 1.0%

Albemarle Corp.

788,954

29,933

Metals & Mining - 2.4%

Aquarius Platinum Ltd. (Australia)

2,254,523

33,377

Steel Dynamics, Inc.

623,900

36,348

69,725

TOTAL MATERIALS

99,658

TELECOMMUNICATION SERVICES - 0.7%

Diversified Telecommunication Services - 0.7%

Time Warner Telecom, Inc. Class A (sub. vtg.) (a)

1,334,807

21,277

UTILITIES - 2.5%

Electric Utilities - 2.1%

PPL Corp.

658,632

29,889

Public Power Corp. of Greece

769,800

32,691

62,580

Independent Power Producers & Energy Traders - 0.4%

Ocean Power Technologies, Inc. (d)(e)

1,007,757

12,536

TOTAL UTILITIES

75,116

TOTAL COMMON STOCKS

(Cost $3,021,502)

2,897,713

Money Market Funds - 8.0%

Fidelity Cash Central Fund, 3.24% (b)

43,992,128

43,992

Fidelity Securities Lending Cash Central Fund, 3.25% (b)(c)

194,213,275

194,213

TOTAL MONEY MARKET FUNDS

(Cost $238,205)

238,205

Cash Equivalents - 0.5%

Maturity Amount (000s)

Value (000s)

Investments in repurchase agreements in a joint trading account at 3.18%, dated 2/29/08 due 3/3/08 (Collateralized by U.S. Government Obligations) #
(Cost $14,464)

$ 14,468

$ 14,464

TOTAL INVESTMENT PORTFOLIO - 106.2%

(Cost $3,274,171)

3,150,382

NET OTHER ASSETS - (6.2)%

(182,715)

NET ASSETS - 100%

$ 2,967,667

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

(e) Affiliated company

(f) Restricted securities - Investment in securities not registered under the Securities Act of 1933 (excluding 144A issues). At the end of the period, the value of restricted securities (excluding 144A issues) amounted to $9,126,000 or 0.3% of net assets.

Additional information on each holding is as follows:

Security

Acquisition Date

Acquisition Cost (000s)

Indiabulls Securities Ltd.

5/2/06 - 1/24/07

$ 3,090

# Additional Information on each counterparty to the repurchase agreement is as follows:

Repurchase Agreement / Counterparty

Value
(Amounts in thousands)

$14,464,000 due 3/03/08 at 3.18%

BNP Paribas Securities Corp.

$ 5,361

Banc of America Securities LLC

4,010

Societe Generale, New York Branch

5,093

$ 14,464

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned
(Amounts in thousands)

Fidelity Cash Central Fund

$ 412

Fidelity Securities Lending Cash Central Fund

1,135

Total

$ 1,547

Other Affiliated Issuers

An affiliated company is a company in which the fund has ownership of at least 5% of the voting securities. Fiscal year to date transactions with companies which are or were affiliates are as follows:

Affiliate
(Amounts in thousands)

Value, beginning of period

Purchases

Sales Proceeds

Dividend Income

Value,
end of
period

Alnylam Pharmaceuticals, Inc.

$ 102,229

$ 21

$ -

$ -

$ 89,021

AMAG Pharmaceuticals, Inc.

65,945

16,822

23,231

-

46,937

ArthroCare Corp.

72,823

74,151

4,884

-

111,988

Conceptus, Inc.

35,899

6,780

25,383

-

-

Cyberonics, Inc.

36,716

-

-

-

34,724

CytRx Corp.

23,616

3,941

-

-

16,475

Dev Property Development PLC

22,197

-

-

-

24,262

MarketAxess Holdings, Inc.

22,188

-

-

-

14,478

NeuroMetrix, Inc.

10,538

-

5,190

-

-

Northstar Neuroscience, Inc.

23,877

-

3,468

-

-

NuVasive, Inc.

146,192

-

-

-

132,352

Ocean Power Technologies, Inc.

14,623

-

-

-

12,536

Omniture, Inc.

57,785

71,643

-

-

109,806

Omrix Biopharmaceuticals, Inc.

40,074

-

34,594

-

-

TranS1, Inc.

30,171

1,845

209

-

21,747

Zumiez, Inc.

-

31,637

-

-

28,841

Total

$ 704,873

$ 206,840

$ 96,959

$ -

$ 643,167

Other Information

The following is a summary of the inputs used, as of February 29, 2008, involving the Fund's assets carried at value. The inputs or methodology used for valuing securities may not be an indication of the risk associated with investing in those securities.

Valuation Inputs at Reporting Date:

Description
(Amounts in thousands)

Total

Level 1

Level 2

Level 3

Investments in Securities

$ 3,150,382

$ 2,858,076

$ 283,180

$ 9,126

The following is a reconciliation of assets for which Level 3 inputs were used in determining value:

(Amounts in thousands)

Investments in Securities

Beginning Balance

$ -

Total Realized Gain (Loss)

-

Total Unrealized Gain (Loss)

6,036

Cost of Purchases

3,090

Proceeds of Sales

-

Amortization/Accretion

-

Transfer in/out of Level 3

-

Ending Balance

$ 9,126

The information used in the above reconciliation represents fiscal year to date activity for any Investment Securities identified as using Level 3 inputs at either the beginning or the end of the current fiscal period. Transfers in or out of Level 3 represents either the beginning value (for transfers in), or the ending value (for transfers out) of any Security or Instrument where a change in the pricing level occurred from the beginning to the end of the period.

Income Tax Information

At February 29, 2008, the aggregate cost of investment securities for income tax purposes was $3,278,689,000. Net unrealized depreciation aggregated $128,307,000, of which $303,581,000 related to appreciated investment securities and $431,888,000 related to depreciated investment securities.

Investment Valuation

Investments are valued as of the close of business of the New York Stock Exchange (NYSE), normally 4:00 p.m. Eastern time. Security transactions are accounted for as of trade date. Wherever possible, the Fund uses independent pricing services approved by the Board of Trustees to value its investments.

Equity securities, including restricted securities, for which market quotations are readily available, are valued at the last reported sale price or official closing price as reported by an independent pricing service on the primary market or exchange on which they are traded. In the event there were no sales during the day or closing prices are not available, securities are valued at the last quoted bid price. Investments in open-end mutual funds, including the Fidelity Central Funds, are valued at their closing net asset value each business day. Short-term securities with remaining maturities of sixty days or less for which quotations are not readily available are valued at amortized cost, which approximates value.

When current market prices or quotations are not readily available or reliable, valuations may be determined in good faith in accordance with procedures adopted by the Board of Trustees. The frequency with which these procedures are used cannot be predicted and may be utilized to a significant extent. The value of securities used for NAV calculation under these procedures may differ from published prices for the same securities.

The Fund adopted the provisions of Statement of Financial Accounting Standards No. 157, "Fair Value Measurements" (SFAS 157), effective with the beginning of the Fund's fiscal year. SFAS 157 establishes a hierarchy that prioritizes the inputs to valuation techniques giving the highest priority to readily available unadjusted quoted prices in active markets for identical assets (level 1 measurements) and the lowest priority to unobservable inputs (level 3 measurements) when market prices are not readily available or reliable. The three levels of the hierarchy under SFAS 157 are described below:

Level 1 - Quoted prices in active markets for identical securities.

Level 2 - Prices determined using other significant observable inputs. Observable inputs are inputs that other market participants would use in pricing a security. These may include quoted prices for similar securities, interest rates, prepayment speeds, credit risk and others.

Level 3 - Prices determined using significant unobservable inputs. In situations where quoted prices or observable inputs are unavailable (for example, when there is little or no market activity for an investment at the end of the period), unobservable inputs may be used. Unobservable inputs reflect the Fund's own assumptions about the factors market participants would use in pricing an investment, and would be based on the best information available.

Changes in valuation techniques may result in transfers in or out of an investment's assigned level within the hierarchy.

For additional information on the fund's policy regarding valuation of investments and other significant accounting policies, please refer to the fund's most recent semiannual or annual shareholder report.

Quarterly Report

The fund's schedule of investments as of the date on the cover of this report has not been audited. This report is provided for the general information of the fund's shareholders. For more information regarding the fund and its holdings, please see the fund's most recent prospectus and annual report.

Third party trademarks and service marks are the property of their respective owners. All other trademarks and service marks are the property of FMR LLC or an affiliate.

Quarterly Report

Quarterly Holdings Report

for

Fidelity ® Growth Company Fund

February 29, 2008

1.797941.104

GCF-QTLY-0408

Investments February 29, 2008 (Unaudited)

Showing Percentage of Net Assets

Common Stocks - 99.5%

Shares

Value (000s)

CONSUMER DISCRETIONARY - 4.8%

Auto Components - 0.1%

Johnson Controls, Inc.

1,000,000

$ 32,860

Hotels, Restaurants & Leisure - 0.4%

Chipotle Mexican Grill, Inc. Class A (a)

40,000

3,972

McDonald's Corp.

1,070,000

57,898

Starbucks Corp. (a)

1,458,400

26,207

Starwood Hotels & Resorts Worldwide, Inc.

725,000

34,314

122,391

Household Durables - 0.0%

Tupperware Brands Corp.

50,000

1,824

Internet & Catalog Retail - 0.4%

Amazon.com, Inc. (a)

2,275,000

146,669

Leisure Equipment & Products - 0.3%

Callaway Golf Co. (e)

5,839,862

88,999

Media - 1.1%

CBS Corp. Class B

1,115,988

25,467

Comcast Corp. Class A

2,187,500

42,744

Morningstar, Inc. (a)(d)

1,435,000

93,548

News Corp. Class A

4,705,000

86,619

The Walt Disney Co.

84,500

2,739

Time Warner, Inc.

2,801,950

43,738

TiVo, Inc. (a)(d)(e)

9,894,029

85,880

380,735

Multiline Retail - 0.6%

Kohl's Corp. (a)

1,415,000

62,883

Nordstrom, Inc.

250,000

9,258

Target Corp.

2,270,000

119,425

191,566

Specialty Retail - 0.9%

Best Buy Co., Inc.

347,500

14,946

Gamestop Corp. Class A (a)

296,300

12,551

Gap, Inc.

717,825

14,479

Home Depot, Inc.

3,635,000

96,509

Lowe's Companies, Inc.

3,335,800

79,959

Staples, Inc.

3,120,602

69,433

287,877

Textiles, Apparel & Luxury Goods - 1.0%

Coach, Inc. (a)

187,200

5,676

Crocs, Inc. (a)(d)

495,000

12,038

Geox SpA

2,285,000

33,566

Common Stocks - continued

Shares

Value (000s)

CONSUMER DISCRETIONARY - continued

Textiles, Apparel & Luxury Goods - continued

Hanesbrands, Inc. (a)(d)

2,275,000

$ 66,203

Lululemon Athletica, Inc. (d)(e)

5,110,219

137,465

NIKE, Inc. Class B (d)

1,579,000

95,056

Under Armour, Inc. Class A (sub. vtg.) (a)(d)

48,017

1,768

351,772

TOTAL CONSUMER DISCRETIONARY

1,604,693

CONSUMER STAPLES - 9.9%

Beverages - 2.2%

PepsiCo, Inc.

3,816,640

265,485

The Coca-Cola Co.

7,832,500

457,888

723,373

Food & Staples Retailing - 2.8%

Costco Wholesale Corp.

1,160,800

71,877

CVS Caremark Corp.

3,012,780

121,656

Safeway, Inc.

1,110,000

31,901

Sysco Corp.

478,200

13,418

Wal-Mart Stores, Inc.

13,310,300

660,058

Walgreen Co.

1,195,000

43,629

Whole Foods Market, Inc.

120,000

4,218

946,757

Food Products - 2.0%

Archer Daniels Midland Co.

4,410,000

198,891

Campbell Soup Co.

1,800,000

58,122

Cosan Ltd. Class A (e)

6,473,600

97,104

General Mills, Inc.

302,800

16,954

Groupe Danone

885,000

67,924

Hershey Co.

320,000

11,866

Kellogg Co.

1,590,000

80,645

Kraft Foods, Inc. Class A

1,296,065

40,398

McCormick & Co., Inc. (non-vtg.)

365,000

12,574

Smithfield Foods, Inc. (a)

730,000

20,112

Tyson Foods, Inc. Class A

550,000

7,926

Wm. Wrigley Jr. Co.

800,000

47,888

660,404

Household Products - 1.4%

Clorox Co.

570,000

33,168

Common Stocks - continued

Shares

Value (000s)

CONSUMER STAPLES - continued

Household Products - continued

Colgate-Palmolive Co.

2,075,000

$ 157,887

Procter & Gamble Co.

4,199,483

277,922

468,977

Personal Products - 0.5%

Avon Products, Inc.

4,600,000

175,076

Tobacco - 1.0%

Altria Group, Inc.

4,375,380

320,015

TOTAL CONSUMER STAPLES

3,294,602

ENERGY - 10.8%

Energy Equipment & Services - 2.3%

Baker Hughes, Inc.

387,400

26,068

Diamond Offshore Drilling, Inc.

827,900

100,035

FMC Technologies, Inc. (a)

1,155,000

65,442

Schlumberger Ltd. (NY Shares)

2,189,000

189,239

Transocean, Inc. (a)

580,668

81,590

Weatherford International Ltd. (a)

4,120,240

283,967

746,341

Oil, Gas & Consumable Fuels - 8.5%

Anadarko Petroleum Corp. (d)

2,685,000

171,142

Apache Corp.

2,510,000

287,922

Cameco Corp.

2,485,000

97,349

Canadian Natural Resources Ltd.

513,500

38,490

ConocoPhillips

453,810

37,535

CONSOL Energy, Inc.

2,734,408

207,760

Devon Energy Corp.

3,540,000

363,629

EnCana Corp.

5,120,000

390,380

EOG Resources, Inc.

1,720,000

204,663

Exxon Mobil Corp.

610,000

53,076

Hess Corp.

7,955,000

741,247

Newfield Exploration Co. (a)

2,990,000

165,586

Noble Energy, Inc.

850,000

65,790

Valero Energy Corp.

325,000

18,775

2,843,344

TOTAL ENERGY

3,589,685

Common Stocks - continued

Shares

Value (000s)

FINANCIALS - 4.6%

Capital Markets - 1.3%

Ameriprise Financial, Inc.

2,345,809

$ 118,792

Charles Schwab Corp.

4,121,475

80,822

Franklin Resources, Inc.

520,000

49,072

Goldman Sachs Group, Inc.

200,000

33,926

Merrill Lynch & Co., Inc.

1,500,000

74,340

State Street Corp.

835,000

65,589

422,541

Commercial Banks - 0.9%

Banco Bradesco SA (PN) sponsored ADR

1,480,000

46,457

Banco Itau Holding Financeira SA sponsored ADR (non-vtg.)

3,680,000

93,251

Commerce Bancorp, Inc.

670,000

25,313

Synovus Financial Corp.

1,261,900

14,550

Uniao de Bancos Brasileiros SA (Unibanco) GDR

805,000

109,174

Wells Fargo & Co.

355,000

10,377

299,122

Consumer Finance - 1.0%

American Express Co.

1,617,548

68,422

Discover Financial Services

16,925,000

255,398

323,820

Diversified Financial Services - 1.2%

Bank of America Corp.

1,390,000

55,239

Bolsa de Mercadorias & Futuros - BM&F SA

9,383,100

101,593

Bovespa Holding SA

11,686,000

186,171

JPMorgan Chase & Co.

1,250,000

50,813

393,816

Insurance - 0.2%

American International Group, Inc.

1,063,750

49,847

Prudential Financial, Inc.

233,000

17,002

66,849

Thrifts & Mortgage Finance - 0.0%

Fannie Mae

730,000

20,185

TOTAL FINANCIALS

1,526,333

HEALTH CARE - 23.0%

Biotechnology - 11.0%

Acadia Pharmaceuticals, Inc. (a)(d)(e)

3,499,994

34,720

Affymax, Inc. (a)(e)

1,255,000

24,159

Alexion Pharmaceuticals, Inc. (a)(d)(e)

3,759,225

227,847

Common Stocks - continued

Shares

Value (000s)

HEALTH CARE - continued

Biotechnology - continued

Alkermes, Inc. (a)(e)

10,182,261

$ 131,758

Alnylam Pharmaceuticals, Inc. (a)

556,247

15,797

Amgen, Inc. (a)

1,227,605

55,881

Amylin Pharmaceuticals, Inc. (a)(d)(e)

13,482,460

356,881

Array Biopharma, Inc. (a)(d)(e)

4,700,070

26,320

Biogen Idec, Inc. (a)

1,104,164

64,439

BioMarin Pharmaceutical, Inc. (a)(d)

3,470,793

132,029

Celgene Corp. (a)

20,046,744

1,130,035

Cepheid, Inc. (a)(d)

2,530,000

70,030

Cougar Biotechnology, Inc. (a)

912,347

24,277

CV Therapeutics, Inc. (a)(d)(e)

5,966,600

34,845

Genentech, Inc. (a)

1,475,800

111,792

Genzyme Corp. (a)

520,000

36,878

Gilead Sciences, Inc. (a)

1,700,000

80,444

GTx, Inc. (a)(d)(e)

2,396,497

39,231

Human Genome Sciences, Inc. (a)(d)(e)

8,074,010

47,717

Immunomedics, Inc. (a)(d)(e)

6,102,700

15,074

InterMune, Inc. (a)(d)(e)

3,897,713

54,880

Isis Pharmaceuticals, Inc. (a)(d)(e)

9,265,033

133,416

Medarex, Inc. (a)(d)

5,936,220

55,207

Millennium Pharmaceuticals, Inc. (a)

185,000

2,588

Momenta Pharmaceuticals, Inc. (a)

406,000

4,125

Myriad Genetics, Inc. (a)(d)(e)

3,224,248

119,362

ONYX Pharmaceuticals, Inc. (a)

1,430,000

39,068

OREXIGEN Therapeutics, Inc. (e)

2,219,921

28,748

PDL BioPharma, Inc. (a)

2,605,450

41,635

Regeneron Pharmaceuticals, Inc. (a)(e)

6,388,948

126,310

Rigel Pharmaceuticals, Inc. (a)(d)(e)

3,365,396

66,702

Seattle Genetics, Inc. (a)(e)

7,643,955

68,719

Sunesis Pharmaceuticals, Inc. (a)(e)

3,156,200

4,419

Transition Therapeutics, Inc. (a)(e)

2,332,446

23,608

Trubion Pharmaceuticals, Inc. (a)(d)(e)

1,775,000

12,620

Vertex Pharmaceuticals, Inc. (a)(d)(e)

13,200,267

231,005

3,672,566

Health Care Equipment & Supplies - 2.3%

Alcon, Inc.

40,000

5,789

Baxter International, Inc.

2,455,000

144,894

Becton, Dickinson & Co.

1,347,900

121,877

Covidien Ltd.

1,525,000

65,255

Common Stocks - continued

Shares

Value (000s)

HEALTH CARE - continued

Health Care Equipment & Supplies - continued

Gen-Probe, Inc. (a)(e)

3,703,676

$ 177,073

Insulet Corp.

951,400

16,297

Kinetic Concepts, Inc. (a)(d)

993,000

51,030

Medtronic, Inc.

1,274,964

62,932

St. Jude Medical, Inc. (a)

834,200

35,854

Thoratec Corp. (a)(d)(e)

5,297,765

76,871

TomoTherapy, Inc.

245,000

3,227

761,099

Health Care Providers & Services - 0.9%

Cardinal Health, Inc.

860,000

50,860

Laboratory Corp. of America Holdings (a)

525,000

40,588

McKesson Corp.

765,000

44,951

Medco Health Solutions, Inc. (a)

1,412,720

62,598

UnitedHealth Group, Inc.

2,186,400

101,624

300,621

Health Care Technology - 0.1%

HLTH Corp. (a)(d)

3,007,800

35,612

Life Sciences Tools & Services - 1.9%

Affymetrix, Inc. (a)(d)(e)

6,903,190

132,403

Applera Corp.:

- Applied Biosystems Group

211,600

7,133

- Celera Genomics Group (a)

9,579,848

132,777

Bruker BioSciences Corp. (a)

150,000

2,051

Exelixis, Inc. (a)(e)

10,507,384

67,037

Illumina, Inc. (a)(d)(e)

3,845,839

278,477

Millipore Corp. (a)

183,194

12,805

632,683

Pharmaceuticals - 6.8%

Abbott Laboratories

2,051,500

109,858

Allergan, Inc.

755,000

44,719

ARYx Therapeutics, Inc.

305,215

2,567

Auxilium Pharmaceuticals, Inc. (a)(e)

2,310,583

74,031

Bristol-Myers Squibb Co.

1,754,700

39,674

Dr. Reddy's Laboratories Ltd. sponsored ADR (d)

800,000

11,448

Elan Corp. PLC sponsored ADR (a)(d)

46,171,262

1,051,320

Eli Lilly & Co.

1,205,000

60,274

Johnson & Johnson

3,488,300

216,135

Map Pharmaceuticals, Inc.

925,000

12,025

Merck & Co., Inc.

4,320,100

191,380

Nastech Pharmaceutical Co., Inc. (a)(d)(e)

2,581,059

5,962

Common Stocks - continued

Shares

Value (000s)

HEALTH CARE - continued

Pharmaceuticals - continued

Pfizer, Inc.

1,451,000

$ 32,328

Schering-Plough Corp.

1,987,600

43,131

Sepracor, Inc. (a)(d)(e)

11,170,129

239,823

Sirtris Pharmaceuticals, Inc. (d)

721,435

8,614

Wyeth

3,075,200

134,140

2,277,429

TOTAL HEALTH CARE

7,680,010

INDUSTRIALS - 7.2%

Aerospace & Defense - 1.6%

Embraer - Empresa Brasileira de Aeronautica SA sponsored ADR

340,000

15,069

Honeywell International, Inc.

2,485,000

142,987

Lockheed Martin Corp.

1,750,100

180,610

Raytheon Co.

2,335,000

151,401

The Boeing Co.

670,000

55,469

545,536

Air Freight & Logistics - 0.3%

United Parcel Service, Inc. Class B

1,619,800

113,775

Airlines - 1.6%

AMR Corp. (a)

127,500

1,633

Continental Airlines, Inc. Class B (a)(e)

6,000,000

145,080

Copa Holdings SA Class A

40,000

1,444

JetBlue Airways Corp. (a)(d)(e)

18,003,143

98,117

Northwest Airlines Corp. (a)

11,453,700

153,823

Ryanair Holdings PLC sponsored ADR (a)

20,000

572

Ryanair Holdings PLC warrants (UBS Warrant Programme) 2/25/10 (a)

3,230,000

30,672

Southwest Airlines Co.

7,273,515

89,173

520,514

Commercial Services & Supplies - 0.1%

EnergySolutions, Inc.

911,400

19,869

Construction & Engineering - 0.2%

Fluor Corp.

195,000

27,154

KBR, Inc. (a)

1,315,000

43,829

70,983

Electrical Equipment - 0.8%

American Superconductor Corp. (a)(d)(e)

2,540,000

57,379

Common Stocks - continued

Shares

Value (000s)

INDUSTRIALS - continued

Electrical Equipment - continued

First Solar, Inc. (a)

997,090

$ 204,603

Sunpower Corp. Class A (a)(d)

82,500

5,422

267,404

Industrial Conglomerates - 1.2%

3M Co.

470,000

36,848

General Electric Co.

10,700,000

354,598

391,446

Machinery - 0.8%

AGCO Corp. (a)

2,460,400

159,582

Caterpillar, Inc.

355,000

25,677

CNH Global NV

230,000

11,845

Deere & Co.

690,000

58,795

ITT Corp.

70,000

3,937

TurboChef Technologies, Inc. (a)(d)(e)

2,898,650

24,639

284,475

Road & Rail - 0.6%

Norfolk Southern Corp.

1,485,000

78,542

Union Pacific Corp.

860,000

107,294

185,836

TOTAL INDUSTRIALS

2,399,838

INFORMATION TECHNOLOGY - 32.9%

Communications Equipment - 4.5%

Blue Coat Systems, Inc. (a)

770,000

18,080

Cisco Systems, Inc. (a)

12,712,600

309,806

Corning, Inc.

2,063,000

47,923

F5 Networks, Inc. (a)

854,446

18,935

Infinera Corp. (e)

7,629,253

89,110

Juniper Networks, Inc. (a)

885,000

23,736

Motorola, Inc.

165,000

1,645

QUALCOMM, Inc.

8,023,400

339,951

Research In Motion Ltd. (a)

3,892,800

404,073

Riverbed Technology, Inc. (a)(e)

7,070,664

141,838

Sonus Networks, Inc. (a)(d)(e)

26,835,720

89,095

1,484,192

Computers & Peripherals - 2.6%

Apple, Inc. (a)

3,345,959

418,312

Dell, Inc. (a)

1,235,600

24,527

Common Stocks - continued

Shares

Value (000s)

INFORMATION TECHNOLOGY - continued

Computers & Peripherals - continued

Hewlett-Packard Co.

3,160,000

$ 150,953

International Business Machines Corp.

49,800

5,670

Network Appliance, Inc. (a)

3,207,208

69,340

Palm, Inc. (d)(e)

10,115,952

65,450

SanDisk Corp. (a)

105,000

2,473

Seagate Technology

2,265,000

48,856

Sun Microsystems, Inc. (a)

449,125

7,366

Synaptics, Inc. (a)(d)(e)

3,085,082

82,649

875,596

Electronic Equipment & Instruments - 0.7%

Agilent Technologies, Inc. (a)

370,000

11,326

Itron, Inc. (a)

638,400

60,859

Mellanox Technologies Ltd. (e)

3,023,300

46,256

Trimble Navigation Ltd. (a)

2,425,000

66,300

Universal Display Corp. (a)(d)(e)

3,526,745

60,096

244,837

Internet Software & Services - 5.1%

Akamai Technologies, Inc. (a)

1,055,000

37,094

Alibaba.com Ltd.

792,500

1,921

Baidu.com, Inc. sponsored ADR (a)

10,000

2,513

eBay, Inc. (a)

2,777,200

73,207

Google, Inc. Class A (sub. vtg.) (a)

2,822,448

1,329,881

Internet Capital Group, Inc. (a)(e)

3,850,000

33,495

Mercadolibre, Inc.

226,300

8,181

Omniture, Inc. (a)

2,004,881

46,072

VeriSign, Inc. (a)

2,340,000

81,432

Yahoo!, Inc. (a)

3,111,340

86,433

1,700,229

IT Services - 0.8%

Cognizant Technology Solutions Corp. Class A (a)

1,584,716

47,874

Fidelity National Information Services, Inc.

375,000

15,559

Hewitt Associates, Inc. Class A (a)

2,045,000

80,696

Mastercard, Inc. Class A

230,000

43,700

The Western Union Co.

1,025,000

21,320

Total System Services, Inc.

2,260,659

50,254

259,403

Semiconductors & Semiconductor Equipment - 5.3%

Analog Devices, Inc.

949,900

25,571

Applied Materials, Inc.

1,770,400

33,939

Common Stocks - continued

Shares

Value (000s)

INFORMATION TECHNOLOGY - continued

Semiconductors & Semiconductor Equipment - continued

ASML Holding NV (NY Shares) (a)

648,888

$ 15,625

Atheros Communications, Inc. (a)(e)

5,678,042

138,090

Atheros Communications, Inc. (a)(e)(g)

1,741,486

42,353

Broadcom Corp. Class A (a)

2,300,000

43,493

Cree, Inc. (a)(d)(e)

8,526,131

263,457

Cypress Semiconductor Corp. (a)(e)

15,976,000

347,318

FEI Co. (a)(d)(e)

1,863,100

37,989

Intel Corp.

15,125,800

301,760

International Rectifier Corp. (a)(d)

710,000

16,167

KLA-Tencor Corp. (d)

510,000

21,425

Linear Technology Corp. (d)

625,800

17,341

Marvell Technology Group Ltd. (a)

498,310

5,636

Power Integrations, Inc. (a)(e)

2,961,308

77,882

Rambus, Inc. (a)(d)(e)

9,135,000

164,704

Rubicon Technology, Inc. (d)(e)

1,845,000

54,261

Samsung Electronics Co. Ltd.

50,000

29,309

Silicon Laboratories, Inc. (a)

1,860,315

57,577

Spansion, Inc. Class A (a)(d)

1,430,000

3,933

Tessera Technologies, Inc. (a)(d)

705,000

16,610

Texas Instruments, Inc.

971,000

29,091

Veeco Instruments, Inc. (a)(d)(e)

1,765,000

27,358

1,770,889

Software - 13.9%

Activision, Inc. (a)

1,660,000

45,235

Adobe Systems, Inc. (a)

843,236

28,375

Blackboard, Inc. (a)

73,200

2,106

BladeLogic, Inc. (d)(e)

2,722,101

51,992

Citrix Systems, Inc. (a)

4,940,000

162,674

Electronic Arts, Inc. (a)

640,000

30,266

Microsoft Corp.

18,810,000

512,008

Nintendo Co. Ltd. (d)

4,571,200

2,281,935

Oracle Corp. (a)

1,175,000

22,090

Red Hat, Inc. (a)(d)(e)

17,667,391

315,010

Salesforce.com, Inc. (a)(d)(e)

11,779,994

703,501

SuccessFactors, Inc. (d)(e)

3,151,600

26,221

Symantec Corp. (a)

2,050,800

34,535

VMware, Inc. Class A (d)

7,207,100

422,841

4,638,789

TOTAL INFORMATION TECHNOLOGY

10,973,935

Common Stocks - continued

Shares

Value (000s)

MATERIALS - 5.8%

Chemicals - 4.5%

Dow Chemical Co.

1,315,000

$ 49,562

E.I. du Pont de Nemours & Co.

500,000

23,210

Minerals Technologies, Inc. (e)

1,927,580

116,195

Monsanto Co.

8,869,978

1,026,079

OM Group, Inc. (a)(e)

1,760,000

106,533

Potash Corp. of Saskatchewan, Inc.

281,200

44,683

Rohm & Haas Co.

900,000

48,249

The Mosaic Co. (a)

575,000

63,998

Uralkali JSC ADR (a)(f)

434,300

17,593

1,496,102

Metals & Mining - 1.3%

Alcoa, Inc.

1,935,000

71,866

Barrick Gold Corp.

4,667,500

242,849

Companhia Vale do Rio Doce sponsored ADR

100,000

3,484

Freeport-McMoRan Copper & Gold, Inc. Class B

745,000

75,141

Nucor Corp.

910,000

58,759

452,099

TOTAL MATERIALS

1,948,201

TELECOMMUNICATION SERVICES - 0.2%

Diversified Telecommunication Services - 0.1%

AT&T, Inc.

1,115,000

38,835

Level 3 Communications, Inc. (a)

3,000,000

6,690

45,525

Wireless Telecommunication Services - 0.1%

Clearwire Corp. (d)

2,313,500

32,412

TOTAL TELECOMMUNICATION SERVICES

77,937

UTILITIES - 0.3%

Electric Utilities - 0.3%

Enernoc, Inc. (d)

125,000

1,848

Exelon Corp.

1,115,000

83,458

85,306

TOTAL COMMON STOCKS

(Cost $27,361,924)

33,180,540

Convertible Preferred Stocks - 0.1%

Shares

Value (000s)

HEALTH CARE - 0.1%

Biotechnology - 0.1%

Perlegen Sciences, Inc. Series D, 8.00% (a)(g)

12,820,512

$ 25,641

TOTAL CONVERTIBLE PREFERRED STOCKS

(Cost $20,000)

25,641

Money Market Funds - 2.8%

Fidelity Cash Central Fund, 3.24% (b)

95,405,442

95,405

Fidelity Securities Lending Cash Central Fund, 3.25% (b)(c)

836,444,015

836,444

TOTAL MONEY MARKET FUNDS

(Cost $931,849)

931,849

TOTAL INVESTMENT PORTFOLIO - 102.4%

(Cost $28,313,773)

34,138,030

NET OTHER ASSETS - (2.4)%

(816,349)

NET ASSETS - 100%

$ 33,321,681

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

(e) Affiliated company

(f) Security exempt from registration under Rule 144A of the Securities Act of 1933. These securities may be resold in transactions exempt from registration, normally to qualified institutional buyers. At the end of the period, the value of these securities amounted to $17,593,000 or 0.1% of net assets.

(g) Restricted securities - Investment in securities not registered under the Securities Act of 1933 (excluding 144A issues). At the end of the period, the value of restricted securities (excluding 144A issues) amounted to $67,994,000 or 0.2% of net assets.

Additional information on each holding is as follows:

Security

Acquisition Date

Acquisition Cost (Amounts in thousands)

Atheros Communications, Inc.

4/18/01

$ 15,000

Perlegen Sciences, Inc. Series D, 8.00%

2/23/05

$ 20,000

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned
(Amounts in thousands)

Fidelity Cash Central Fund

$ 1,597

Fidelity Securities Lending Cash Central Fund

9,301

Total

$ 10,898

Other Affiliated Issuers

An affiliated company is a company in which the fund has ownership of at least 5% of the voting securities. Fiscal year to date transactions with companies which are or were affiliates are as follows:

Affiliate
(Amounts in thousands)

Value, beginning of period

Purchases

Sales Proceeds

Dividend Income

Value,
end of
period

Acadia Pharmaceuticals, Inc.

$ 39,300

$ 368

$ -

$ -

$ 34,720

Affymax, Inc.

31,952

-

-

-

24,159

Affymetrix, Inc.

143,047

977

-

-

132,403

Alexion Pharmaceuticals, Inc.

270,064

3,208

-

-

227,847

Alkermes, Inc.

145,199

-

-

-

131,758

Affiliate
(Amounts in thousands)

Value, beginning of period

Purchases

Sales Proceeds

Dividend Income

Value,
end of
period

American Superconductor Corp.

$ 75,075

$ -

$ 13,249

$ -

$ 57,379

Amylin Pharmaceuticals, Inc.

514,819

55

-

-

356,881

Array Biopharma, Inc.

52,030

-

-

-

26,320

Atheros Communications, Inc.

167,445

-

-

-

138,090

Atheros Communications, Inc.

51,356

-

-

-

42,353

Auxilium Pharmaceuticals, Inc.

-

71,731

-

-

74,031

Blackboard, Inc.

93,335

-

76,737

-

-

BladeLogic, Inc.

37,853

25,748

-

-

51,992

Blue Coat Systems, Inc.

106,638

-

60,799

-

-

Callaway Golf Co.

97,496

4,731

3,057

395

88,999

Celgene Corp.

1,686,270

-

377,483

-

-

Continental Airlines, Inc. Class B

-

144,995

-

-

145,080

Cosan Ltd. Class A

61,799

10,096

-

-

97,104

Cree, Inc.

209,972

35

-

-

263,457

CV Therapeutics, Inc.

51,892

140

-

-

34,845

Cypress Semiconductor Corp.

530,723

-

-

-

347,318

Exelixis, Inc.

91,145

452

-

-

67,037

F5 Networks, Inc.

135,520

-

100,493

-

-

FEI Co.

46,410

-

-

-

37,989

Gen-Probe, Inc.

282,856

-

26,399

-

177,073

GTx, Inc.

32,999

1,890

-

-

39,231

Human Genome Sciences, Inc.

83,009

19,381

16,686

-

47,717

Illumina, Inc.

178,053

51,936

-

-

278,477

Immunomedics, Inc.

13,792

-

-

-

15,074

Infinera Corp.

92,112

61,683

-

-

89,110

InterMune, Inc.

63,299

-

-

-

54,880

Internet Capital Group, Inc.

43,005

-

-

-

33,495

Isis Pharmaceuticals, Inc.

151,242

11,076

-

-

133,416

JetBlue Airways Corp.

126,022

-

-

-

98,117

Lululemon Athletica, Inc.

126,886

53,376

-

-

137,465

Mellanox Technologies Ltd.

53,633

-

-

-

46,256

Minerals Technologies, Inc.

128,955

-

-

96

116,195

Affiliate
(Amounts in thousands)

Value, beginning of period

Purchases

Sales Proceeds

Dividend Income

Value,
end of
period

Myriad Genetics, Inc.

$ 192,523

$ -

$ 33,736

$ -

$ 119,362

Nastech Pharmaceutical Co., Inc.

9,808

-

-

-

5,962

OM Group, Inc.

63,889

36,490

-

-

106,533

Omniture, Inc.

90,056

-

33,933

-

-

OREXIGEN Therapeutics, Inc.

14,523

13,856

-

-

28,748

Palm, Inc.

68,591

1,595

-

-

65,450

Power Integrations, Inc.

86,294

2,540

-

-

77,882

Rambus, Inc.

158,804

18,979

-

-

164,704

Red Hat, Inc.

228,076

127,347

7,372

-

315,010

Regeneron Pharmaceuticals, Inc.

139,151

-

-

-

126,310

Rigel Pharmaceuticals, Inc.

21,587

8,512

-

-

66,702

Riverbed Technology, Inc.

199,958

-

-

-

141,838

Rubicon Technology, Inc.

24,150

12,445

-

-

54,261

Salesforce.com, Inc.

662,606

5,774

-

-

703,501

Seattle Genetics, Inc.

73,220

9,018

-

-

68,719

Sepracor, Inc.

282,362

15,227

-

-

239,823

Sonus Networks, Inc.

176,579

-

-

-

89,095

SuccessFactors, Inc.

28,946

14,473

-

-

26,221

Sunesis Pharmaceuticals, Inc.

6,691

-

-

-

4,419

Synaptics, Inc.

144,712

23,063

-

-

82,649

Thoratec Corp.

103,783

-

-

-

76,871

TiVo, Inc.

73,514

694

-

-

85,880

Transition Therapeutics, Inc.

22,721

-

-

-

23,608

Trubion Pharmaceuticals, Inc.

17,732

-

-

-

12,620

TurboChef Technologies, Inc.

43,103

-

-

-

24,639

Universal Display Corp.

57,811

631

-

-

60,096

Veeco Instruments, Inc.

29,617

-

-

-

27,358

Vertex Pharmaceuticals, Inc.

321,122

9,702

-

-

231,005

Total

$ 9,357,132

$ 762,224

$ 749,944

$ 491

$ 6,675,534

Other Information

The following is a summary of the inputs used, as of February 29, 2008, involving the Fund's assets carried at value. The inputs or methodology used for valuing securities may not be an indication of the risk associated with investing in those securities.

Valuation Inputs at Reporting Date:

Description
(Amounts in thousands)

Total

Level 1

Level 2

Level 3

Investments in Securities

$ 34,138,030

$ 31,668,983

$ 2,443,406

$ 25,641

The following is a reconciliation of assets for which Level 3 inputs were used in determining value:

(Amounts in thousands)

Investments in Securities

Beginning Balance

$ 25,641

Total Realized Gain (Loss)

-

Total Unrealized Gain (Loss)

-

Cost of Purchases

-

Proceeds of Sales

-

Amortization/Accretion

-

Transfer in/out of Level 3

-

Ending Balance

$ 25,641

The information used in the above reconciliation represents fiscal year to date activity for any Investment Securities identified as using Level 3 inputs at either the beginning or the end of the current fiscal period. Transfers in or out of Level 3 represents either the beginning value (for transfers in), or the ending value (for transfers out) of any Security or Instrument where a change in the pricing level occured from the beginning to the end of the period.

Income Tax Information

At February 29, 2008, the aggregate cost of investment securities for income tax purposes was $28,555,745,000. Net unrealized appreciation aggregated $5,582,285,000, of which $8,961,313,000 related to appreciated investment securities and $3,379,028,000 related to depreciated investment securities.

Investment Valuation

Investments are valued as of the close of business of the New York Stock Exchange (NYSE), normally 4:00 p.m. Eastern time. Security transactions are accounted for as of trade date. Wherever possible, the Fund uses independent pricing services approved by the Board of Trustees to value its investments.

Equity securities, including restricted securities, for which market quotations are readily available, are valued at the last reported sale price or official closing price as reported by an independent pricing service on the primary market or exchange on which they are traded. In the event there were no sales during the day or closing prices are not available, securities are valued at the last quoted bid price. Investments in open-end mutual funds, including the Fidelity Central Funds, are valued at their closing net asset value each business day. Short-term securities with remaining maturities of sixty days or less for which quotations are not readily available are valued at amortized cost, which approximates value.

When current market prices or quotations are not readily available or reliable, valuations may be determined in good faith in accordance with procedures adopted by the Board of Trustees. The frequency with which these procedures are used cannot be predicted and may be utilized to a significant extent. The value of securities used for NAV calculation under these procedures may differ from published prices for the same securities.

The Fund adopted the provisions of Statement of Financial Accounting Standards No. 157, "Fair Value Measurements" (SFAS 157), effective with the beginning of the Fund's fiscal year. SFAS 157 establishes a hierarchy that prioritizes the inputs to valuation techniques giving the highest priority to readily available unadjusted quoted prices in active markets for identical assets (level 1 measurements) and the lowest priority to unobservable inputs (level 3 measurements) when market prices are not readily available or reliable. The three levels of the hierarchy under SFAS 157 are described below:

Level 1 - Quoted prices in active markets for identical securities.

Level 2 - Prices determined using other significant observable inputs. Observable inputs are inputs that other market participants would use in pricing a security. These may include quoted prices for similar securities, interest rates, prepayment speeds, credit risk and others.

Level 3 - Prices determined using significant unobservable inputs. In situations where quoted prices or observable inputs are unavailable (for example, when there is little or no market activity for an investment at the end of the period), unobservable inputs may be used. Unobservable inputs reflect the Fund's own assumptions about the factors market participants would use in pricing an investment, and would be based on the best information available.

Changes in valuation techniques may result in transfers in or out of an investment's assigned level within the hierarchy.

For additional information on the fund's policy regarding valuation of investments and other significant accounting policies, please refer to the fund's most recent semiannual or annual shareholder report.

Quarterly Report

The fund's schedule of investments as of the date on the cover of this report has not been audited. This report is provided for the general information of the fund's shareholders. For more information regarding the fund and its holdings, please see the fund's most recent prospectus and annual report.

Third party trademarks and service marks are the property of their respective owners. All other trademarks and service marks are the property of FMR LLC or an affiliate.

Quarterly Report

Quarterly Holdings Report

for

Fidelity ® New Millennium Fund®

February 29, 2008

1.797943.104

NMF-QTLY-0408

Investments February 29, 2008 (Unaudited)

Showing Percentage of Net Assets

Common Stocks - 96.6%

Shares

Value (000s)

CONSUMER DISCRETIONARY - 11.0%

Auto Components - 0.5%

Gentex Corp.

165,300

$ 2,665

Johnson Controls, Inc.

250,000

8,215

10,880

Automobiles - 0.3%

Fiat SpA

133,400

2,817

Renault SA

30,900

3,301

6,118

Distributors - 1.9%

Li & Fung Ltd.

9,483,600

34,495

LKQ Corp. (a)

131,600

2,795

37,290

Diversified Consumer Services - 0.3%

New Oriental Education & Technology Group, Inc. sponsored ADR (a)

94,100

5,867

Hotels, Restaurants & Leisure - 1.0%

Ctrip.com International Ltd. sponsored ADR

139,500

8,456

Famous Dave's of America, Inc. (a)

230,000

2,335

Life Time Fitness, Inc. (a)(d)

65,500

1,903

Red Robin Gourmet Burgers, Inc. (a)(d)

232,500

7,749

20,443

Household Durables - 0.7%

D.R. Horton, Inc.

304,200

4,268

Ryland Group, Inc.

91,600

2,591

Whirlpool Corp.

74,300

6,269

13,128

Internet & Catalog Retail - 0.1%

NutriSystem, Inc. (a)(d)

186,800

2,673

Leisure Equipment & Products - 0.6%

MarineMax, Inc. (a)(d)

298,900

3,670

Pool Corp. (d)

249,500

4,753

Summer Infant, Inc. (a)(e)

910,784

3,825

12,248

Media - 1.6%

Central European Media Enterprises Ltd. Class A (a)

70,400

6,471

Lee Enterprises, Inc. (d)

202,600

2,089

Sports Properties Acquisition Corp. unit

315,300

3,121

Common Stocks - continued

Shares

Value (000s)

CONSUMER DISCRETIONARY - continued

Media - continued

The Walt Disney Co.

280,800

$ 9,101

Time Warner, Inc.

675,000

10,537

31,319

Multiline Retail - 0.4%

Kohl's Corp. (a)

68,900

3,062

Nordstrom, Inc. (d)

138,000

5,110

8,172

Specialty Retail - 1.2%

Lowe's Companies, Inc.

355,000

8,509

OfficeMax, Inc.

131,400

2,795

PetSmart, Inc.

132,100

2,844

TJX Companies, Inc.

250,900

8,029

Zumiez, Inc. (a)(d)

163,100

2,867

25,044

Textiles, Apparel & Luxury Goods - 2.4%

Carter's, Inc. (a)

431,800

6,663

Crocs, Inc. (a)(d)

168,200

4,091

LVMH Moet Hennessy - Louis Vuitton

55,000

5,650

Polo Ralph Lauren Corp. Class A

56,500

3,514

Ports Design Ltd.

2,390,100

7,066

Under Armour, Inc. Class A (sub. vtg.) (a)(d)

564,200

20,774

47,758

TOTAL CONSUMER DISCRETIONARY

220,940

CONSUMER STAPLES - 7.1%

Beverages - 1.0%

Coca-Cola Femsa SA de CV sponsored ADR

130,100

6,822

Diageo PLC sponsored ADR

98,700

8,103

Molson Coors Brewing Co. Class B

99,300

5,358

20,283

Food & Staples Retailing - 1.9%

Costco Wholesale Corp.

49,100

3,040

CVS Caremark Corp.

159,300

6,433

Sysco Corp.

263,200

7,385

United Natural Foods, Inc. (a)

515,600

8,724

Whole Foods Market, Inc. (d)

229,700

8,074

X5 Retail Group NV GDR (a)(f)

112,500

3,791

37,447

Common Stocks - continued

Shares

Value (000s)

CONSUMER STAPLES - continued

Food Products - 3.3%

Bunge Ltd.

57,300

$ 6,351

Corn Products International, Inc.

153,000

5,617

Green Mountain Coffee Roasters, Inc. (a)(d)

1,145,700

34,955

Hain Celestial Group, Inc. (a)

329,600

8,899

Marine Harvest ASA (a)

4,107,000

2,433

Tyson Foods, Inc. Class A

276,500

3,984

Wimm-Bill-Dann Foods OJSC sponsored ADR

40,000

4,210

66,449

Household Products - 0.5%

Procter & Gamble Co.

158,000

10,456

Personal Products - 0.4%

Avon Products, Inc.

179,700

6,839

Physicians Formula Holdings, Inc. (a)

166,100

1,309

8,148

TOTAL CONSUMER STAPLES

142,783

ENERGY - 10.1%

Energy Equipment & Services - 0.9%

ION Geophysical Corp. (a)(d)

352,800

4,689

Nabors Industries Ltd. (a)

162,200

5,114

North American Energy Partners, Inc. (a)

251,400

4,168

SBM Offshore NV

142,996

4,610

18,581

Oil, Gas & Consumable Fuels - 9.2%

Canadian Natural Resources Ltd.

46,600

3,493

Denbury Resources, Inc. (a)

230,600

7,354

Enbridge, Inc.

118,500

4,883

Energy Transfer Equity LP

161,300

5,365

EOG Resources, Inc.

102,600

12,208

Exxon Mobil Corp.

243,800

21,213

Hess Corp.

67,200

6,262

Lukoil Oil Co. sponsored ADR

77,100

5,721

Niko Resources Ltd.

60,100

5,651

Noble Energy, Inc.

183,500

14,203

OAO Gazprom sponsored ADR

149,500

7,481

OJSC Rosneft unit

765,700

6,263

Petroleo Brasileiro SA - Petrobras sponsored ADR

112,400

13,189

Plains Exploration & Production Co. (a)

175,695

9,488

Quicksilver Resources, Inc. (a)

360,600

12,405

Common Stocks - continued

Shares

Value (000s)

ENERGY - continued

Oil, Gas & Consumable Fuels - continued

SouthGobi Energy Resources Ltd. (a)

607,700

$ 7,960

Southwestern Energy Co. (a)

124,500

8,121

Suncor Energy, Inc.

46,200

4,765

Ultra Petroleum Corp. (a)

135,200

10,609

Williams Companies, Inc.

161,400

5,814

XTO Energy, Inc.

176,250

10,876

183,324

TOTAL ENERGY

201,905

FINANCIALS - 10.6%

Capital Markets - 1.7%

Charles Schwab Corp.

1,002,900

19,667

EFG International

113,920

3,828

GFI Group, Inc. (a)(d)

39,100

2,993

T. Rowe Price Group, Inc.

143,000

7,226

33,714

Commercial Banks - 2.4%

Associated Banc-Corp.

127,900

3,187

BOK Financial Corp.

65,600

3,393

Boston Private Financial Holdings, Inc. (d)

268,000

3,690

City National Corp.

83,900

4,300

East West Bancorp, Inc. (d)

27,600

519

First Community Bancorp, California

57,300

1,633

M&T Bank Corp. (d)

159,300

13,075

PNC Financial Services Group, Inc.

128,100

7,869

Seacoast Banking Corp., Florida (d)

50,029

500

UMB Financial Corp.

258,700

9,903

48,069

Diversified Financial Services - 2.8%

Bolsa de Mercadorias & Futuros - BM&F SA

476,400

5,158

Bovespa Holding SA

491,000

7,822

CME Group, Inc.

44,500

22,842

Deutsche Boerse AG

51,000

8,083

Hong Kong Exchanges & Clearing Ltd.

165,000

3,135

NYMEX Holdings, Inc. (d)

92,300

9,118

56,158

Insurance - 3.3%

AFLAC, Inc.

164,800

10,285

Common Stocks - continued

Shares

Value (000s)

FINANCIALS - continued

Insurance - continued

American International Group, Inc.

266,700

$ 12,498

Assured Guaranty Ltd.

170,000

4,361

Berkshire Hathaway, Inc. Class A (a)

177

24,780

IPC Holdings Ltd.

182,200

4,941

The Chubb Corp.

178,600

9,091

65,956

Real Estate Investment Trusts - 0.4%

Thornburg Mortgage, Inc. (SBI) (d)

865,900

7,707

TOTAL FINANCIALS

211,604

HEALTH CARE - 15.3%

Biotechnology - 4.4%

Alexion Pharmaceuticals, Inc. (a)

59,900

3,631

Alnylam Pharmaceuticals, Inc. (a)

266,600

7,571

Amylin Pharmaceuticals, Inc. (a)

136,200

3,605

Celgene Corp. (a)

158,300

8,923

Cepheid, Inc. (a)

230,600

6,383

CSL Ltd.

402,424

13,497

CytRx Corp. (a)

3,063,304

5,514

Diagnocure, Inc. (a)

920,700

2,302

Genentech, Inc. (a)

139,800

10,590

Genomic Health, Inc. (a)(d)

288,300

5,498

Gilead Sciences, Inc. (a)

115,800

5,480

GTx, Inc. (a)

297,068

4,863

Memory Pharmaceuticals Corp. (a)

1,606,400

1,028

Myriad Genetics, Inc. (a)

49,700

1,840

Omrix Biopharmaceuticals, Inc. (a)

61,700

1,490

OREXIGEN Therapeutics, Inc.

226,300

2,931

Vertex Pharmaceuticals, Inc. (a)

134,100

2,347

87,493

Health Care Equipment & Supplies - 3.8%

Becton, Dickinson & Co.

95,400

8,626

C.R. Bard, Inc.

67,500

6,398

Gen-Probe, Inc. (a)

96,100

4,595

Heartware Ltd. (a)

3,210,821

1,380

Hologic, Inc. (a)

136,052

8,205

Immucor, Inc. (a)

124,900

3,722

Insulet Corp.

192,400

3,296

Common Stocks - continued

Shares

Value (000s)

HEALTH CARE - continued

Health Care Equipment & Supplies - continued

IRIS International, Inc. (a)

233,234

$ 2,629

Meridian Bioscience, Inc.

320,995

11,000

Mindray Medical International Ltd. sponsored ADR

224,500

8,239

Quidel Corp. (a)

505,500

8,305

ThermoGenesis Corp. (a)(e)

5,465,243

9,072

75,467

Health Care Providers & Services - 1.6%

Assisted Living Concepts, Inc. Class A (a)

508,300

3,060

athenahealth, Inc. (d)

119,500

3,886

Capital Senior Living Corp. (a)

611,000

4,607

Dialysis Corp. of America (a)

254,254

2,118

Emeritus Corp. (a)

175,208

3,879

Henry Schein, Inc. (a)

123,300

7,376

LHC Group, Inc. (a)(d)

227,133

3,861

VCA Antech, Inc. (a)

74,400

2,389

31,176

Health Care Technology - 0.8%

Cerner Corp. (a)

308,300

13,396

MedAssets, Inc.

202,000

3,668

17,064

Life Sciences Tools & Services - 2.5%

Affymetrix, Inc. (a)

268,500

5,150

Applera Corp. - Celera Genomics Group (a)

364,200

5,048

Covance, Inc. (a)

74,700

6,305

Illumina, Inc. (a)

101,400

7,342

Pharmaceutical Product Development, Inc.

142,100

6,404

QIAGEN NV (a)

740,900

16,292

Techne Corp. (a)

52,100

3,563

50,104

Pharmaceuticals - 2.2%

BioMimetic Therapeutics, Inc. (a)

61,672

847

Elan Corp. PLC sponsored ADR (a)

251,200

5,720

Inspire Pharmaceuticals, Inc. (a)

302,500

1,358

Nexmed, Inc. (a)(e)

6,399,067

9,791

Perrigo Co.

167,700

5,605

Schering-Plough Corp.

207,000

4,492

Teva Pharmaceutical Industries Ltd. sponsored ADR

100,000

4,907

Common Stocks - continued

Shares

Value (000s)

HEALTH CARE - continued

Pharmaceuticals - continued

Wyeth

150,700

$ 6,574

XenoPort, Inc. (a)

112,000

5,731

45,025

TOTAL HEALTH CARE

306,329

INDUSTRIALS - 16.2%

Aerospace & Defense - 2.9%

General Dynamics Corp.

200,300

16,395

Lockheed Martin Corp.

63,000

6,502

Raytheon Co.

197,000

12,773

Rockwell Collins, Inc.

178,300

10,502

Taser International, Inc. (a)(d)

332,800

3,751

United Technologies Corp.

118,100

8,327

58,250

Air Freight & Logistics - 1.5%

C.H. Robinson Worldwide, Inc.

164,700

8,362

Expeditors International of Washington, Inc.

114,200

4,490

Forward Air Corp.

182,700

5,362

Hub Group, Inc. Class A (a)

233,916

7,015

UTI Worldwide, Inc.

268,000

4,497

29,726

Commercial Services & Supplies - 4.1%

China Security & Surveillance Technology, Inc. (a)(d)

120,500

1,800

Clean Harbors, Inc. (a)

294,600

18,124

Corrections Corp. of America (a)

473,700

12,724

Equifax, Inc.

126,600

4,332

Fuel Tech, Inc. (a)(d)

197,387

3,962

Healthcare Services Group, Inc.

566,600

11,207

Monster Worldwide, Inc. (a)(d)

158,600

4,217

PHH Corp. (a)

200,000

4,048

Robert Half International, Inc.

165,500

4,460

Stericycle, Inc. (a)

80,200

4,322

Tetra Tech, Inc. (a)

268,500

5,067

Waste Management, Inc.

227,800

7,479

81,742

Construction & Engineering - 1.0%

Chicago Bridge & Iron Co. NV (NY Shares)

125,300

5,829

Doosan Heavy Industries & Construction Co. Ltd.

30,540

4,411

Common Stocks - continued

Shares

Value (000s)

INDUSTRIALS - continued

Construction & Engineering - continued

Quanta Services, Inc. (a)

255,201

$ 6,094

URS Corp. (a)

77,200

3,110

19,444

Electrical Equipment - 4.5%

Carmanah Technologies Corp. (a)

1,294,800

1,105

Conergy AG (d)

101,996

2,285

Evergreen Solar, Inc. (a)(d)

1,738,759

16,709

First Solar, Inc. (a)

30,800

6,320

JA Solar Holdings Co. Ltd. ADR

210,000

3,001

Orion Energy Systems, Inc.

123,300

1,112

Q-Cells AG (a)(d)

251,200

20,181

Renewable Energy Corp. AS (a)(d)

1,005,500

24,592

SolarWorld AG

170,500

7,731

Sunpower Corp. Class A (a)

51,500

3,385

Suntech Power Holdings Co. Ltd. sponsored ADR (a)

104,800

3,895

90,316

Industrial Conglomerates - 0.5%

3M Co.

131,100

10,278

Machinery - 1.3%

Badger Meter, Inc.

93,900

3,584

ITT Corp.

160,400

9,021

Kadant, Inc. (a)

167,500

4,204

Pentair, Inc.

128,500

4,192

Valmont Industries, Inc.

62,900

5,026

26,027

Road & Rail - 0.4%

J.B. Hunt Transport Services, Inc.

166,000

4,543

Landstar System, Inc.

91,300

4,234

8,777

TOTAL INDUSTRIALS

324,560

INFORMATION TECHNOLOGY - 18.7%

Communications Equipment - 3.9%

Ciena Corp. (a)

224,400

5,796

Cisco Systems, Inc. (a)

1,724,100

42,021

Corning, Inc.

558,800

12,981

Harris Corp.

133,100

6,499

Infinera Corp.

6,800

79

Common Stocks - continued

Shares

Value (000s)

INFORMATION TECHNOLOGY - continued

Communications Equipment - continued

Juniper Networks, Inc. (a)

265,400

$ 7,118

Nokia Corp. sponsored ADR

100,000

3,601

78,095

Computers & Peripherals - 2.2%

Apple, Inc. (a)

84,500

10,564

EMC Corp. (a)

603,500

9,378

Hewlett-Packard Co.

485,700

23,202

43,144

Electronic Equipment & Instruments - 2.0%

Acacia Research Corp. - Acacia Technologies (a)

562,800

3,675

FLIR Systems, Inc. (a)

265,700

7,562

Hon Hai Precision Industry Co. Ltd. (Foxconn)

1,395,080

8,323

Itron, Inc. (a)(d)

112,000

10,677

Motech Industries, Inc.

578,674

3,905

Roth & Rau AG (a)

13,700

3,302

Universal Display Corp. (a)

200,000

3,408

40,852

Internet Software & Services - 3.0%

Akamai Technologies, Inc. (a)

220,800

7,763

Constant Contact, Inc. (d)

205,300

3,654

DealerTrack Holdings, Inc. (a)

127,300

2,606

Equinix, Inc. (a)

104,500

7,247

Google, Inc. Class A (sub. vtg.) (a)

33,200

15,643

Greenfield Online, Inc. (a)

209,100

2,829

Omniture, Inc. (a)

483,690

11,115

Terremark Worldwide, Inc. (a)

779,900

3,900

VistaPrint Ltd. (a)

178,707

5,615

60,372

IT Services - 1.5%

Cognizant Technology Solutions Corp. Class A (a)

157,200

4,749

Fidelity National Information Services, Inc.

103,200

4,282

Hewitt Associates, Inc. Class A (a)

150,800

5,951

Paychex, Inc.

200,000

6,292

The Western Union Co.

413,100

8,592

29,866

Semiconductors & Semiconductor Equipment - 3.9%

Advanced Analog Technology, Inc.

539,000

1,806

Altera Corp.

228,800

3,915

Applied Materials, Inc.

527,000

10,103

Common Stocks - continued

Shares

Value (000s)

INFORMATION TECHNOLOGY - continued

Semiconductors & Semiconductor Equipment - continued

ARM Holdings PLC sponsored ADR

1,409,400

$ 7,597

ASML Holding NV (NY Shares) (a)

247,566

5,961

Global Mixed-mode Technology, Inc.

469,000

2,625

Hittite Microwave Corp. (a)

133,900

4,433

Intel Corp.

818,500

16,329

MEMC Electronic Materials, Inc. (a)

156,600

11,945

Richtek Technology Corp.

512,050

4,518

Siliconware Precision Industries Co. Ltd. sponsored ADR

684,178

5,597

Volterra Semiconductor Corp. (a)

363,600

3,145

77,974

Software - 2.2%

Adobe Systems, Inc. (a)

209,600

7,053

Autonomy Corp. PLC (a)

578,300

10,757

Concur Technologies, Inc. (a)

135,000

3,947

Gameloft (a)

351,800

1,503

Nuance Communications, Inc. (a)

274,700

4,519

Quality Systems, Inc. (d)

167,700

5,464

Salesforce.com, Inc. (a)

187,800

11,215

44,458

TOTAL INFORMATION TECHNOLOGY

374,761

MATERIALS - 4.5%

Chemicals - 1.7%

ADA-ES, Inc. (a)

203,246

2,175

Airgas, Inc.

148,500

7,216

Calgon Carbon Corp. (a)

262,500

4,289

Ecolab, Inc.

214,700

10,046

Monsanto Co.

56,900

6,582

Zoltek Companies, Inc. (a)(d)

117,800

2,695

33,003

Metals & Mining - 2.6%

ArcelorMittal SA (NY Reg.) Class A

105,800

8,043

BHP Billiton PLC

127,000

4,063

Equigold NL

1,882,701

7,982

Goldcorp, Inc.

250,800

10,829

Ivanhoe Mines Ltd. (a)

481,800

6,282

Rio Tinto PLC (Reg.)

71,300

8,101

Common Stocks - continued

Shares

Value (000s)

MATERIALS - continued

Metals & Mining - continued

Searchlight Minerals Corp. (a)

488,161

$ 1,616

Steel Dynamics, Inc.

98,700

5,750

52,666

Paper & Forest Products - 0.2%

Stella-Jones, Inc.

127,800

4,792

TOTAL MATERIALS

90,461

TELECOMMUNICATION SERVICES - 1.4%

Diversified Telecommunication Services - 1.0%

Qwest Communications International, Inc.

804,500

4,344

Time Warner Telecom, Inc. Class A (sub. vtg.) (a)

327,200

5,216

Verizon Communications, Inc.

289,700

10,522

20,082

Wireless Telecommunication Services - 0.4%

America Movil SAB de CV Series L sponsored ADR

119,200

7,207

TOTAL TELECOMMUNICATION SERVICES

27,289

UTILITIES - 1.7%

Electric Utilities - 0.1%

Westar Energy, Inc.

120,050

2,729

Independent Power Producers & Energy Traders - 0.5%

AES Corp. (a)

528,300

9,499

Multi-Utilities - 0.9%

NSTAR

249,400

7,706

Veolia Environnement sponsored ADR

75,200

6,697

YTL Corp. Bhd

993,500

2,371

16,774

Water Utilities - 0.2%

Cascal BV

340,000

4,097

TOTAL UTILITIES

33,099

TOTAL COMMON STOCKS

(Cost $1,819,876)

1,933,731

Money Market Funds - 12.7%

Shares

Value (000s)

Fidelity Cash Central Fund, 3.24% (b)

80,008,380

$ 80,008

Fidelity Securities Lending Cash Central Fund, 3.25% (b)(c)

174,802,741

174,803

TOTAL MONEY MARKET FUNDS

(Cost $254,811)

254,811

TOTAL INVESTMENT PORTFOLIO - 109.3%

(Cost $2,074,687)

2,188,542

NET OTHER ASSETS - (9.3)%

(186,347)

NET ASSETS - 100%

$ 2,002,195

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

(e) Affiliated company

(f) Security exempt from registration under Rule 144A of the Securities Act of 1933. These securities may be resold in transactions exempt from registration, normally to qualified institutional buyers. At the end of the period, the value of these securities amounted to $3,791,000 or 0.2% of net assets.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned
(Amounts in thousands)

Fidelity Cash Central Fund

$ 583

Fidelity Securities Lending Cash Central Fund

902

Total

$ 1,485

Other Affiliated Issuers

An affiliated company is a company in which the fund has ownership of at least 5% of the voting securities. Fiscal year to date transactions with companies which are or were affiliates are as follows:

Affiliate
(Amounts in thousands)

Value, beginning of period

Purchases

Sales Proceeds

Dividend Income

Value,
end of
period

Nexmed, Inc.

$ 8,895

$ -

$ -

$ -

$ 9,791

Summer Infant, Inc.

4,545

-

-

-

3,825

ThermoGenesis Corp.

11,750

-

-

-

9,072

Total

$ 25,190

$ -

$ -

$ -

$ 22,688

Other Information

The following is a summary of the inputs used, as of February 29, 2008, involving the Fund's assets carried at value. The inputs or methodology used for valuing securities may not be an indication of the risk associated with investing in those securities.

Valuation Inputs at Reporting Date:

Description
(Amounts in thousands)

Total

Level 1

Level 2

Level 3

Investments in Securities

$ 2,188,542

$ 1,979,791

$ 208,751

$ 0

Income Tax Information

At February 29, 2008, the aggregate cost of investment securities for income tax purposes was $2,079,827,000. Net unrealized appreciation aggregated $108,715,000, of which $316,829,000 related to appreciated investment securities and $208,114,000 related to depreciated investment securities.

Investment Valuation

Investments are valued as of the close of business of the New York Stock Exchange (NYSE), normally 4:00 p.m. Eastern time. Security transactions are accounted for as of trade date. Wherever possible, the Fund uses independent pricing services approved by the Board of Trustees to value its investments.

Equity securities, including restricted securities, for which market quotations are readily available, are valued at the last reported sale price or official closing price as reported by an independent pricing service on the primary market or exchange on which they are traded. In the event there were no sales during the day or closing prices are not available, securities are valued at the last quoted bid price. Investments in open-end mutual funds, including the Fidelity Central Funds, are valued at their closing net asset value each business day. Short-term securities with remaining maturities of sixty days or less for which quotations are not readily available are valued at amortized cost, which approximates value.

When current market prices or quotations are not readily available or reliable, valuations may be determined in good faith in accordance with procedures adopted by the Board of Trustees. The frequency with which these procedures are used cannot be predicted and may be utilized to a significant extent. The value of securities used for NAV calculation under these procedures may differ from published prices for the same securities.

The Fund adopted the provisions of Statement of Financial Accounting Standards No. 157, "Fair Value Measurements" (SFAS 157), effective with the beginning of the Fund's fiscal year. SFAS 157 establishes a hierarchy that prioritizes the inputs to valuation techniques giving the highest priority to readily available unadjusted quoted prices in active markets for identical assets (level 1 measurements) and the lowest priority to unobservable inputs (level 3 measurements) when market prices are not readily available or reliable. The three levels of the hierarchy under SFAS 157 are described below:

Level 1 - Quoted prices in active markets for identical securities.

Level 2 - Prices determined using other significant observable inputs. Observable inputs are inputs that other market participants would use in pricing a security. These may include quoted prices for similar securities, interest rates, prepayment speeds, credit risk and others.

Level 3 - Prices determined using significant unobservable inputs. In situations where quoted prices or observable inputs are unavailable (for example, when there is little or no market activity for an investment at the end of the period), unobservable inputs may be used. Unobservable inputs reflect the Fund's own assumptions about the factors market participants would use in pricing an investment, and would be based on the best information available.

Changes in valuation techniques may result in transfers in or out of an investment's assigned level within the hierarchy.

For additional information on the fund's policy regarding valuation of investments and other significant accounting policies, please refer to the fund's most recent semiannual or annual shareholder report.

Quarterly Report

The fund's schedule of investments as of the date on the cover of this report has not been audited. This report is provided for the general information of the fund's shareholders. For more information regarding the fund and its holdings, please see the fund's most recent prospectus and annual report.

Third party trademarks and service marks are the property of their respective owners. All other trademarks and service marks are the property of FMR LLC or an affiliate.

Quarterly Report

Item 2. Controls and Procedures

(a)(i) The President and Treasurer and the Chief Financial Officer have concluded that the Fidelity Mt. Vernon Street Trust's (the "Trust") disclosure controls and procedures (as defined in Rule 30a-3(c) under the Investment Company Act) provide reasonable assurances that material information relating to the Trust is made known to them by the appropriate persons, based on their evaluation of these controls and procedures as of a date within 90 days of the filing date of this report.

(a)(ii) There was no change in the Trust's internal control over financial reporting (as defined in Rule 30a-3(d) under the Investment Company Act) that occurred during the Trust's last fiscal quarter that has materially affected, or is reasonably likely to materially affect, the Trust's internal control over financial reporting.

Item 3. Exhibits

Certification pursuant to Rule 30a-2(a) under the Investment Company Act of 1940 (17 CFR 270.30a-2(a)) is filed and attached hereto as Exhibit 99.CERT.

SIGNATURES

Pursuant to the requirements of the Securities Exchange Act of 1934 and the Investment Company Act of 1940, the registrant has duly caused this report to be signed on its behalf by the undersigned, thereunto duly authorized.

Fidelity Mt. Vernon Street Trust

By:

/s/Kimberley Monasterio

Kimberley Monasterio

President and Treasurer

Date:

April 29, 2008

Pursuant to the requirements of the Securities Exchange Act of 1934 and the Investment Company Act of 1940, this report has been signed below by the following persons on behalf of the registrant and in the capacities and on the dates indicated.

By:

/s/ Kimberley Monasterio

Kimberley Monasterio

President and Treasurer

Date:

April 29, 2008

By:

/s/Joseph B. Hollis

Joseph B. Hollis

Chief Financial Officer

Date:

April 29, 2008