N-CSR 1 main.htm

UNITED STATES
SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

FORM N-CSR

CERTIFIED SHAREHOLDER REPORT OF REGISTERED

MANAGEMENT INVESTMENT COMPANIES

Investment Company Act file number 811-1352

Fidelity Devonshire Trust
(Exact name of registrant as specified in charter)

82 Devonshire St., Boston, Massachusetts 02109
(Address of principal executive offices) (Zip code)

Eric D. Roiter, Secretary

82 Devonshire St.

Boston, Massachusetts 02109
(Name and address of agent for service)

Registrant's telephone number, including area code: 617-563-7000

Date of fiscal year end:

January 31

Date of reporting period:

January 31, 2008

Item 1. Reports to Stockholders

Fidelity®

Equity-Income

Fund

Annual Report

January 31, 2008

(2_fidelity_logos) (Registered_Trademark)

Contents

Chairman's Message

<Click Here>

Ned Johnson's message to shareholders.

Performance

<Click Here>

How the fund has done over time.

Management's Discussion

<Click Here>

The manager's review of fund performance, strategy and outlook.

Shareholder Expense Example

<Click Here>

An example of shareholder expenses.

Investment Changes

<Click Here>

A summary of major shifts in the fund's investments over the past six months.

Investments

<Click Here>

A complete list of the fund's investments with their market values.

Financial Statements

<Click Here>

Statements of assets and liabilities, operations, and changes in net assets, as well as financial highlights.

Notes

<Click Here>

Notes to the financial statements.

Report of Independent Registered Public Accounting Firm

<Click Here>

Trustees and Officers

<Click Here>

Distributions

<Click Here>

To view a fund's proxy voting guidelines and proxy voting record for the 12-month period ended June 30, visit http://www.fidelity.com (search for "proxy voting guidelines") or visit the Securities and Exchange Commission's (SEC) web site at http://www.sec.gov. You may also call 1-800-544-8544 to request a free copy of the proxy voting guidelines.

Standard & Poor's, S&P and S&P 500 are registered service marks of The McGraw-Hill Companies, Inc. and have been licensed for use by Fidelity Distributors Corporation.

Other third party marks appearing herein are the property of their respective owners.

All other marks appearing herein are registered or unregistered trademarks or service marks of FMR LLC or an affiliated company.

Annual Report

This report and the financial statements contained herein are submitted for the general information of the shareholders of the fund. This report is not authorized for distribution to prospective investors in the fund unless preceded or accompanied by an effective prospectus.

A fund files its complete schedule of portfolio holdings with the SEC for the first and third quarters of each fiscal year on Form N-Q. Forms N-Q are available on the SEC's web site at http://www.sec.gov. A fund's Forms N-Q listing may be reviewed and copied at the SEC's Public Reference Room in Washington, DC. Information regarding the operation of the SEC's Public Reference Room may be obtained by calling 1-800-SEC-0330. For a complete list of a fund's portfolio holdings, view the most recent holdings listing, semiannual report, or annual report on Fidelity's web site at http://www.fidelity.com or http://www.advisor.fidelity.com, as applicable.

NOT FDIC INSURED · MAY LOSE VALUE · NO BANK GUARANTEE

Neither the fund nor Fidelity Distributors Corporation is a bank.

Annual Report

Chairman's Message

(photo_of_Edward_C_Johnson_3d)

Dear Shareholder:

Stocks got off to a poor start in 2008, while investment-grade bonds and money markets showed positive returns, once again underscoring the importance of a diversified portfolio. Financial markets are always unpredictable, but there are a number of time-tested principles that can put the historical odds in your favor.

One of the basic tenets is to invest for the long term. Over time, riding out the markets' inevitable ups and downs has proven much more effective than selling into panic or chasing the hottest trend. Even missing only a few of the markets' best days can significantly diminish investor returns. Patience also affords the benefits of compounding - of earning interest on additional income or reinvested dividends and capital gains. There are tax advantages and cost benefits to consider as well. The more you sell, the more taxes you pay, and the more you trade, the higher the costs. While staying the course doesn't eliminate risk, it can considerably lessen the effect of short-term declines.

You can further manage your investing risk through diversification. And today, more than ever, geographic diversification should be taken into account. Studies indicate that asset allocation is the single most important determinant of a portfolio's long-term success. The right mix of stocks, bonds and cash - aligned to your particular risk tolerance and investment objective - is very important. Age-appropriate rebalancing is also an essential aspect of asset allocation. For younger investors, an emphasis on equities - which historically have been the best-performing asset class over time - is encouraged. As investors near their specific goal, such as retirement or sending a child to college, consideration may be given to replacing volatile assets (e.g. common stocks) with more-stable fixed investments (bonds or savings plans).

A third investment principle - investing regularly - can help lower the average cost of your purchases. Investing a certain amount of money each month or quarter helps ensure you won't pay for all your shares at market highs. This strategy - known as dollar cost averaging - also reduces unconstructive "emotion" from investing, helping shareholders avoid selling weak performers just prior to an upswing, or chasing a hot performer just before a correction.

We invite you to contact us via the Internet, through our Investor Centers or over the phone. It is our privilege to provide you the information you need to make the investments that are right for you.

Sincerely,

/s/Edward C. Johnson 3d

Edward C. Johnson 3d

Annual Report

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the fund's dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table and the fund's returns do not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Past 10
years

Fidelity® Equity-Income Fund

-5.21%

12.75%

6.17%

$10,000 Over 10 Years

Let's say hypothetically that $10,000 was invested in Fidelity® Equity-Income Fund on January 31, 1998. The chart shows how the value of your investment would have changed, and also shows how the Russell 3000® Value Index performed over the same period.



Annual Report

Management's Discussion of Fund Performance

Comments from Stephen Petersen, Portfolio Manager of Fidelity® Equity-Income Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the 12 months ending January 31, 2008, the fund declined 5.21%. By comparison, the Russell 3000® Value Index was down 6.20%. Underweighting financials - and, in particular, mid-sized banks - versus the Russell had the greatest positive impact on the fund's relative results. In general, the fund's emphasis on larger-cap issues also helped because the index holds more small- and mid-cap stocks, which generally struggled during the past year. Additionally, good stock picks in diversified financials, industrials and, particularly, technology all contributed to results, though overweighting the poor-performing tech sector offset much of the gain. Conversely, our overweighting and poor security selection in consumer discretionary detracted, as did stock selection in health care. Underweighting poor-performing financial giant Citigroup, a casualty of the subprime blowup, helped our results the most relative to the index. Honeywell, a large supplier to the aerospace industry, got a boost from the huge backlog in orders for new planes to be built and delivered over the next several years. Detractors included newspaper company McClatchy, which encountered problems following its 2006 purchase of newspaper publisher Knight Ridder, including a slower-than-expected integration process. Money transfer provider MoneyGram International, an out-of-benchmark holding, also hurt. The company's thriving wire transfer business was offset by losses in its investment loan portfolio, stemming primarily from its subprime mortgage holdings.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Shareholder Expense Example

As a shareholder of the Fund, you incur two types of costs: (1) transaction costs, and (2) ongoing costs, including management fees and other Fund expenses. This Example is intended to help you understand your ongoing costs (in dollars) of investing in the Fund and to compare these costs with the ongoing costs of investing in other mutual funds.

The Example is based on an investment of $1,000 invested at the beginning of the period and held for the entire period (August 1, 2007 to January 31, 2008).

Actual Expenses

The first line of the accompanying table provides information about actual account values and actual expenses. You may use the information in this line, together with the amount you invested, to estimate the expenses that you paid over the period. Simply divide your account value by $1,000.00 (for example, an $8,600 account value divided by $1,000.00 = 8.6), then multiply the result by the number in the first line under the heading entitled "Expenses Paid During Period" to estimate the expenses you paid on your account during this period. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, the Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Hypothetical Example for Comparison Purposes

The second line of the accompanying table provides information about hypothetical account values and hypothetical expenses based on the Fund's actual expense ratio and an assumed rate of return of 5% per year before expenses, which is not the Fund's actual return. The hypothetical account values and expenses may not be used to estimate the actual ending account balance or expenses you paid for the period. You may use this information to compare the ongoing costs of investing in the Fund and other funds. To do so, compare this 5% hypothetical example with the 5% hypothetical examples that appear in the shareholder reports of the other funds. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, the Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Annual Report

Please note that the expenses shown in the table are meant to highlight your ongoing costs only and do not reflect any transaction costs. Therefore, the second line of the table is useful in comparing ongoing costs only, and will not help you determine the relative total costs of owning different funds.


Beginning
Account Value
August 1, 2007


Ending
Account Value
January 31, 2008

Expenses Paid
During Period
*
August 1, 2007 to
January 31, 2008

Actual

$ 1,000.00

$ 920.00

$ 3.24

Hypothetical (5% return per year before expenses)

$ 1,000.00

$ 1,021.83

$ 3.41

* Expenses are equal to the Fund's annualized expense ratio of .67%; multiplied by the average account value over the period, multiplied by 184/365 (to reflect the one-half year period).

Annual Report

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Exxon Mobil Corp.

6.0

5.7

AT&T, Inc.

3.6

4.1

Bank of America Corp.

3.1

3.0

JPMorgan Chase & Co.

2.5

2.0

American International Group, Inc.

2.4

2.3

Pfizer, Inc.

1.9

1.7

General Electric Co.

1.9

1.5

Citigroup, Inc.

1.8

2.2

Chevron Corp.

1.8

1.6

Verizon Communications, Inc.

1.5

1.4

26.5

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Financials

26.0

25.6

Energy

16.9

15.2

Consumer Discretionary

12.2

10.5

Industrials

10.3

10.0

Information Technology

9.4

10.1

Asset Allocation (% of fund's net assets)

As of January 31, 2008 *

As of July 31, 2007 **

Stocks 97.2%

Stocks 96.1%

Bonds 0.1%

Bonds 0.1%

Convertible
Securities 2.2%

Convertible
Securities 1.4%

Short-Term
Investments and
Net Other Assets 0.5%

Short-Term
Investments and
Net Other Assets 2.4%

* Foreign investments

8.4%

** Foreign investments

10.3%

Annual Report

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 97.2%

Shares

Value
(000s)

CONSUMER DISCRETIONARY - 11.6%

Auto Components - 0.4%

American Axle & Manufacturing Holdings, Inc.

1,790,229

$ 38,937

The Goodyear Tire & Rubber Co. (a)

2,331,200

58,676

TRW Automotive Holdings Corp. (a)

999,457

22,918

120,531

Automobiles - 1.2%

Ford Motor Co. (a)

8,909,190

59,157

General Motors Corp. (d)

1,301,300

36,840

Harley-Davidson, Inc.

1,172,400

47,576

Hyundai Motor Co.

197,040

15,323

Monaco Coach Corp.

1,321,209

13,437

Peugeot Citroen SA

465,500

34,465

Renault SA

65,121

7,434

Toyota Motor Corp. sponsored ADR

942,600

102,319

Winnebago Industries, Inc.

881,737

18,569

335,120

Diversified Consumer Services - 0.2%

H&R Block, Inc.

3,049,150

58,757

Hotels, Restaurants & Leisure - 0.4%

McDonald's Corp.

906,100

48,522

Starbucks Corp. (a)

1,625,300

30,734

Wyndham Worldwide Corp.

1,123,260

26,464

105,720

Household Durables - 1.5%

Beazer Homes USA, Inc. (d)

1,655,900

14,439

Black & Decker Corp.

935,400

67,854

Centex Corp.

2,082,900

57,863

KB Home

566,900

15,590

Lennar Corp. Class A (d)

1,913,100

39,410

The Stanley Works

1,196,941

61,475

Whirlpool Corp.

1,833,548

156,053

412,684

Internet & Catalog Retail - 0.2%

Liberty Media Corp. - Interactive Series A (a)

3,683,764

58,609

Leisure Equipment & Products - 0.6%

Brunswick Corp.

3,531,000

67,054

Eastman Kodak Co. (d)

4,283,600

85,372

Polaris Industries, Inc. (d)

521,500

22,654

175,080

Common Stocks - continued

Shares

Value
(000s)

CONSUMER DISCRETIONARY - continued

Media - 3.5%

Belo Corp. Series A

917,600

$ 15,241

Citadel Broadcasting Corp.

4,466,355

6,521

Clear Channel Communications, Inc.

4,659,142

143,082

Comcast Corp. Class A (a)

7,797,855

141,609

E.W. Scripps Co. Class A

1,257,033

51,186

Gannett Co., Inc.

1,424,700

52,714

News Corp. Class B

3,667,900

71,304

The McClatchy Co. Class A (d)

3,694,535

39,790

The New York Times Co. Class A (d)

3,000,891

50,235

The Walt Disney Co.

2,631,310

78,755

Time Warner Cable, Inc. (a)

1,817,495

45,728

Time Warner, Inc.

16,769,290

263,949

Virgin Media, Inc.

1,279,250

21,389

981,503

Multiline Retail - 1.7%

Family Dollar Stores, Inc.

1,922,800

40,436

JCPenney Co., Inc.

972,800

46,120

Kohl's Corp. (a)

2,725,347

124,385

Macy's, Inc.

2,916,000

80,598

Nordstrom, Inc.

935,100

36,375

Sears Holdings Corp. (a)(d)

677,700

74,879

Target Corp.

1,296,500

72,059

Tuesday Morning Corp.

1,513,113

9,139

483,991

Specialty Retail - 1.7%

Advance Auto Parts, Inc.

1,034,700

36,918

AnnTaylor Stores Corp. (a)

919,759

23,132

Chico's FAS, Inc. (a)(d)

2,652,900

28,625

Foot Locker, Inc.

1,954,358

26,755

Home Depot, Inc.

5,961,000

182,824

OfficeMax, Inc.

1,415,127

35,053

RadioShack Corp. (d)

1,609,400

27,923

Staples, Inc.

2,688,155

64,354

Williams-Sonoma, Inc.

2,212,200

59,464

485,048

Textiles, Apparel & Luxury Goods - 0.2%

Liz Claiborne, Inc.

2,144,400

46,941

TOTAL CONSUMER DISCRETIONARY

3,263,984

Common Stocks - continued

Shares

Value
(000s)

CONSUMER STAPLES - 5.3%

Beverages - 0.2%

Heineken NV (Bearer)

1,146,446

$ 63,742

Food & Staples Retailing - 1.8%

CVS Caremark Corp.

2,130,100

83,223

Rite Aid Corp. (a)

7,331,532

21,701

Wal-Mart Stores, Inc.

7,383,000

375,647

Winn-Dixie Stores, Inc. (a)

644,464

11,420

491,991

Food Products - 0.7%

Hershey Co.

2,326,700

84,227

Kraft Foods, Inc. Class A

1,538,140

45,006

Lighthouse Caledonia ASA (d)

308,730

237

Marine Harvest ASA (a)

49,497,600

26,662

Tyson Foods, Inc. Class A

3,053,600

43,514

199,646

Household Products - 0.7%

Energizer Holdings, Inc. (a)

261,400

24,472

Kimberly-Clark Corp.

965,200

63,365

Procter & Gamble Co.

1,808,017

119,239

207,076

Personal Products - 0.9%

Avon Products, Inc.

5,790,200

202,773

Estee Lauder Companies, Inc. Class A

1,104,100

46,593

249,366

Tobacco - 1.0%

Altria Group, Inc.

3,711,295

281,390

TOTAL CONSUMER STAPLES

1,493,211

ENERGY - 16.9%

Energy Equipment & Services - 3.2%

Baker Hughes, Inc.

2,002,400

130,016

Expro International Group PLC

383,300

7,109

Halliburton Co.

3,286,100

109,000

Nabors Industries Ltd. (a)

3,229,551

87,908

Noble Corp.

4,714,844

206,369

Pride International, Inc. (a)

1,100,800

34,906

Schlumberger Ltd. (NY Shares)

4,403,763

332,308

907,616

Common Stocks - continued

Shares

Value
(000s)

ENERGY - continued

Oil, Gas & Consumable Fuels - 13.7%

Apache Corp.

1,430,710

$ 136,547

Boardwalk Pipeline Partners, LP

649,600

20,190

BP PLC sponsored ADR

547,400

34,897

Chevron Corp.

5,922,882

500,484

ConocoPhillips

5,123,800

411,544

CONSOL Energy, Inc.

649,600

47,421

Devon Energy Corp.

211,300

17,956

EOG Resources, Inc.

1,911,600

167,265

Exxon Mobil Corp.

19,577,817

1,691,522

Hess Corp.

1,950,000

177,119

Occidental Petroleum Corp.

2,767,100

187,803

Peabody Energy Corp.

875,439

47,291

Royal Dutch Shell PLC Class A sponsored ADR

596,000

42,560

Spectra Energy Corp.

1,106,050

25,262

Total SA sponsored ADR

2,366,102

172,205

Valero Energy Corp.

1,903,000

112,639

Williams Companies, Inc.

1,365,500

43,655

3,836,360

TOTAL ENERGY

4,743,976

FINANCIALS - 25.3%

Capital Markets - 3.9%

Ares Capital Corp.

1,595,142

22,300

Bank of New York Mellon Corp.

8,332,549

388,547

Goldman Sachs Group, Inc.

618,821

124,241

KKR Private Equity Investors, LP

947,000

15,862

KKR Private Equity Investors, LP Restricted Depositary Units (f)

1,714,600

28,720

Legg Mason, Inc.

963,550

69,376

Lehman Brothers Holdings, Inc.

1,135,343

72,855

Merrill Lynch & Co., Inc.

2,518,300

142,032

Morgan Stanley

3,864,760

191,035

State Street Corp.

607,397

49,879

1,104,847

Commercial Banks - 5.3%

Associated Banc-Corp.

3,328,222

93,789

Barclays PLC Sponsored ADR

1,695,300

63,964

HSBC Holdings PLC sponsored ADR

1,796,761

135,080

KeyCorp

1,723,800

45,077

Lloyds TSB Group PLC

6,659,701

58,163

Common Stocks - continued

Shares

Value
(000s)

FINANCIALS - continued

Commercial Banks - continued

Marshall & Ilsley Corp.

1,385,898

$ 38,667

PNC Financial Services Group, Inc.

3,036,300

199,242

Royal Bank of Scotland Group PLC

4,356,927

33,591

Societe Generale Series A

228,000

28,644

Sterling Financial Corp., Washington

1,774,124

31,562

U.S. Bancorp, Delaware

3,674,302

124,743

Wachovia Corp.

7,163,618

278,880

Wells Fargo & Co.

10,863,436

369,465

1,500,867

Consumer Finance - 0.7%

American Express Co.

1,261,200

62,202

Capital One Financial Corp.

130,000

7,125

Discover Financial Services

6,330,232

110,779

SLM Corp.

195,400

4,250

184,356

Diversified Financial Services - 7.4%

Bank of America Corp.

20,067,245

889,982

Citigroup, Inc.

18,108,885

511,033

JPMorgan Chase & Co.

14,580,849

693,319

2,094,334

Insurance - 5.6%

ACE Ltd.

3,680,901

214,744

Allstate Corp.

887,600

43,732

American International Group, Inc.

12,184,157

672,078

Hartford Financial Services Group, Inc.

2,057,600

166,192

MetLife, Inc. unit

2,020,300

58,851

Montpelier Re Holdings Ltd.

3,835,478

65,740

Paris RE Holdings Ltd.

293,599

6,244

PartnerRe Ltd.

1,199,126

95,067

The Travelers Companies, Inc.

4,471,640

215,086

XL Capital Ltd. Class A

545,651

24,554

1,562,288

Real Estate Investment Trusts - 0.6%

Annaly Capital Management, Inc.

1,500,239

29,585

Developers Diversified Realty Corp.

934,400

38,451

HCP, Inc.

1,961,300

59,643

Senior Housing Properties Trust (SBI)

1,341,468

30,035

157,714

Common Stocks - continued

Shares

Value
(000s)

FINANCIALS - continued

Real Estate Management & Development - 0.3%

CB Richard Ellis Group, Inc. Class A (a)

4,167,751

$ 80,896

Thrifts & Mortgage Finance - 1.5%

Countrywide Financial Corp. (d)

1,988,700

13,841

Fannie Mae

6,590,500

223,154

Freddie Mac

2,986,300

90,754

MGIC Investment Corp.

1,537,100

28,436

New York Community Bancorp, Inc.

2,730,782

50,656

People's United Financial, Inc.

625,000

10,556

417,397

TOTAL FINANCIALS

7,102,699

HEALTH CARE - 6.6%

Biotechnology - 0.3%

Amgen, Inc. (a)

2,078,778

96,850

Health Care Equipment & Supplies - 0.6%

Covidien Ltd.

2,765,190

123,410

Medtronic, Inc.

921,300

42,905

166,315

Health Care Providers & Services - 0.2%

UnitedHealth Group, Inc.

1,075,466

54,677

Pharmaceuticals - 5.5%

Bristol-Myers Squibb Co.

4,530,800

105,069

Johnson & Johnson

4,547,700

287,688

Merck & Co., Inc.

4,941,200

228,679

Pfizer, Inc.

23,034,900

538,786

Schering-Plough Corp.

9,030,740

176,732

Wyeth

5,474,900

217,901

1,554,855

TOTAL HEALTH CARE

1,872,697

INDUSTRIALS - 10.2%

Aerospace & Defense - 3.0%

General Dynamics Corp.

756,000

63,852

Honeywell International, Inc.

5,506,950

325,296

Lockheed Martin Corp.

1,303,600

140,685

Northrop Grumman Corp.

521,200

41,362

Common Stocks - continued

Shares

Value
(000s)

INDUSTRIALS - continued

Aerospace & Defense - continued

The Boeing Co.

814,100

$ 67,717

United Technologies Corp.

2,778,460

203,967

842,879

Airlines - 0.2%

AMR Corp. (a)

1,601,500

22,325

Delta Air Lines, Inc. (a)

981,900

16,525

US Airways Group, Inc. (a)

309,300

4,271

43,121

Building Products - 0.3%

Masco Corp.

3,455,947

79,245

Commercial Services & Supplies - 0.4%

Cintas Corp.

817,200

26,821

Equifax, Inc.

931,619

34,554

Waste Management, Inc.

1,658,500

53,802

115,177

Electrical Equipment - 0.2%

Emerson Electric Co.

1,202,702

61,145

Industrial Conglomerates - 2.9%

3M Co.

2,445,470

194,782

General Electric Co.

14,980,850

530,472

Tyco International Ltd.

2,563,490

100,899

826,153

Machinery - 2.1%

Briggs & Stratton Corp. (e)

2,945,485

61,413

Caterpillar, Inc.

813,100

57,844

Dover Corp.

2,531,334

102,165

Eaton Corp.

599,400

49,606

Illinois Tool Works, Inc.

853,100

42,996

Ingersoll-Rand Co. Ltd. Class A

2,051,792

81,087

SPX Corp.

1,873,985

188,523

583,634

Marine - 0.1%

Alexander & Baldwin, Inc.

551,486

25,170

Road & Rail - 1.0%

Burlington Northern Santa Fe Corp.

1,708,900

147,854

Common Stocks - continued

Shares

Value
(000s)

INDUSTRIALS - continued

Road & Rail - continued

Ryder System, Inc.

1,306,600

$ 68,022

Union Pacific Corp.

599,500

74,955

290,831

TOTAL INDUSTRIALS

2,867,355

INFORMATION TECHNOLOGY - 9.3%

Communications Equipment - 1.2%

Alcatel-Lucent SA sponsored ADR

7,162,299

45,337

Cisco Systems, Inc. (a)

5,391,900

132,102

Harris Corp.

1,464,600

80,099

Motorola, Inc.

7,393,470

85,247

342,785

Computers & Peripherals - 2.7%

EMC Corp. (a)

6,091,100

96,666

Hewlett-Packard Co.

6,818,461

298,308

International Business Machines Corp.

3,058,300

328,278

Sun Microsystems, Inc. (a)

1,980,150

34,653

757,905

Electronic Equipment & Instruments - 1.0%

Agilent Technologies, Inc. (a)

2,051,900

69,580

Arrow Electronics, Inc. (a)

1,876,900

64,228

Avnet, Inc. (a)

1,895,215

67,489

Tyco Electronics Ltd.

2,765,190

93,491

294,788

Internet Software & Services - 0.2%

Google, Inc. Class A (sub. vtg.) (a)

89,420

50,460

IT Services - 0.6%

Electronic Data Systems Corp.

2,052,600

41,257

Metavante Holding Co. (a)

461,966

10,233

MoneyGram International, Inc.

2,437,904

13,018

The Western Union Co.

3,056,800

68,472

Unisys Corp. (a)

5,456,100

22,697

155,677

Office Electronics - 0.3%

Xerox Corp.

5,237,398

80,656

Semiconductors & Semiconductor Equipment - 2.5%

Analog Devices, Inc.

3,045,200

86,362

Applied Materials, Inc.

6,382,900

114,382

Common Stocks - continued

Shares

Value
(000s)

INFORMATION TECHNOLOGY - continued

Semiconductors & Semiconductor Equipment - continued

Atmel Corp. (a)

2,613,756

$ 8,259

Intel Corp.

12,248,560

259,669

Micron Technology, Inc. (a)(d)

2,748,200

19,320

National Semiconductor Corp.

5,507,364

101,501

Novellus Systems, Inc. (a)

1,306,800

31,050

Teradyne, Inc. (a)

5,676,700

62,273

Varian Semiconductor Equipment Associates, Inc. (a)

911,800

29,369

712,185

Software - 0.8%

Microsoft Corp.

4,747,687

154,775

Symantec Corp. (a)

3,608,900

64,708

219,483

TOTAL INFORMATION TECHNOLOGY

2,613,939

MATERIALS - 2.6%

Chemicals - 1.5%

Arkema sponsored ADR (a)

582,570

32,944

Celanese Corp. Class A

1,646,500

61,217

Chemtura Corp.

7,174,364

48,068

Dow Chemical Co.

1,748,000

67,578

E.I. du Pont de Nemours & Co.

1,040,800

47,023

Georgia Gulf Corp. (d)(e)

2,149,060

16,763

H.B. Fuller Co.

1,053,558

21,872

Hercules, Inc.

1,637,959

28,713

Linde AG

503,879

65,759

PolyOne Corp. (a)

2,892,376

17,817

407,754

Containers & Packaging - 0.2%

Smurfit-Stone Container Corp. (a)

7,180,121

68,139

Metals & Mining - 0.7%

Alcoa, Inc.

4,681,121

154,945

Century Aluminum Co. (a)

437,000

22,720

Nucor Corp.

390,800

22,588

200,253

Common Stocks - continued

Shares

Value
(000s)

MATERIALS - continued

Paper & Forest Products - 0.2%

Glatfelter

993,529

$ 14,366

Weyerhaeuser Co.

502,300

34,016

48,382

TOTAL MATERIALS

724,528

TELECOMMUNICATION SERVICES - 6.3%

Diversified Telecommunication Services - 5.6%

AT&T, Inc.

26,272,950

1,011,246

Qwest Communications International, Inc.

16,770,500

98,611

Telkom SA Ltd. sponsored ADR

737,959

56,631

Verizon Communications, Inc.

10,728,949

416,712

1,583,200

Wireless Telecommunication Services - 0.7%

Sprint Nextel Corp.

9,275,300

97,669

Vodafone Group PLC sponsored ADR

3,098,925

107,843

205,512

TOTAL TELECOMMUNICATION SERVICES

1,788,712

UTILITIES - 3.1%

Electric Utilities - 1.5%

Allegheny Energy, Inc.

1,361,807

74,613

Entergy Corp.

1,590,600

172,071

Exelon Corp.

1,700,700

129,576

PPL Corp.

1,224,100

59,883

436,143

Independent Power Producers & Energy Traders - 0.4%

AES Corp. (a)

5,213,721

99,478

Multi-Utilities - 1.2%

Public Service Enterprise Group, Inc.

2,310,200

221,779

Wisconsin Energy Corp.

2,323,900

105,807

327,586

TOTAL UTILITIES

863,207

TOTAL COMMON STOCKS

(Cost $21,989,460)

27,334,308

Convertible Preferred Stocks - 1.4%

Shares

Value
(000s)

CONSUMER DISCRETIONARY - 0.2%

Automobiles - 0.1%

General Motors Corp.:

Series B, 5.25%

1,122,600

$ 22,295

Series C, 6.25%

781,200

16,194

38,489

Hotels, Restaurants & Leisure - 0.1%

Six Flags, Inc. 7.25% PIERS

1,101,300

14,317

TOTAL CONSUMER DISCRETIONARY

52,806

FINANCIALS - 0.7%

Diversified Financial Services - 0.5%

Bank of America Corp. Series L, 7.25%

48,800

54,283

CIT Group, Inc. 7.75%

1,105,800

22,868

Citigroup, Inc. Series T, 6.50%

1,181,898

63,929

141,080

Thrifts & Mortgage Finance - 0.2%

Washington Mutual, Inc. Series R, 7.75%

32,600

37,971

TOTAL FINANCIALS

179,051

HEALTH CARE - 0.2%

Pharmaceuticals - 0.2%

Schering-Plough Corp. 6.00%

260,400

49,486

MATERIALS - 0.3%

Chemicals - 0.0%

Celanese Corp. 4.25%

157,200

7,579

Metals & Mining - 0.3%

Freeport-McMoRan Copper & Gold, Inc. 6.75%

651,000

85,887

TOTAL MATERIALS

93,466

TELECOMMUNICATION SERVICES - 0.0%

Diversified Telecommunication Services - 0.0%

Cincinnati Bell, Inc. Series B, 6.75%

15,600

652

TOTAL CONVERTIBLE PREFERRED STOCKS

(Cost $381,161)

375,461

Corporate Bonds - 0.9%

Principal Amount
(000s)

Value
(000s)

Convertible Bonds - 0.8%

CONSUMER DISCRETIONARY - 0.4%

Automobiles - 0.1%

Ford Motor Co. 4.25% 12/15/36

$ 28,010

$ 27,724

Hotels, Restaurants & Leisure - 0.0%

Six Flags, Inc. 4.5% 5/15/15

9,950

6,269

Media - 0.3%

Liberty Media Corp.:

3.5% 1/15/31

7,175

6,247

4% 11/15/29 (f)

8,890

5,725

3.5% 1/15/31 (f)

20,342

17,711

News America, Inc. liquid yield option note:

0% 2/28/21 (f)

57,550

34,170

0% 2/28/21

7,600

4,513

68,366

TOTAL CONSUMER DISCRETIONARY

102,359

INDUSTRIALS - 0.1%

Airlines - 0.1%

UAL Corp.:

4.5% 6/30/21 (f)

20,550

27,110

4.5% 6/30/21

3,320

4,380

31,490

INFORMATION TECHNOLOGY - 0.1%

Semiconductors & Semiconductor Equipment - 0.1%

Advanced Micro Devices, Inc.:

6% 5/1/15 (f)

39,060

28,221

6% 5/1/15

13,020

9,407

37,628

TELECOMMUNICATION SERVICES - 0.2%

Diversified Telecommunication Services - 0.2%

Level 3 Communications, Inc.:

3.5% 6/15/12

17,200

15,599

5.25% 12/15/11 (f)

28,080

29,523

5.25% 12/15/11

15,880

16,696

61,818

TOTAL CONVERTIBLE BONDS

233,295

Corporate Bonds - continued

Principal Amount
(000s)

Value
(000s)

Nonconvertible Bonds - 0.1%

MATERIALS - 0.1%

Chemicals - 0.1%

Hercules, Inc. 6.5% 6/30/29 unit

$ 31,600

$ 26,706

TOTAL CORPORATE BONDS

(Cost $282,581)

260,001

Money Market Funds - 1.1%

Shares

Fidelity Cash Central Fund, 3.79% (b)

116,488,509

116,489

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

188,505,785

188,506

TOTAL MONEY MARKET FUNDS

(Cost $304,995)

304,995

TOTAL INVESTMENT PORTFOLIO - 100.6%

(Cost $22,958,197)

28,274,765

NET OTHER ASSETS - (0.6)%

(159,978)

NET ASSETS - 100%

$ 28,114,787

Security Type Abbreviations

PIERS -

Preferred Income Equity Redeemable Securities

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

(e) Affiliated company

(f) Security exempt from registration under Rule 144A of the Securities Act of 1933. These securities may be resold in transactions exempt from registration, normally to qualified institutional buyers. At the end of the period, the value of these securities amounted to $171,180,000 or 0.6% of net assets.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned
(Amounts in thousands)

Fidelity Cash Central Fund

$ 32,962

Fidelity Securities Lending Cash Central Fund

4,263

Total

$ 37,225

Other Affiliated Issuers

An affiliated company is a company in which the fund has ownership of at least 5% of the voting securities. Fiscal year to date transactions with companies which are or were affiliates are as follows:

Affiliates
(Amounts in thousands)

Value, beginning of period

Purchases

Sales Proceeds

Dividend Income

Value, end of period

Briggs & Stratton Corp.

$ 71,205

$ 15,116

$ -

$ 2,305

$ 61,413

Georgia Gulf Corp.

39,180

3,877

-

623

16,763

Total

$ 110,385

$ 18,993

$ -

$ 2,928

$ 78,176

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Statements

Statement of Assets and Liabilities

Amounts in thousands (except per-share amount)

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $192,248) - See accompanying schedule:

Unaffiliated issuers (cost $22,493,820)

$ 27,891,594

Fidelity Central Funds (cost $304,995)

304,995

Other affiliated issuers (cost $159,382)

78,176

Total Investments (cost $22,958,197)

$ 28,274,765

Foreign currency held at value (cost $202)

202

Receivable for investments sold

108,926

Receivable for fund shares sold

29,659

Dividends receivable

41,815

Interest receivable

1,753

Distributions receivable from Fidelity Central Funds

596

Prepaid expenses

94

Other receivables

664

Total assets

28,458,474

Liabilities

Payable for investments purchased

$ 110,800

Payable for fund shares redeemed

27,450

Accrued management fee

10,541

Other affiliated payables

5,162

Other payables and accrued expenses

1,228

Collateral on securities loaned, at value

188,506

Total liabilities

343,687

Net Assets

$ 28,114,787

Net Assets consist of:

Paid in capital

$ 22,165,178

Undistributed net investment income

31,169

Accumulated undistributed net realized gain (loss) on investments and foreign currency transactions

601,855

Net unrealized appreciation (depreciation) on investments and assets and liabilities in foreign currencies

5,316,585

Net Assets, for 538,038 shares outstanding

$ 28,114,787

Net Asset Value, offering price and redemption price per share ($28,114,787 ÷ 538,038 shares)

$ 52.25

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Statements - continued

Statement of Operations

Amounts in thousands

Year ended January 31, 2008

Investment Income

Dividends (including $2,928 earned from other
affiliated issuers)

$ 691,077

Interest

12,327

Income from Fidelity Central Funds

37,225

Total income

740,629

Expenses

Management fee

$ 145,120

Transfer agent fees

61,274

Accounting and security lending fees

2,063

Custodian fees and expenses

489

Independent trustees' compensation

115

Registration fees

60

Audit

282

Legal

169

Interest

36

Miscellaneous

987

Total expenses before reductions

210,595

Expense reductions

(2,625)

207,970

Net investment income (loss)

532,659

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

1,809,320

Foreign currency transactions

112

Total net realized gain (loss)

1,809,432

Change in net unrealized appreciation (depreciation) on:

Investment securities

(3,847,126)

Assets and liabilities in foreign currencies

90

Total change in net unrealized appreciation (depreciation)

(3,847,036)

Net gain (loss)

(2,037,604)

Net increase (decrease) in net assets resulting from operations

$ (1,504,945)

See accompanying notes which are an integral part of the financial statements.

Annual Report

Statement of Changes in Net Assets

Amounts in thousands

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ 532,659

$ 469,920

Net realized gain (loss)

1,809,432

1,948,593

Change in net unrealized appreciation (depreciation)

(3,847,036)

2,155,906

Net increase (decrease) in net assets resulting
from operations

(1,504,945)

4,574,419

Distributions to shareholders from net investment income

(540,755)

(456,678)

Distributions to shareholders from net realized gain

(1,682,275)

(1,722,300)

Total distributions

(2,223,030)

(2,178,978)

Share transactions
Proceeds from sales of shares

6,781,847

5,873,092

Reinvestment of distributions

2,173,858

2,127,167

Cost of shares redeemed

(8,335,462)

(5,215,678)

Net increase (decrease) in net assets resulting from share transactions

620,243

2,784,581

Total increase (decrease) in net assets

(3,107,732)

5,180,022

Net Assets

Beginning of period

31,222,519

26,042,497

End of period (including undistributed net investment income of $31,169 and undistributed net investment income of $36,873, respectively)

$ 28,114,787

$ 31,222,519

Other Information

Shares

Sold

115,779

103,776

Issued in reinvestment of distributions

38,488

37,508

Redeemed

(142,454)

(92,804)

Net increase (decrease)

11,813

48,480

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights

Years ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 59.33

$ 54.51

$ 51.52

$ 50.27

$ 38.32

Income from Investment Operations

Net investment income (loss) B

1.00

.96

.82

.79

.71

Net realized and unrealized gain (loss)

(3.86)

8.30

5.14

2.93

12.88

Total from investment operations

(2.86)

9.26

5.96

3.72

13.59

Distributions from net investment income

(1.02)

(.94)

(.84)

(.81)

(.71)

Distributions from net realized gain

(3.20)

(3.50)

(2.13)

(1.66)

(.93)

Total distributions

(4.22)

(4.44)

(2.97)

(2.47)

(1.64)

Net asset value, end of period

$ 52.25

$ 59.33

$ 54.51

$ 51.52

$ 50.27

Total Return A

(5.21)%

17.55%

11.87%

7.51%

35.95%

Ratios to Average Net Assets C, E

Expenses before reductions

.66%

.68%

.69%

.70%

.71%

Expenses net of fee waivers, if any

.66%

.68%

.69%

.70%

.71%

Expenses net of all reductions

.66%

.67%

.67%

.69%

.70%

Net investment income (loss)

1.68%

1.71%

1.57%

1.56%

1.63%

Supplemental Data

Net assets, end of period
(in millions)

$ 28,115

$ 31,223

$ 26,042

$ 25,730

$ 23,693

Portfolio turnover rate D

23%

24%

19%

19%

25%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Expense ratios reflect operating expenses of the Fund. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the Fund during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the Fund.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Notes to Financial Statements

For the period ended January 31, 2008

(Amounts in thousands except ratios)

1. Organization.

Fidelity Equity-Income Fund (the Fund) is a fund of Fidelity Devonshire Trust (the trust) and is authorized to issue an unlimited number of shares. The trust is registered under the Investment Company Act of 1940, as amended (the 1940 Act), as an open-end management investment company organized as a Massachusetts business trust.

2. Investments in Fidelity Central Funds.

The Fund may invest in Fidelity Central Funds, which are open-end investment companies available only to other investment companies and accounts managed by Fidelity Management & Research Company (FMR) and its affiliates. The Fund's Schedule of Investments lists each of the Fidelity Central Funds as an investment of the Fund, but does not include the underlying holdings of each Fidelity Central Fund. As an Investing Fund, the Fund indirectly bears its proportionate share of the expenses of the underlying Fidelity Central Funds.

The Money Market Central Funds seek preservation of capital and current income and are managed by Fidelity Investments Money Management, Inc. (FIMM), an affiliate of FMR.

A complete list of holdings for each Fidelity Central Fund is available upon request or at the SEC's web site at www.sec.gov. In addition, the financial statements of the Fidelity Central Funds, which are not covered by the Fund's Report of Independent Registered Public Accounting Firm, are available on the SEC's web site or upon request.

3. Significant Accounting Policies.

The financial statements have been prepared in conformity with accounting principles generally accepted in the United States of America, which require management to make certain estimates and assumptions at the date of the financial statements. Actual results could differ from those estimates. The following summarizes the significant accounting policies of the Fund:

Security Valuation. Investments are valued and net asset value (NAV) per share is calculated (NAV calculation) as of the close of business of the New York Stock Exchange (NYSE), normally 4:00 p.m. Eastern time. Wherever possible, the Fund uses independent pricing services approved by the Board of Trustees to value its investments.

Equity securities, including restricted securities, for which market quotations are readily available, are valued at the last reported sale price or official closing price as reported by an independent pricing service on the primary market or exchange on which they are traded. In the event there were no sales during the day or closing prices are not available, securities are valued at the last quoted bid price. Debt securities, including restricted securities, for which quotations are readily available, are valued by independent pricing services or by dealers who make markets in such securities. Pricing services consider

Annual Report

Notes to Financial Statements - continued

(Amounts in thousands except ratios)

3. Significant Accounting Policies - continued

Security Valuation - continued

yield or price of bonds of comparable quality, coupon, maturity and type as well as available dealer supplied prices. Investments in open-end mutual funds, including the Fidelity Central Funds, are valued at their closing net asset value each business day. Short-term securities with remaining maturities of sixty days or less for which quotations are not readily available are valued at amortized cost, which approximates value.

When current market prices or quotations are not readily available or do not accurately reflect fair value, valuations may be determined in accordance with procedures adopted by the Board of Trustees. For example, when developments occur between the close of a market and the close of the NYSE that may materially affect the value of some or all of the securities, or when trading in a security is halted, those securities may be fair valued. Factors used in the determination of fair value may include monitoring news to identify significant market or security specific events such as changes in the value of U.S. securities markets, reviewing developments in foreign markets and evaluating the performance of ADRs, futures contracts and exchange-traded funds. Because the Fund's utilization of fair value pricing depends on market activity, the frequency with which fair value pricing is used cannot be predicted and may be utilized to a significant extent. The value of securities used for NAV calculation under fair value pricing may differ from published prices for the same securities.

Foreign Currency. The Fund uses foreign currency contracts to facilitate transactions in foreign-denominated securities. Losses from these transactions may arise from changes in the value of the foreign currency or if the counterparties do not perform under the contracts' terms.

Foreign-denominated assets, including investment securities, and liabilities are translated into U.S. dollars at the exchange rate at period end. Purchases and sales of investment securities, income and dividends received and expenses denominated in foreign currencies are translated into U.S. dollars at the exchange rate in effect on the transaction date.

The effects of exchange rate fluctuations on investments are included with the net realized and unrealized gain (loss) on investment securities. Other foreign currency transactions resulting in realized and unrealized gain (loss) are disclosed separately.

Investment Transactions and Income. For financial reporting purposes, the Fund's investment holdings and NAV include trades executed through the end of the last business day of the period. The NAV for processing shareholder transactions includes trades executed through the end of the prior business day. Gains and losses on securities sold are determined on the basis of identified cost and may include proceeds received

Annual Report

3. Significant Accounting Policies - continued

Investment Transactions and Income - continued

from litigation. Dividend income is recorded on the ex-dividend date, except for certain dividends from foreign securities where the ex-dividend date may have passed, which are recorded as soon as the Fund is informed of the ex-dividend date. Non-cash dividends included in dividend income, if any, are recorded at the fair market value of the securities received. Distributions received on securities that represent a return of capital or capital gain are recorded as a reduction of cost of investments and/or as a realized gain. The Fund estimates the components of distributions received that may be considered return of capital distributions or capital gain distributions. Interest income and distributions from the Fidelity Central Funds are accrued as earned. Interest income includes coupon interest and amortization of premium and accretion of discount on debt securities. Investment income is recorded net of foreign taxes withheld where recovery of such taxes is uncertain.

Expenses. Most expenses of the trust can be directly attributed to a fund. Expenses which cannot be directly attributed are apportioned among each Fund in the trust. Expense estimates are accrued in the period to which they relate and adjustments are made when actual amounts are known.

Deferred Trustee Compensation. Under a Deferred Compensation Plan (the Plan), Independent Trustees must defer receipt of a portion of, and may elect to defer receipt of an additional portion of, their annual compensation. Deferred amounts are invested in a cross-section of Fidelity funds, are marked-to-market and remain in the Fund until distributed in accordance with the Plan. The investment of deferred amounts and the offsetting payable to the Trustees are included in the accompanying Statement of Assets and Liabilities.

Income Tax Information and Distributions to Shareholders. Each year, the Fund intends to qualify as a regulated investment company by distributing substantially all of its taxable income and realized gains under Subchapter M of the Internal Revenue Code and filing its U.S. federal tax return. As a result, no provision for income taxes is required. The Fund adopted the provisions of FASB Interpretation No. 48, Accounting for Uncertainties in Income Taxes (FIN 48), on June 29, 2007. FIN 48 sets forth a minimum threshold for financial statement recognition of the benefit of a tax position taken or expected to be taken in a tax return. The implementation of FIN 48 did not result in any unrecognized tax benefits in the accompanying financial statements. Each of the Fund's federal tax returns for the prior three fiscal years remains subject to examination by the Internal Revenue Service. Foreign taxes are provided for based on the Fund's understanding of the tax rules and rates that exist in the foreign markets in which it invests.

Annual Report

Notes to Financial Statements - continued

(Amounts in thousands except ratios)

3. Significant Accounting Policies - continued

Income Tax Information and Distributions to Shareholders - continued

Distributions are recorded on the ex-dividend date. Income and capital gain distributions are determined in accordance with income tax regulations, which may differ from generally accepted accounting principles. In addition, the Fund will claim a portion of the payment made to redeeming shareholders as a distribution for income tax purposes.

Capital accounts within the financial statements are adjusted for permanent book-tax differences. Certain adjustments have been made to the accounts relating to prior periods. Collectively, these adjustments have no impact on net assets or the results of operations. Temporary book-tax differences will reverse in a subsequent period.

Book-tax differences are primarily due to foreign currency transactions, market discount, partnerships, deferred trustees compensation and losses deferred due to wash sales.

The tax-basis components of distributable earnings and the federal tax cost as of period end were as follows:

Unrealized appreciation

$ 7,746,582

Unrealized depreciation

(2,455,619)

Net unrealized appreciation (depreciation)

5,290,963

Undistributed ordinary income

30,508

Undistributed long-term capital gain

453,907

Cost for federal income tax purposes

$ 22,983,802

The tax character of distributions paid was as follows:

January 31, 2008

January 31, 2007

Ordinary Income

$ 546,038

$ 509,068

Long-term Capital Gains

1,676,992

1,669,910

Total

$ 2,223,030

$ 2,178,978

New Accounting Pronouncement. In September 2006, Statement of Financial Accounting Standards No. 157, Fair Value Measurements (SFAS 157), was issued and is effective for fiscal years beginning after November 15, 2007. SFAS 157 defines fair value, establishes a framework for measuring fair value and expands disclosures about fair value measurements. Management is currently evaluating the impact the adoption of SFAS 157 will have on the Fund's financial statement disclosures.

4. Operating Policies.

Repurchase Agreements. FMR has received an Exemptive Order from the Securities and Exchange Commission (the SEC) which permits the Fund and other affiliated entities of FMR to transfer uninvested cash balances into joint trading accounts which

Annual Report

4. Operating Policies - continued

are then invested in repurchase agreements. The Fund may also invest directly with institutions in repurchase agreements. Repurchase agreements are collateralized by government or non-government securities. Upon settlement date, collateral is held in segregated accounts with custodian banks and may be obtained in the event of a default of the counterparty. The Fund monitors, on a daily basis, the value of the collateral to ensure it is at least equal to the principal amount of the repurchase agreement (including accrued interest). In the event of a default by the counterparty, realization of the collateral proceeds could be delayed, during which time the value of the collateral may decline.

Restricted Securities. The Fund may invest in securities that are subject to legal or contractual restrictions on resale. These securities generally may be resold in transactions exempt from registration or to the public if the securities are registered. Disposal of these securities may involve time-consuming negotiations and expense, and prompt sale at an acceptable price may be difficult. Information regarding restricted securities is included at the end of the Fund's Schedule of Investments.

5. Purchases and Sales of Investments.

Purchases and sales of securities, other than short-term securities, aggregated $7,104,567 and $7,282,962, respectively.

6. Fees and Other Transactions with Affiliates.

Management Fee. FMR and its affiliates provide the Fund with investment management related services for which the Fund pays a monthly management fee. The management fee is the sum of an individual fund fee rate that is based on an annual rate of .20% of the Fund's average net assets and a group fee rate that averaged .26% during the period. The group fee rate is based upon the average net assets of all the mutual funds advised by FMR. The group fee rate decreases as assets under management increase and increases as assets under management decrease. For the period, the total annual management fee rate was .46% of the Fund's average net assets.

Transfer Agent Fees. Fidelity Investments Institutional Operations Company, Inc. (FIIOC), an affiliate of FMR, is the Fund's transfer, dividend disbursing and shareholder servicing agent. FIIOC receives account fees and asset-based fees that vary according to account size and type of account. FIIOC pays for typesetting, printing and mailing of shareholder reports, except proxy statements. For the period the transfer agent fees were equivalent to an annual rate of .19% of average net assets. Prior to January 1, 2008, Fidelity Service Company, Inc. (FSC), also an affiliate of FMR was the Fund's transfer agent.

Accounting and Security Lending Fees. FSC maintains the Fund's accounting records. The accounting fee is based on the level of average net assets for the month.

Annual Report

Notes to Financial Statements - continued

(Amounts in thousands except ratios)

6. Fees and Other Transactions with Affiliates - continued

Accounting and Security Lending Fees - continued

Under a separate contract, FSC administers the security lending program. The security lending fee is based on the number and duration of lending transactions.

Brokerage Commissions. The Fund placed a portion of its portfolio transactions with brokerage firms which are affiliates of the investment adviser. The commissions paid to these affiliated firms were $33 for the period.

Interfund Lending Program. Pursuant to an Exemptive Order issued by the SEC, the Fund, along with other registered investment companies having management contracts with FMR, may participate in an interfund lending program. This program provides an alternative credit facility allowing the funds to borrow from, or lend money to, other participating affiliated funds. At period end, there were no interfund loans outstanding. The Fund's activity in this program during the period for which loans were outstanding was as follows:

Borrower or Lender

Average Daily
Loan Balance

Weighted Average
Interest Rate

Interest
Expense

Borrower

$ 42,426

4.38%

$ 36

7. Committed Line of Credit.

The Fund participates with other funds managed by FMR in a $4.2 billion credit facility (the "line of credit") to be utilized for temporary or emergency purposes to fund shareholder redemptions or for other short-term liquidity purposes. The Fund has agreed to pay commitment fees on its pro rata portion of the line of credit, which amounted to $61 and is reflected in Miscellaneous Expense on the Statement of Operations. During the period, there were no borrowings on this line of credit.

8. Security Lending.

The Fund lends portfolio securities from time to time in order to earn additional income. On the settlement date of the loan, the Fund receives collateral (in the form of U.S. Treasury obligations, letters of credit and/or cash) against the loaned securities and maintains collateral in an amount not less than 100% of the market value of the loaned securities during the period of the loan. The market value of the loaned securities is determined at the close of business of the Fund and any additional required collateral is delivered to the Fund on the next business day. If the borrower defaults on its obligation to return the securities loaned because of insolvency or other reasons, a fund could experience delays and costs in recovering the securities loaned or in gaining access to the collateral. Any cash collateral received is invested in the Fidelity Securities Lending Cash Central Fund. The value of loaned securities and cash collateral at period end are disclosed on the Fund's Statement of Assets and Liabilities. Security lending income represents the income earned on investing cash collateral, less fees and expenses associated with the loan, plus any premium payments that may be received on the loan of

Annual Report

8. Security Lending - continued

certain types of securities. Security lending income is presented in the Statement of Operations as a component of income from Fidelity Central Funds. Net income from lending portfolio securities during the period amounted to $4,263.

9. Expense Reductions.

Many of the brokers with whom FMR places trades on behalf of the Fund provided services to the Fund in addition to trade execution. These services included payments of certain expenses on behalf of the Fund totaling $157 for the period. In addition, through arrangements with the Fund's custodian and transfer agent, credits realized as a result of uninvested cash balances were used to reduce the Fund's expenses. During the period, these credits reduced the Fund's custody and transfer agent expenses by $30 and $2,431, respectively.

10. Other.

The Fund's organizational documents provide former and current trustees and officers with a limited indemnification against liabilities arising in connection with the performance of their duties to the Fund. In the normal course of business, the Fund may also enter into contracts that provide general indemnifications. The Fund's maximum exposure under these arrangements is unknown as this would be dependent on future claims that may be made against the Fund. The risk of material loss from such claims is considered remote.

In December 2006, the Independent Trustees, with the assistance of independent counsel, completed an investigation regarding gifts, gratuities and business entertainment provided by certain brokers to certain individuals who were employed on FMR's domestic equity trading desk during the period 2002 to 2004. The Independent Trustees and FMR agreed that, despite the absence of proof that the Fidelity mutual funds experienced diminished execution quality as a result of the improper receipt of gifts and business entertainment, the conduct at issue was serious and was worthy of redress. Accordingly, the Independent Trustees requested, and FMR agreed to make a payment of $42 million plus accrued interest, which equaled approximately $7.3 million, to certain Fidelity mutual funds. FMR also reimbursed the related legal expenses, which are recorded in the accompanying Statement of Operations as an expense reduction.

Subsequent to period end, the Trustees approved a method for allocating this payment among the funds and, in total, FMR paid the fund $935.

In a related administrative order dated March 5, 2008, the U.S. Securities and Exchange Commission ("SEC") announced a settlement with FMR and FMR Co., Inc. (an affiliate of FMR) involving the SEC's regulatory rules for investment advisers and the improper receipt of gifts, gratuities and business entertainment. Without admitting or denying the SEC's findings, FMR agreed to pay an $8 million civil penalty to the United States Treasury.

Annual Report

Report of Independent Registered Public Accounting Firm

To the Trustees of Fidelity Devonshire Trust and the Shareholders of Fidelity Equity-Income Fund:

In our opinion, the accompanying statement of assets and liabilities, including the schedule of investments, and the related statements of operations and of changes in net assets and the financial highlights present fairly, in all material respects, the financial position of Fidelity Equity-Income Fund (a fund of Fidelity Devonshire Trust) at January 31, 2008, the results of its operations for the year then ended, the changes in its net assets for each of the two years in the period then ended and the financial highlights for each of the five years in the period then ended, in conformity with accounting principles generally accepted in the United States of America. These financial statements and financial highlights (hereafter referred to as "financial statements") are the responsibility of the Fidelity Equity-Income Fund's management. Our responsibility is to express an opinion on these financial statements based on our audits. We conducted our audits of these financial statements in accordance with the standards of the Public Company Accounting Oversight Board (United States). Those standards require that we plan and perform the audit to obtain reasonable assurance about whether the financial statements are free of material misstatement. An audit includes examining, on a test basis, evidence supporting the amounts and disclosures in the financial statements, assessing the accounting principles used and significant estimates made by management, and evaluating the overall financial statement presentation. We believe that our audits, which included confirmation of securities at January 31, 2008 by correspondence with the custodian and brokers, provide a reasonable basis for our opinion.

/s/ PricewaterhouseCoopers LLP

PricewaterhouseCoopers LLP

Boston, Massachusetts

March 24, 2008

Annual Report

Trustees and Officers

The Trustees, Members of the Advisory Board, and executive officers of the trust and fund, as applicable, are listed below. The Board of Trustees governs the fund and is responsible for protecting the interests of shareholders. The Trustees are experienced executives who meet periodically throughout the year to oversee the fund's activities, review contractual arrangements with companies that provide services to the fund, and review the fund's performance. Except for James C. Curvey, each of the Trustees oversees 374 funds advised by FMR or an affiliate. Mr. Curvey oversees 369 funds advised by FMR or an affiliate.

The Trustees hold office without limit in time except that (a) any Trustee may resign; (b) any Trustee may be removed by written instrument, signed by at least two-thirds of the number of Trustees prior to such removal; (c) any Trustee who requests to be retired or who has become incapacitated by illness or injury may be retired by written instrument signed by a majority of the other Trustees; and (d) any Trustee may be removed at any special meeting of shareholders by a two-thirds vote of the outstanding voting securities of the trust. Each Trustee who is not an interested person (as defined in the 1940 Act) (Independent Trustee), shall retire not later than the last day of the calendar year in which his or her 72nd birthday occurs. The Independent Trustees may waive this mandatory retirement age policy with respect to individual Trustees. The executive officers and Advisory Board Members hold office without limit in time, except that any officer and Advisory Board Member may resign or may be removed by a vote of a majority of the Trustees at any regular meeting or any special meeting of the Trustees. Except as indicated, each individual has held the office shown or other offices in the same company for the past five years.

The fund's Statement of Additional Information (SAI) includes more information about the Trustees. To request a free copy, call Fidelity at 1-800-544-8544.

Interested Trustees*:

Correspondence intended for each Trustee who is an interested person may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Edward C. Johnson 3d (77)

Year of Election or Appointment: 1985

Mr. Johnson is Chairman of the Board of Trustees. Mr. Johnson serves as President (2006-present), Chief Executive Officer, Chairman, and a Director of FMR LLC; Chairman and a Director of FMR; Chairman and a Director of Fidelity Research & Analysis Company (FRAC); Chairman and a Director of Fidelity Investments Money Management, Inc.; and Chairman (2001-present) and a Director of FMR Co., Inc. In addition, Mr. Johnson serves as Chairman and Director of FIL Limited. Mr. Edward C. Johnson 3d and Mr. Arthur E. Johnson are not related.

James C. Curvey (72)

Year of Election or Appointment: 2007

Mr. Curvey also serves as Trustee (2007-present) or Member of the Advisory Board (2007-present) of other investment companies advised by FMR. Mr. Curvey is a Director of FMR and FMR Co., Inc. (2007-present). Mr. Curvey is also Vice Chairman (2006-present) and Director of FMR LLC. Mr. Curvey joined Fidelity in 1982 and served in numerous senior management positions, including President and Chief Operating Officer of FMR LLC (1997-2000) and President of Fidelity Strategic Investments (2000-2002). In addition, he serves as a member of the Board of Directors of Geerlings & Wade, Inc. (wine distribution).

* Trustees have been determined to be "Interested Trustees" by virtue of, among other things, their affiliation with the trust or various entities under common control with FMR. FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Independent Trustees:

Correspondence intended for each Independent Trustee (that is, the Trustees other than the Interested Trustees) may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235.

Name, Age; Principal Occupation

Dennis J. Dirks (59)

Year of Election or Appointment: 2005

Prior to his retirement in May 2003, Mr. Dirks was Chief Operating Officer and a member of the Board of The Depository Trust & Clearing Corporation (DTCC) (1999-2003). He also served as President, Chief Operating Officer, and Board member of The Depository Trust Company (DTC) (1999-2003) and President and Board member of the National Securities Clearing Corporation (NSCC) (1999-2003). In addition, Mr. Dirks served as Chief Executive Officer and Board member of the Government Securities Clearing Corporation (2001-2003) and Chief Executive Officer and Board member of the Mortgage-Backed Securities Clearing Corporation (2001-2003). Mr. Dirks also serves as a Trustee and a member of the Finance Committee of Manhattan College (2005-present) and a Trustee and a member of the Finance Committee of AHRC of Nassau County (2006-present).

Albert R. Gamper, Jr. (65)

Year of Election or Appointment: 2006

Prior to his retirement in December 2004, Mr. Gamper served as Chairman of the Board of CIT Group Inc. (commercial finance). During his tenure with CIT Group Inc. Mr. Gamper served in numerous senior management positions, including Chairman (1987-1989; 1999-2001; 2002-2004), Chief Executive Officer (1987-2004), and President (1989-2002). He currently serves as a member of the Board of Directors of Public Service Enterprise Group (utilities, 2001-present), Chairman of the Board of Governors, Rutgers University (2004-present), and Chairman of the Board of Saint Barnabas Health Care System.

George H. Heilmeier (71)

Year of Election or Appointment: 2004

Dr. Heilmeier is Chairman Emeritus of Telcordia Technologies (communication software and systems), where prior to his retirement, he served as company Chairman and Chief Executive Officer. He currently serves on the Boards of Directors of The Mitre Corporation (systems engineering and information technology support for the government), and HRL Laboratories (private research and development, 2004-present). He is Chairman of the General Motors Science & Technology Advisory Board and a Life Fellow of the Institute of Electrical and Electronics Engineers (IEEE). Dr. Heilmeier is a member of the Defense Science Board and the National Security Agency Advisory Board. He is also a member of the National Academy of Engineering, the American Academy of Arts and Sciences, and the Board of Overseers of the School of Engineering and Applied Science of the University of Pennsylvania. Previously, Dr. Heilmeier served as a Director of TRW Inc. (automotive, space, defense, and information technology, 1992-2002), Compaq (1994-2002), Automatic Data Processing, Inc. (ADP) (technology-based business outsourcing, 1995-2002), INET Technologies Inc. (telecommunications network surveillance, 2001-2004), and Teletech Holdings (customer management services). He is the recipient of the 2005 Kyoto Prize in Advanced Technology for his invention of the liquid crystal display, and a member of the Consumer Electronics Hall of Fame.

James H. Keyes (67)

Year of Election or Appointment: 2007

Prior to his retirement in 2003, Mr. Keyes was Chairman, President, and Chief Executive Officer of Johnson Controls, Inc. (automotive supplier, 1993-2003). He currently serves as a member of the boards of LSI Logic Corporation (semiconductor technologies), Navistar International Corporation (manufacture and sale of trucks, buses, and diesel engines, 2002-present), and Pitney Bowes, Inc. (integrated mail, messaging, and document management solutions).

Marie L. Knowles (61)

Year of Election or Appointment: 2001

Prior to Ms. Knowles' retirement in June 2000, she served as Executive Vice President and Chief Financial Officer of Atlantic Richfield Company (ARCO) (diversified energy, 1996-2000). From 1993 to 1996, she was a Senior Vice President of ARCO and President of ARCO Transportation Company. She served as a Director of ARCO from 1996 to 1998. She currently serves as a Director of Phelps Dodge Corporation (copper mining and manufacturing) and McKesson Corporation (healthcare service, 2002-present). Ms. Knowles is a Trustee of the Brookings Institution and the Catalina Island Conservancy and also serves as a member of the Advisory Board for the School of Engineering of the University of Southern California.

Ned C. Lautenbach (63)

Year of Election or Appointment: 2000

Mr. Lautenbach is Chairman of the Independent Trustees (2006-present). Mr. Lautenbach has been a partner of Clayton, Dubilier & Rice, Inc. (private equity investment firm) since September 1998. Previously, Mr. Lautenbach was with the International Business Machines Corporation (IBM) from 1968 until his retirement in 1998. Mr. Lautenbach serves as a Director of Sony Corporation (2006-present) and Eaton Corporation (diversified industrial) as well as the Philharmonic Center for the Arts in Naples, Florida. He also is a member of the Board of Trustees of Fairfield University (2005-present), as well as a member of the Council on Foreign Relations.

Cornelia M. Small (63)

Year of Election or Appointment: 2005

Ms. Small is a member (2000-present) and Chairperson (2002-present) of the Investment Committee, and a member (2002-present) of the Board of Trustees of Smith College. Previously, she served as Chief Investment Officer (1999-2000), Director of Global Equity Investments (1996-1999), and a member of the Board of Directors of Scudder, Stevens & Clark (1990-1997) and Scudder Kemper Investments (1997-1999). In addition, Ms. Small served as Co-Chair (2000-2003) of the Annual Fund for the Fletcher School of Law and Diplomacy.

William S. Stavropoulos (68)

Year of Election or Appointment: 2002

Mr. Stavropoulos is Chairman Emeritus of the Board of Directors of The Dow Chemical Company. Since joining The Dow Chemical Company in 1967, Mr. Stavropoulos served in numerous senior management positions, including President (1993-2000; 2002-2003), CEO (1995-2000; 2002-2004), and Chairman of the Executive Committee (2000-2004). Currently, he is a Director of NCR Corporation (data warehousing and technology solutions), Chemical Financial Corporation, Maersk Inc. (industrial conglomerate, 2002-present), Tyco International, Inc. (multinational manufacturing and services, 2007-present), and a member of the Advisory Board for Metalmark Capital (private equity investment firm, 2005-present). He is a special advisor to Clayton, Dubilier & Rice, Inc., a private equity investment firm. He also serves as a member of the Board of Trustees of the American Enterprise Institute for Public Policy Research. In addition, Mr. Stavropoulos is a member of The Business Council, J.P. Morgan International Council and the University of Notre Dame Advisory Council for the College of Science.

Kenneth L. Wolfe (68)

Year of Election or Appointment: 2005

Mr. Wolfe is Chairman and a Director of Hershey Foods Corporation (2007-present), where prior to his retirement in 2001, he was Chairman and Chief Executive Officer. Mr. Wolfe currently serves as a member of the board of Revlon Inc. (2004-present). Previously, Mr. Wolfe served as a member of the boards of Adelphia Communications Corporation (2003-2006) and Bausch & Lomb, Inc. (1993-2007).

Advisory Board Members and Executive Officers**:

Correspondence intended for Mr. Mauriello, Mr. Thomas, Mr. Wiley, Mr. Lacy, and Mr. Arthur Johnson may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235. Correspondence intended for each executive officer and Mr. Lynch may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Annual Report

Trustees and Officers - continued

Name, Age; Principal Occupation

Arthur E. Johnson (61)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Johnson serves as Senior Vice President of Corporate Strategic Development of Lockheed Martin Corporation (defense contractor). In addition, Mr. Johnson serves as a member of the Board of Directors of AGL Resources, Inc. (holding company, 2002-present), and IKON Office Solutions, Inc. (document management systems and services). Mr. Arthur E. Johnson and Mr. Edward C. Johnson 3d are not related.

Alan J. Lacy (54)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lacy serves as Senior Adviser (2007-present) of Oak Hill Capital Partners, L.P. (a private equity firm). Mr. Lacy also served as Vice Chairman and Chief Executive Officer of Sears Holdings Corporation and Sears, Roebuck and Co. (retail, 2005-2006; 2000-2005). In addition, Mr. Lacy serves as a member of the Board of Directors of The Western Union Company (global money transfer, 2006-present) and Bristol-Myers Squibb (global pharmaceuticals, 2007-present). Mr. Lacy is a Trustee of the National Parks Conservation Association and The Field Museum of Natural History.

Peter S. Lynch (64)

Year of Election or Appointment: 2003

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lynch is Vice Chairman and a Director of FMR, and Vice Chairman (2001-present) and a Director of FMR Co., Inc. Previously, Mr. Lynch served as a Trustee of the Fidelity funds (1990-2003). In addition, he serves as a Trustee of Boston College and as the Chairman of the Inner-City Scholarship Fund.

Joseph Mauriello (63)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Prior to his retirement in January 2006, Mr. Mauriello served in numerous senior management positions including Deputy Chairman and Chief Operating Officer (2004-2005), and Vice Chairman of Financial Services (2002-2004) of KPMG LLP US (professional services firm, 1965-2005). Mr. Mauriello currently serves as a member of the Board of Directors of XL Capital Ltd. (global insurance and re-insurance company, 2006-present) and of Arcadia Resources Inc. (health care services and products, 2007-present). He also served as a Director of the Hamilton Funds of the Bank of New York (2006-2007).

David M. Thomas (58)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Previously, Mr. Thomas served as Executive Chairman (2005-2006) and Chairman and Chief Executive Officer (2000-2005) of IMS Health, Inc. (pharmaceutical and healthcare information solutions). In addition, Mr. Thomas serves as a member of the Board of Directors of Fortune Brands, Inc. (consumer products holding company), and Interpublic Group of Companies, Inc. (marketing communication, 2004-present).

Michael E. Wiley (57)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Wiley also serves as a member of the Board of Trustees of the University of Tulsa (2000-2006; 2007-present). He serves as a Director of Tesoro Corporation (independent oil refiner and marketer, 2005-present), and a Director of Bill Barrett Corporation (exploration and production company, 2005-present). In addition, he also serves as a Director of Post Oak Bank (privately-held bank, 2004-present). Previously, Mr. Wiley served as a Sr. Energy Advisor of Katzenbach Partners, LLC (consulting firm, 2006-2007), as an Advisory Director of Riverstone Holdings (private investment firm), Chairman, President, and CEO of Baker Hughes, Inc. (oilfield services company, 2000-2004), and as Director of Spinnaker Exploration Company (exploration and production company, 2001-2005).

Kimberley H. Monasterio (44)

Year of Election or Appointment: 2007

President and Treasurer of Equity-Income. Ms. Monasterio also serves as President and Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2004-present). Previously, Ms. Monasterio served as Deputy Treasurer of the Fidelity funds (2004-2006). Before joining Fidelity Investments, Ms. Monasterio served as Treasurer (2000-2004) and Chief Financial Officer (2002-2004) of the Franklin Templeton Funds and Senior Vice President of Franklin Templeton Services, LLC (2000-2004).

Name, Age; Principal Occupation

Walter C. Donovan (45)

Year of Election or Appointment: 2007

Vice President of Equity-Income. Mr. Donovan also serves as Vice President of Fidelity's Equity Funds (2007-present). Mr. Donovan also serves as Executive Vice President of FMR (2005-present) and FMR Co., Inc. (2005-present). Previously, Mr. Donovan served as Vice President of Fidelity's High Income Funds (2005-2007), Fixed-Income Funds (2005-2006), certain Asset Allocation Funds (2005-2006), certain Balanced Funds (2005-2006), and as Vice President and Director of Fidelity's International Equity Trading group (1998-2005).

Bruce T. Herring (42)

Year of Election or Appointment: 2006

Vice President of Equity-Income. Mr. Herring also serves as Vice President of certain Equity Funds (2006-present). Mr. Herring is Senior Vice President of FMR (2006-present) and Vice President of FMR Co., Inc. (2001-present). Previously, Mr. Herring served as a portfolio manager for Fidelity U.S. Equity Funds (2001-2005).

Eric D. Roiter (59)

Year of Election or Appointment: 1998

Secretary of Equity-Income. He also serves as Secretary of other Fidelity funds; Vice President, General Counsel, and Secretary of FMR Co., Inc. (2001-present) and FMR; Assistant Secretary of Fidelity Management & Research (U.K.) Inc. (2001-present), Fidelity Research & Analysis Company (2001-present), and Fidelity Investments Money Management, Inc. (2001-present). Mr. Roiter is an Adjunct Member, Faculty of Law, at Boston College Law School (2003-present). Previously, Mr. Roiter served as Vice President and Secretary of Fidelity Distributors Corporation (FDC) (1998-2005).

John B. McGinty, Jr. (45)

Year of Election or Appointment: 2008

Assistant Secretary of Equity-Income. Mr. McGinty also serves as Assistant Secretary of other Fidelity funds (2008-present) and is an employee of FMR LLC (2004-present). Mr. McGinty also serves as Senior Vice President, Secretary, and Chief Legal Officer of FDC (2007-present). Before joining Fidelity Investments, Mr. McGinty practiced law at Ropes & Gray, LLP.

R. Stephen Ganis (41)

Year of Election or Appointment: 2006

Anti-Money Laundering (AML) officer of Equity-Income. Mr. Ganis also serves as AML officer of other Fidelity funds (2006-present) and FMR LLC (2003-present). Before joining Fidelity Investments, Mr. Ganis practiced law at Goodwin Procter, LLP (2000-2002).

Joseph B. Hollis (59)

Year of Election or Appointment: 2006

Chief Financial Officer of Equity-Income. Mr. Hollis also serves as Chief Financial Officer of other Fidelity funds. Mr. Hollis is President of Fidelity Pricing and Cash Management Services (FPCMS) (2005-present). Mr. Hollis also serves as President and Director of Fidelity Service Company, Inc. (2006-present). Previously, Mr. Hollis served as Senior Vice President of Cash Management Services (1999-2002) and Investment Management Operations (2002-2005).

Kenneth A. Rathgeber (60)

Year of Election or Appointment: 2004

Chief Compliance Officer of Equity-Income. Mr. Rathgeber also serves as Chief Compliance Officer of other Fidelity funds (2004-present) and Executive Vice President of Risk Oversight for Fidelity Investments (2002-present). He is Chief Compliance Officer of FMR (2005-present), FMR Co., Inc. (2005-present), Fidelity Management & Research (U.K.) Inc. (2005-present), Fidelity Research & Analysis Company (2005-present), Fidelity Investments Money Management, Inc. (2005-present), and Strategic Advisers, Inc. (2005-present). Previously, Mr. Rathgeber served as Executive Vice President and Chief Operating Officer for Fidelity Investments Institutional Services Company, Inc. (1998-2002).

Bryan A. Mehrmann (46)

Year of Election or Appointment: 2005

Deputy Treasurer of Equity-Income. Mr. Mehrmann also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR. Previously, Mr. Mehrmann served as Vice President of Fidelity Investments Institutional Services Group (FIIS)/Fidelity Investments Institutional Operations Corporation, Inc. (FIIOC) Client Services (1998-2004).

Kenneth B. Robins (38)

Year of Election or Appointment: 2005

Deputy Treasurer of Equity-Income. Mr. Robins also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2004-present). Before joining Fidelity Investments, Mr. Robins worked at KPMG LLP, where he was a partner in KPMG's department of professional practice (2002-2004) and a Senior Manager (1999-2000). In addition, Mr. Robins served as Assistant Chief Accountant, United States Securities and Exchange Commission (2000-2002).

Robert G. Byrnes (41)

Year of Election or Appointment: 2005

Assistant Treasurer of Equity-Income. Mr. Byrnes also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Byrnes served as Vice President of FPCMS (2003-2005). Before joining Fidelity Investments, Mr. Byrnes worked at Deutsche Asset Management where he served as Vice President of the Investment Operations Group (2000-2003).

Peter L. Lydecker (54)

Year of Election or Appointment: 2004

Assistant Treasurer of Equity-Income. Mr. Lydecker also serves as Assistant Treasurer of other Fidelity funds (2004) and is an employee of FMR.

Paul M. Murphy (60)

Year of Election or Appointment: 2007

Assistant Treasurer of Equity-Income. Mr. Murphy also serves as Assistant Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2007-present). Previously, Mr. Murphy served as Chief Financial Officer of the Fidelity Funds (2005-2006), Vice President and Associate General Counsel of FMR (2007), and Senior Vice President of Fidelity Pricing and Cash Management Services Group (FPCMS) (1994-2007).

Gary W. Ryan (49)

Year of Election or Appointment: 2005

Assistant Treasurer of Equity-Income. Mr. Ryan also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Ryan served as Vice President of Fund Reporting in FPCMS (1999-2005).

** FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Annual Report

Distributions

The Board of Trustees of Fidelity Equity-Income Fund voted to pay on March 10, 2008, to shareholders of record at the opening of business on March 7, 2008, a distribution of $.85 per share derived from capital gains realized from sales of portfolio securities.

The fund hereby designates as a capital gain dividend with respect to the taxable year ended January 31, 2008, $1,706,050,156, or, if subsequently determined to be different, the net capital gain of such year.

A total of 0.18% of the dividends distributed during the fiscal year was derived from interest on U.S. Government securities which is generally exempt from state income tax.

The fund designates 100% of the dividends distributed during the fiscal year as qualifying for the dividends-received deduction for corporate shareholders.

The fund designates 100% of the dividends distributed during the fiscal year as amounts which may be taken into account as a dividend for the purposes of the maximum rate under section 1(h)(11) of the Internal Revenue Code.

The fund will notify shareholders in January 2009 of amounts for use in preparing 2008 income tax returns.

Annual Report

Managing Your Investments

Fidelity offers several ways to conveniently manage your personal investments via your telephone or PC. You can access your account information, conduct trades and research your investments 24 hours a day.

By Phone

Fidelity Automated Service Telephone provides a single toll-free number to access account balances, positions, quotes and trading. It's easy to navigate the service, and on your first call, the system will help you create a personal identification number (PIN) for security.

(phone_graphic)

Fidelity Automated
Service Telephone (FAST
®)
1-800-544-5555

Press

1   For mutual fund and brokerage trading.

2   For quotes.*

3   For account balances and holdings.

4   To review orders and mutual
fund activity.

5   To change your PIN.

*0   To speak to a Fidelity representative.

By PC

Fidelity's web site on the Internet provides a wide range of information, including daily financial news, fund performance, interactive planning tools and news about Fidelity products and services.

(computer_graphic)

Fidelity's Web Site
www.fidelity.com

* When you call the quotes line, please remember that a fund's yield and return will vary and, except for money market funds, share price will also vary. This means that you may have a gain or loss when you sell your shares. There is no assurance that money market funds will be able to maintain a stable $1 share price; an investment in a money market fund is not insured or guaranteed by the U.S. government. Total returns are historical and include changes in share price, reinvestment of dividends and capital gains, and the effects of any sales charges.

Annual Report

Investment Adviser

Fidelity Management & Research Company

Boston, MA

Investment Sub-Advisers

FMR Co., Inc.

Fidelity Research & Analysis Company

Fidelity Management & Research
(U.K.) Inc.

Fidelity Investments Japan Limited

Fidelity International Investment Advisors

Fidelity International Investment Advisors
(U.K.) Limited

General Distributor

Fidelity Distributors Corporation

Boston, MA

Transfer and Service Agents

Fidelity Investments Institutional
Operations Company, Inc.

Boston, MA

Fidelity Service Company, Inc.

Boston, MA

Custodian

The Northern Trust Company
Chicago, IL

The Fidelity Telephone Connection

Mutual Fund 24-Hour Service

Exchanges/Redemptions
and Account Assistance 1-800-544-6666

Product Information 1-800-544-6666

Retirement Accounts 1-800-544-4774
(8 a.m. - 9 p.m.)

TDD Service 1-800-544-0118
for the deaf and hearing impaired
(9 a.m. - 9 p.m. Eastern time)

Fidelity Automated Service
Telephone (FAST®) (automated graphic)    1-800-544-5555

(automated graphic)    Automated line for quickest service

(Fidelity Investment logo)(registered trademark)
Corporate Headquarters
82 Devonshire St., Boston, MA 02109
www.fidelity.com

EQU-UANN-0308
1.789253.106

Fidelity® Large Cap Value

Fidelity Mid Cap Value

Fidelity Large Cap Growth

Fidelity Mid Cap Growth

Funds

Annual Report

January 31, 2008

(2_fidelity_logos) (Registered_Trademark)

Contents

Chairman's Message

<Click Here>

Ned Johnson's message to shareholders.

Shareholder Expense Example

<Click Here>

An example of shareholder expenses.

Fidelity Large Cap Value Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investments Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Mid Cap Value Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Large Cap Growth Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Mid Cap Growth Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Notes

<Click Here>

Notes to the Financial Statements

Report of Independent Registered Public Accounting Firm

<Click Here>

Trustees and Officers

<Click Here>

Distributions

<Click Here>

To view a fund's proxy voting guidelines and proxy voting record for the 12-month period ended June 30, visit http://www.fidelity.com (search for "proxy voting guidelines") or visit the Securities and Exchange Commission's (SEC) web site at http://www.sec.gov. You may also call 1-800-544-8544 to request a free copy of the proxy voting guidelines.

Standard & Poor's, S&P and S&P 500 are registered service marks of The McGraw-Hill Companies, Inc. and have been licensed for use by Fidelity Distributors Corporation.

Other third party marks appearing herein are the property of their respective owners.

All other marks appearing herein are registered or unregistered trademarks or service marks of FMR LLC or an affiliated company.

This report and the financial statements contained herein are submitted for the general information of the shareholders of the funds. This report is not authorized for distribution to prospective investors in the funds unless preceded or accompanied by an effective prospectus.

A fund files its complete schedule of portfolio holdings with the SEC for the first and third quarters of each fiscal year on Form N-Q. Forms N-Q are available on the SEC's web site at http://www.sec.gov. A fund's Forms N-Q may be reviewed and copied at the SEC's Public Reference Room in Washington, DC. Information regarding the operation of the SEC's Public Reference Room may be obtained by calling 1-800-SEC-0330. For a complete list of a fund's portfolio holdings, view the most recent holdings listing, semiannual report, or annual report on Fidelity's web site at http://www.fidelity.com or http://www.advisor.fidelity.com, as applicable.

NOT FDIC INSURED · MAY LOSE VALUE · NO BANK GUARANTEE

Neither the funds nor Fidelity Distributors Corporation is a bank.

Annual Report

Chairman's Message

(Photograph of Edward C. Johnson 3d.)

Dear Shareholder:

Stocks got off to a poor start in 2008, while investment-grade bonds and money markets showed positive returns, once again underscoring the importance of a diversified portfolio. Financial markets are always unpredictable, but there are a number of time-tested principles that can put the historical odds in your favor.

One of the basic tenets is to invest for the long term. Over time, riding out the markets' inevitable ups and downs has proven much more effective than selling into panic or chasing the hottest trend. Even missing only a few of the markets' best days can significantly diminish investor returns. Patience also affords the benefits of compounding - of earning interest on additional income or reinvested dividends and capital gains. There are tax advantages and cost benefits to consider as well. The more you sell, the more taxes you pay, and the more you trade, the higher the costs. While staying the course doesn't eliminate risk, it can considerably lessen the effect of short-term declines.

You can further manage your investing risk through diversification. And today, more than ever, geographic diversification should be taken into account. Studies

indicate that asset allocation is the single most important determinant of a portfolio's long-term success. The right mix of stocks, bonds and cash - aligned to your particular risk tolerance and investment objective - is very important. Age-appropriate rebalancing is also an essential aspect of asset allocation. For younger investors, an emphasis on equities - which historically have been the best-performing asset class over time - is encouraged. As investors near their specific goal, such as retirement or sending a child to college, consideration may be given to replacing volatile assets (e.g. common stocks) with more-stable fixed investments (bonds or savings plans).

A third investment principle - investing regularly - can help lower the average cost of your purchases. Investing a certain amount of money each month or quarter helps ensure you won't pay for all your shares at market highs. This strategy - known as dollar cost averaging - also reduces unconstructive "emotion" from investing, helping shareholders avoid selling weak performers just prior to an upswing, or chasing a hot performer just before a correction.

We invite you to contact us via the Internet, through our Investor Centers or over the phone. It is our privilege to provide you the information you need to make the investments that are right for you.

Sincerely,

/s/Edward C. Johnson 3d

Edward C. Johnson 3d

Annual Report

Shareholder Expense Example

As a shareholder of a Fund, you incur two types of costs: (1) transaction costs, including sales charges (loads) on purchase payments or redemption proceeds, redemption fees, and (2) ongoing costs, including management fees, distribution and/or service (12b-1) fees and other Fund expenses. This Example is intended to help you understand your ongoing costs (in dollars) of investing in the Funds and to compare these costs with the ongoing costs of investing in other mutual funds.

The Example is based on an investment of $1,000 invested at the beginning of the period and held for the entire period (August 1, 2007 to January 31, 2008).

Actual Expenses

The first line of the accompanying table for each class of each fund provides information about actual account values and actual expenses. You may use the information in this line, together with the amount you invested, to estimate the expenses that you paid over the period. Simply divide your account value by $1,000.00 (for example, an $8,600 account value divided by $1,000.00 = 8.6), then multiply the result by the number in the first line for a class of the fund under the heading entitled "Expenses Paid During Period" to estimate the expenses you paid on your account during this period. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, each Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Hypothetical Example for Comparison Purposes

The second line of the accompanying table for each class of each fund provides information about hypothetical account values and hypothetical expenses based on a Class' actual expense ratio and an assumed rate of return of 5% per year before expenses, which is not the Class' actual return. The hypothetical account values and expenses may not be used to estimate the actual ending account balance or expenses you paid for the period. You may use this information to compare the ongoing costs of investing in the Fund and other funds. To do so, compare this 5% hypothetical example with the 5% hypothetical examples that appear in the shareholder reports of the other funds. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, each Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Please note that the expenses shown in the table are meant to highlight your ongoing costs only and do not reflect any transaction costs. Therefore, the second line of the table is useful in comparing ongoing costs only, and will not help you determine the relative total costs of owning different funds. In addition, if these transactional costs were included, your costs would have been higher.

Beginning
Account Value
August 1, 2007

Ending
Account Value
January 31, 2008

Expenses Paid
During Period
*
August 1, 2007
to January 31, 2008

Fidelity Large Cap Value Fund

Class A

Actual

$ 1,000.00

$ 946.40

$ 6.13

HypotheticalA

$ 1,000.00

$ 1,018.90

$ 6.36

Class T

Actual

$ 1,000.00

$ 944.70

$ 7.35

HypotheticalA

$ 1,000.00

$ 1,017.64

$ 7.63

Class B

Actual

$ 1,000.00

$ 943.10

$ 9.80

HypotheticalA

$ 1,000.00

$ 1,015.12

$ 10.16

Class C

Actual

$ 1,000.00

$ 943.20

$ 9.65

HypotheticalA

$ 1,000.00

$ 1,015.27

$ 10.01

Large Cap Value

Actual

$ 1,000.00

$ 948.00

$ 4.27

HypotheticalA

$ 1,000.00

$ 1,020.82

$ 4.43

Institutional Class

Actual

$ 1,000.00

$ 948.00

$ 4.22

HypotheticalA

$ 1,000.00

$ 1,020.87

$ 4.38

Fidelity Mid Cap Value Fund

Class A

Actual

$ 1,000.00

$ 912.50

$ 5.54

HypotheticalA

$ 1,000.00

$ 1,019.41

$ 5.85

Beginning
Account Value
August 1, 2007

Ending
Account Value
January 31, 2008

Expenses Paid
During Period
*
August 1, 2007
to January 31, 2008

Class T

Actual

$ 1,000.00

$ 911.80

$ 6.79

HypotheticalA

$ 1,000.00

$ 1,018.10

$ 7.17

Class B

Actual

$ 1,000.00

$ 909.00

$ 9.09

HypotheticalA

$ 1,000.00

$ 1,015.68

$ 9.60

Class C

Actual

$ 1,000.00

$ 909.00

$ 9.24

HypotheticalA

$ 1,000.00

$ 1,015.53

$ 9.75

Mid Cap Value

Actual

$ 1,000.00

$ 913.40

$ 4.05

HypotheticalA

$ 1,000.00

$ 1,020.97

$ 4.28

Institutional Class

Actual

$ 1,000.00

$ 913.70

$ 4.34

HypotheticalA

$ 1,000.00

$ 1,020.67

$ 4.58

Fidelity Large Cap Growth Fund

Class A

Actual

$ 1,000.00

$ 955.40

$ 5.91

HypotheticalA

$ 1,000.00

$ 1,019.16

$ 6.11

Class T

Actual

$ 1,000.00

$ 955.60

$ 7.30

HypotheticalA

$ 1,000.00

$ 1,017.74

$ 7.53

Class B

Actual

$ 1,000.00

$ 952.20

$ 9.74

HypotheticalA

$ 1,000.00

$ 1,015.22

$ 10.06

Class C

Actual

$ 1,000.00

$ 953.00

$ 9.65

HypotheticalA

$ 1,000.00

$ 1,015.32

$ 9.96

Large Cap Growth

Actual

$ 1,000.00

$ 956.60

$ 4.78

HypotheticalA

$ 1,000.00

$ 1,020.32

$ 4.94

Institutional Class

Actual

$ 1,000.00

$ 958.20

$ 4.10

HypotheticalA

$ 1,000.00

$ 1,021.02

$ 4.23

Fidelity Mid Cap Growth Fund

Class A

Actual

$ 1,000.00

$ 909.40

$ 5.29

HypotheticalA

$ 1,000.00

$ 1,019.66

$ 5.60

Class T

Actual

$ 1,000.00

$ 905.80

$ 6.58

HypotheticalA

$ 1,000.00

$ 1,018.30

$ 6.97

Class B

Actual

$ 1,000.00

$ 904.90

$ 8.79

HypotheticalA

$ 1,000.00

$ 1,015.98

$ 9.30

Class C

Actual

$ 1,000.00

$ 904.20

$ 8.88

HypotheticalA

$ 1,000.00

$ 1,015.88

$ 9.40

Mid Cap Growth

Actual

$ 1,000.00

$ 909.50

$ 3.75

HypotheticalA

$ 1,000.00

$ 1,021.27

$ 3.97

Institutional Class

Actual

$ 1,000.00

$ 909.60

$ 3.32

HypotheticalA

$ 1,000.00

$ 1,021.73

$ 3.52

A 5% return per year before expenses

* Expenses are equal to each Class' annualized expense ratio (shown in the table below); multiplied by the average account value over the period, multiplied by 184/ 365 (to reflect the one-half year period).

Annual Report

Annualized
Expense Ratio

Fidelity Large Cap Value Fund

Class A

1.25%

Class T

1.50%

Class B

2.00%

Class C

1.97%

Large Cap Value

.87%

Institutional Class

.86%

Fidelity Mid Cap Value Fund

Class A

1.15%

Class T

1.41%

Class B

1.89%

Class C

1.92%

Mid Cap Value

.84%

Institutional Class

.90%

Fidelity Large Cap Growth Fund

Class A

1.20%

Class T

1.48%

Class B

1.98%

Class C

1.96%

Large Cap Growth

.97%

Institutional Class

.83%

Fidelity Mid Cap Growth Fund

Class A

1.10%

Class T

1.37%

Class B

1.83%

Class C

1.85%

Mid Cap Growth

.78%

Institutional Class

.69%

Annual Report

Fidelity Large Cap Value Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the fund's dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table and the fund's returns do not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Large Cap Value

-4.39%

14.11%

7.82%

A From November 15, 2001.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Large Cap Value, a class of the fund, on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell 1000® Value Index performed over the same period.



Annual Report

Fidelity Large Cap Value Fund

Management's Discussion of Fund Performance

Comments from Bruce Dirks, Portfolio Manager of Fidelity® Large Cap Value Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the year, Large Cap Value returned -4.39%, which, while disappointing in absolute terms, was ahead of the -5.38% return of the Russell 1000® Value Index. During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among banks and real estate investment trusts (REITs). Favorable industry weightings within the weak financials and consumer discretionary sectors also added value, as did good stock picking in health care and pockets of information technology. Underweighting diversified financials giant Citigroup and domestic banking firm Wachovia added to relative performance, as both stocks fared poorly within an environment of deteriorating business fundamentals for financials. Among other strong contributors were Cummins, a diesel engine manufacturer, and CF Industries, an out-of-index company that produces fertilizer products. On the flip side, relative results were held back by unfavorable stock picks in consumer staples - especially among food retailers - energy and the transportation industry. Among the biggest detractors were Tyson Foods, a large processor of poultry and meat products, and YRC Worldwide, a transportation company. Underweighting such strong-performing energy stocks as Occidental Petroleum also hurt. Tyson, YRC and Occidental were no longer in the portfolio at period end.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Large Cap Value Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Exxon Mobil Corp.

4.3

4.5

AT&T, Inc.

3.8

3.8

General Electric Co.

3.4

3.0

ConocoPhillips

3.1

0.0

JPMorgan Chase & Co.

3.0

3.0

American International Group, Inc.

2.7

2.3

Chevron Corp.

2.7

2.5

Bank of America Corp.

2.4

2.4

Altria Group, Inc.

2.3

1.8

Verizon Communications, Inc.

2.2

2.3

29.9

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Financials

30.7

31.7

Energy

15.6

14.0

Industrials

10.6

10.7

Consumer Staples

8.5

7.3

Health Care

7.6

7.6

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.7%

Stocks 99.7%

Short-Term
Investments and
Net Other Assets 0.3%

Short-Term
Investments and
Net Other Assets 0.3%

* Foreign investments

5.3%

**Foreign investments

2.0%

Annual Report

Fidelity Large Cap Value Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.7%

Shares

Value

CONSUMER DISCRETIONARY - 7.3%

Automobiles - 0.5%

General Motors Corp. (d)

283,401

$ 8,023,082

Hotels, Restaurants & Leisure - 2.0%

McDonald's Corp.

557,500

29,854,125

Household Durables - 0.3%

Whirlpool Corp.

56,000

4,766,160

Internet & Catalog Retail - 0.3%

Expedia, Inc. (a)

198,600

4,571,772

Leisure Equipment & Products - 0.4%

Eastman Kodak Co.

261,400

5,209,702

Media - 1.2%

Liberty Media Corp. - Capital
Series A (a)

64,300

6,919,966

Viacom, Inc. Class B (non-vtg.) (a)

296,500

11,492,340

18,412,306

Multiline Retail - 0.7%

Macy's, Inc.

387,800

10,718,792

Specialty Retail - 0.8%

Home Depot, Inc.

414,300

12,706,581

Textiles, Apparel & Luxury Goods - 1.1%

NIKE, Inc. Class B (d)

255,700

15,792,032

TOTAL CONSUMER DISCRETIONARY

110,054,552

CONSUMER STAPLES - 8.5%

Beverages - 1.1%

Molson Coors Brewing Co. Class B

392,400

17,528,508

Food & Staples Retailing - 2.9%

BJ's Wholesale Club, Inc. (a)

147,100

4,771,924

Kroger Co.

621,500

15,817,175

Safeway, Inc.

267,700

8,296,023

SUPERVALU, Inc.

484,600

14,567,076

43,452,198

Food Products - 0.3%

Del Monte Foods Co.

459,500

4,121,715

Household Products - 1.9%

Procter & Gamble Co.

433,300

28,576,135

Tobacco - 2.3%

Altria Group, Inc.

450,000

34,119,000

TOTAL CONSUMER STAPLES

127,797,556

ENERGY - 15.6%

Energy Equipment & Services - 1.9%

National Oilwell Varco, Inc. (a)

232,500

14,003,475

Transocean, Inc. (a)

112,200

13,755,720

27,759,195

Oil, Gas & Consumable Fuels - 13.7%

Chevron Corp.

482,300

40,754,350

ConocoPhillips

574,900

46,175,968

Shares

Value

Exxon Mobil Corp.

744,700

$ 64,342,081

Hess Corp.

265,100

24,079,033

Marathon Oil Corp.

308,900

14,471,965

Valero Energy Corp.

274,801

16,265,471

206,088,868

TOTAL ENERGY

233,848,063

FINANCIALS - 30.7%

Capital Markets - 6.5%

GLG Partners, Inc. (a)(d)

1,247,500

14,708,025

Goldman Sachs Group, Inc.

143,200

28,750,264

Janus Capital Group, Inc. (d)

575,300

15,538,853

Lehman Brothers Holdings, Inc. (d)

459,300

29,473,281

State Street Corp.

120,500

9,895,460

98,365,883

Commercial Banks - 3.8%

Associated Banc-Corp.

181,900

5,125,942

PNC Financial Services Group, Inc.

113,200

7,428,184

Regions Financial Corp. (d)

222,500

5,615,900

Wachovia Corp.

507,800

19,768,654

Wells Fargo & Co.

359,100

12,212,991

Zions Bancorp

132,200

7,236,628

57,388,299

Consumer Finance - 0.9%

Capital One Financial Corp. (d)

252,700

13,850,487

Diversified Financial Services - 6.7%

Bank of America Corp.

813,700

36,087,595

Citigroup, Inc.

546,460

15,421,101

JPMorgan Chase & Co.

947,400

45,048,870

MSCI, Inc. Class A

115,197

3,798,045

100,355,611

Insurance - 10.5%

ACE Ltd.

281,000

16,393,540

American International Group, Inc.

742,300

40,945,268

Assurant, Inc.

142,600

9,253,314

Axis Capital Holdings Ltd.

226,400

9,065,056

Endurance Specialty Holdings Ltd.

252,800

10,243,456

Loews Corp.

357,400

16,687,006

PartnerRe Ltd.

157,400

12,478,672

Prudential Financial, Inc.

255,600

21,564,972

RenaissanceRe Holdings Ltd.

215,900

12,304,141

W.R. Berkley Corp.

284,000

8,593,840

157,529,265

Real Estate Investment Trusts - 2.0%

Annaly Capital Management, Inc.

716,400

14,127,408

Boston Properties, Inc.

77,000

7,077,840

ProLogis Trust

91,600

5,436,460

Public Storage

46,700

3,654,275

30,295,983

Common Stocks - continued

Shares

Value

FINANCIALS - continued

Thrifts & Mortgage Finance - 0.3%

Astoria Financial Corp.

167,400

$ 4,549,932

TOTAL FINANCIALS

462,335,460

HEALTH CARE - 7.6%

Biotechnology - 1.0%

Biogen Idec, Inc. (a)

239,600

14,603,620

Health Care Equipment & Supplies - 0.5%

Baxter International, Inc.

118,200

7,179,468

Health Care Providers & Services - 1.8%

Humana, Inc. (a)

169,900

13,642,970

Medco Health Solutions, Inc. (a)

275,400

13,792,032

27,435,002

Life Sciences Tools & Services - 1.4%

Charles River Laboratories International, Inc. (a)

160,000

9,936,000

Invitrogen Corp. (a)

127,800

10,948,626

20,884,626

Pharmaceuticals - 2.9%

Johnson & Johnson

349,000

22,077,740

Pfizer, Inc.

960,700

22,470,773

44,548,513

TOTAL HEALTH CARE

114,651,229

INDUSTRIALS - 10.6%

Aerospace & Defense - 3.2%

Honeywell International, Inc.

94,300

5,570,301

Northrop Grumman Corp.

165,400

13,126,144

Raytheon Co.

170,600

11,112,884

The Boeing Co.

212,800

17,700,704

47,510,033

Airlines - 0.5%

Northwest Airlines Corp. (a)

193,200

3,614,772

UAL Corp.

121,400

4,607,130

8,221,902

Commercial Services & Supplies - 0.6%

Allied Waste Industries, Inc. (a)

920,000

9,062,000

Construction & Engineering - 1.4%

KBR, Inc. (a)

488,600

15,434,874

URS Corp. (a)

133,200

5,847,480

21,282,354

Industrial Conglomerates - 3.4%

General Electric Co.

1,424,000

50,423,840

Machinery - 1.2%

AGCO Corp. (a)

127,000

7,647,940

Cummins, Inc.

220,900

10,665,052

18,312,992

Shares

Value

Road & Rail - 0.3%

Hertz Global Holdings, Inc. (a)

266,933

$ 3,982,640

TOTAL INDUSTRIALS

158,795,761

INFORMATION TECHNOLOGY - 3.1%

Computers & Peripherals - 1.6%

Hewlett-Packard Co.

147,600

6,457,500

International Business Machines Corp.

124,200

13,331,628

Western Digital Corp. (a)

176,300

4,663,135

24,452,263

Electronic Equipment & Instruments - 0.8%

Tyco Electronics Ltd.

337,100

11,397,351

IT Services - 0.5%

Accenture Ltd. Class A

215,500

7,460,610

Software - 0.2%

Oracle Corp. (a)

172,000

3,534,600

TOTAL INFORMATION TECHNOLOGY

46,844,824

MATERIALS - 4.1%

Chemicals - 2.2%

Celanese Corp. Class A

164,400

6,112,392

CF Industries Holdings, Inc.

76,500

8,180,145

Chemtura Corp.

1,255,300

8,410,510

The Mosaic Co. (a)

108,600

9,883,686

32,586,733

Containers & Packaging - 0.6%

Ball Corp.

91,600

4,203,524

Owens-Illinois, Inc. (a)

94,300

4,752,720

8,956,244

Metals & Mining - 1.3%

Freeport-McMoRan Copper & Gold, Inc. Class B

104,800

9,330,344

United States Steel Corp.

99,300

10,139,523

19,469,867

TOTAL MATERIALS

61,012,844

TELECOMMUNICATION SERVICES - 6.0%

Diversified Telecommunication Services - 6.0%

AT&T, Inc.

1,480,362

56,979,133

Verizon Communications, Inc.

842,100

32,707,164

89,686,297

UTILITIES - 6.2%

Electric Utilities - 1.7%

Entergy Corp.

112,000

12,116,160

PPL Corp.

179,000

8,756,680

Reliant Energy, Inc. (a)

255,100

5,425,977

26,298,817

Gas Utilities - 1.0%

Energen Corp.

244,700

15,391,630

Common Stocks - continued

Shares

Value

UTILITIES - continued

Independent Power Producers & Energy Traders - 2.4%

Constellation Energy Group, Inc.

92,200

$ 8,663,112

Mirant Corp. (a)

324,700

11,961,948

NRG Energy, Inc. (a)

382,800

14,772,252

35,397,312

Multi-Utilities - 1.1%

Public Service Enterprise Group, Inc.

170,100

16,329,600

TOTAL UTILITIES

93,417,359

TOTAL COMMON STOCKS

(Cost $1,476,386,384)

1,498,443,945

Money Market Funds - 5.2%

Fidelity Cash Central Fund, 3.79% (b)

9,464,505

9,464,505

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

69,571,507

69,571,507

TOTAL MONEY MARKET FUNDS

(Cost $79,036,012)

79,036,012

TOTAL INVESTMENT PORTFOLIO - 104.9%

(Cost $1,555,422,396)

1,577,479,957

NET OTHER ASSETS - (4.9)%

(74,027,771)

NET ASSETS - 100%

$ 1,503,452,186

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 625,964

Fidelity Securities Lending Cash Central Fund

89,576

Total

$ 715,540

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $42,711,239 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Value Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $69,644,477) - See accompanying schedule:

Unaffiliated issuers (cost $1,476,386,384)

$ 1,498,443,945

Fidelity Central Funds (cost $79,036,012)

79,036,012

Total Investments (cost $1,555,422,396)

$ 1,577,479,957

Receivable for investments sold

23,766,119

Receivable for fund shares sold

2,565,058

Dividends receivable

1,696,869

Distributions receivable from Fidelity Central Funds

33,811

Prepaid expenses

4,921

Total assets

1,605,546,735

Liabilities

Payable for investments purchased

$ 23,494,925

Payable for fund shares redeemed

7,847,405

Accrued management fee

720,873

Distribution fees payable

6,898

Other affiliated payables

363,497

Other payables and accrued expenses

89,444

Collateral on securities loaned, at value

69,571,507

Total liabilities

102,094,549

Net Assets

$ 1,503,452,186

Net Assets consist of:

Paid in capital

$ 1,523,198,617

Undistributed net investment income

4,864,259

Accumulated undistributed net realized gain (loss) on investments

(46,668,251)

Net unrealized appreciation (depreciation) on investments

22,057,561

Net Assets

$ 1,503,452,186

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering
Price

Class A:
Net Asset Value
and redemption price per share ($9,774,232 ÷ 721,934 shares)

$ 13.54

Maximum offering price per share (100/94.25 of $13.54)

$ 14.37

Class T:
Net Asset Value
and redemption price per share ($5,975,654 ÷ 441,693 shares)

$ 13.53

Maximum offering price per share (100/96.50 of $13.53)

$ 14.02

Class B:
Net Asset Value
and offering price per share ($1,859,809 ÷ 137,404 shares)A

$ 13.54

Class C:
Net Asset Value
and offering price per share ($1,208,181 ÷ 89,368 shares)A

$ 13.52

Large Cap Value:
Net Asset Value
, offering price and redemption price per share ($1,483,574,281 ÷ 109,324,284 shares)

$ 13.57

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($1,060,029 ÷ 78,261 shares)

$ 13.54

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Value Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 32,039,820

Interest

4,832

Income from Fidelity Central Funds (including $89,576 from security lending)

715,540

Total income

32,760,192

Expenses

Management fee
Basic fee

$ 9,024,987

Performance adjustment

288,595

Transfer agent fees

3,768,695

Distribution fees

73,728

Accounting and security lending fees

507,630

Custodian fees and expenses

37,007

Independent trustees' compensation

5,753

Registration fees

122,860

Audit

60,620

Legal

7,461

Interest

5,226

Miscellaneous

53,563

Total expenses before reductions

13,956,125

Expense reductions

(136,896)

13,819,229

Net investment income (loss)

18,940,963

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

18,026,031

Change in net unrealized appreciation (depreciation) on investment securities

(121,407,791)

Net gain (loss)

(103,381,760)

Net increase (decrease) in net assets resulting from operations

$ (84,440,797)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ 18,940,963

$ 9,973,926

Net realized gain (loss)

18,026,031

34,353,479

Change in net unrealized appreciation (depreciation)

(121,407,791)

98,674,962

Net increase (decrease) in net assets resulting from operations

(84,440,797)

143,002,367

Distributions to shareholders from net investment income

(14,286,599)

(10,401,399)

Distributions to shareholders from net realized gain

(90,733,678)

(14,021,724)

Total distributions

(105,020,277)

(24,423,123)

Share transactions - net increase (decrease)

320,162,513

685,040,997

Redemption fees

-

21,396

Total increase (decrease) in net assets

130,701,439

803,641,637

Net Assets

Beginning of period

1,372,750,747

569,109,110

End of period (including undistributed net investment income of $4,864,259 and undistributed net investment income
of $274,346, respectively)

$ 1,503,452,186

$ 1,372,750,747

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.12

Net realized and unrealized gain (loss)

(1.00)

Total from investment operations

(.88)

Distributions from net investment income

(.12)

Distributions from net realized gain

(.87)

Total distributions

(.99)

Net asset value, end of period

$ 13.54

Total Return B,C

(6.04)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.22% A

Expenses net of fee waivers, if any

1.22% A

Expenses net of all reductions

1.22% A

Net investment income (loss)

.81% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 9,774

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the sales charges.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.08

Net realized and unrealized gain (loss)

(1.01)

Total from investment operations

(.93)

Distributions from net investment income

(.08)

Distributions from net realized gain

(.87)

Total distributions

(.95)

Net asset value, end of period

$ 13.53

Total Return B,C

(6.34)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.47% A

Expenses net of fee waivers, if any

1.47% A

Expenses net of all reductions

1.47% A

Net investment income (loss)

.56% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 5,976

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the sales charges.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.01

Net realized and unrealized gain (loss)

(1.00)

Total from investment operations

(.99)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.87)

Total distributions

(.88)

Net asset value, end of period

$ 13.54

Total Return B,C

(6.74)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.99% A

Expenses net of fee waivers, if any

1.99% A

Expenses net of all reductions

1.99% A

Net investment income (loss)

.04% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,860

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the contingent deferred sales charge.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.01

Net realized and unrealized gain (loss)

(.98)

Total from investment operations

(.97)

Distributions from net investment income

(.05)

Distributions from net realized gain

(.87)

Total distributions

(.92)

Net asset value, end of period

$ 13.52

Total Return B,C

(6.61)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.94% A

Expenses net of fee waivers, if any

1.94% A

Expenses net of all reductions

1.94% A

Net investment income (loss)

.09% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,208

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the contingent deferred sales charge.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Large Cap Value

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 15.19

$ 13.62

$ 12.04

$ 10.64

$ 8.17

Income from Investment Operations

Net investment income (loss) B

.18

.16

.17

.09 E

.09

Net realized and unrealized gain (loss)

(.80)

1.80

1.87

1.47

2.47

Total from investment operations

(.62)

1.96

2.04

1.56

2.56

Distributions from net investment income

(.13)

(.13)

(.11)

(.05)

(.09)

Distributions from net realized gain

(.87)

(.26)

(.35)

(.11)

-

Total distributions

(1.00)

(.39)

(.46)

(.16)

(.09)

Redemption fees added to paid in capital B

-

- G,H

- G

- G

- G

Net asset value, end of period

$ 13.57

$ 15.19

$ 13.62

$ 12.04

$ 10.64

Total Return A

(4.39)%

14.63%

17.09%

14.68%

31.44%

Ratios to Average Net Assets C,F

Expenses before reductions

.86%

.89%

.89%

1.07%

1.45%

Expenses net of fee waivers, if any

.85%

.89%

.89%

1.07%

1.20%

Expenses net of all reductions

.85%

.89%

.84%

1.05%

1.18%

Net investment income (loss)

1.18%

1.10%

1.32%

.79% E

.99%

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,483,574

$ 1,372,751

$ 569,109

$ 177,004

$ 25,168

Portfolio turnover rate D

204%

164%

175%

170%

72%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been .59%.

F Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

G Amount represents less than $.01 per share.

H The redemption fee was eliminated during the year January 31, 2007.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.17

Net realized and unrealized gain (loss)

(1.02)

Total from investment operations

(.85)

Distributions from net investment income

(.15)

Distributions from net realized gain

(.87)

Total distributions

(1.02)

Net asset value, end of period

$ 13.54

Total Return B,C

(5.82)%

Ratios to Average Net Assets E,H

Expenses before reductions

.85% A

Expenses net of fee waivers, if any

.85% A

Expenses net of all reductions

.84% A

Net investment income (loss)

1.19% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,060

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Value Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the fund's dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table and the fund's returns do not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Mid Cap Value

-7.67%

15.63%

10.38%

A From November 15, 2001.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Mid Cap Value, a class of the fund, on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell Midcap® Value Index performed over the same period.



Annual Report

Fidelity Mid Cap Value Fund

Management's Discussion of Fund Performance

Comments from Bruce Dirks, Portfolio Manager of Fidelity® Mid Cap Value Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the year, Mid Cap Value returned -7.67%, which, while disappointing in absolute terms, was better than the -8.73% return of the Russell Midcap® Value Index. During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among bank and asset management companies. Favorable industry weightings within the weak financials and consumer discretionary sectors also added value. Good stock picking in materials, energy and the equipment/services group of health care also helped. Among the fund's strongest contributors for the period were CF Industries, an out-of-index company that produces fertilizer products; Cummins, a diesel engine manufacturer; Annaly Capital Management, a real estate investment trust that owned AAA-rated mortgages; and National Oilwell Varco, an energy services contractor. Conversely, relative results were held back by some unfavorable stock selection decisions in the utilities, consumer staples and consumer discretionary sectors. The fund's poorest performers included transportation firm YRC Worldwide; original equipment manufacturer TRW Automotive; and photographic imaging company Eastman Kodak. YRC and TRW were no longer held at period end.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Mid Cap Value Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Hess Corp.

2.5

2.3

Edison International

2.1

1.5

Everest Re Group Ltd.

1.8

0.0

Annaly Capital Management, Inc.

1.7

0.8

Boston Properties, Inc.

1.7

0.0

Questar Corp.

1.7

0.0

Axis Capital Holdings Ltd.

1.6

0.8

Assurant, Inc.

1.6

1.2

PartnerRe Ltd.

1.6

1.2

Energen Corp.

1.6

1.8

17.9

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Financials

30.0

29.5

Utilities

14.0

12.5

Consumer Discretionary

13.1

13.1

Industrials

10.4

10.7

Energy

8.1

7.0

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.7%

Stocks 99.4%

Short-Term
Investments and
Net Other Assets 0.3%

Short-Term
Investments and
Net Other Assets 0.6%

* Foreign investments

13.1%

**Foreign investments

4.2%

Annual Report

Fidelity Mid Cap Value Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.7%

Shares

Value

CONSUMER DISCRETIONARY - 13.1%

Diversified Consumer Services - 0.5%

Apollo Group, Inc. Class A (non-vtg.) (a)

50,800

$ 4,050,792

Hotels, Restaurants & Leisure - 1.1%

Burger King Holdings, Inc.

303,600

7,999,860

Household Durables - 2.3%

Leggett & Platt, Inc. (d)

296,500

5,639,430

Whirlpool Corp. (d)

133,500

11,362,185

17,001,615

Internet & Catalog Retail - 1.2%

Expedia, Inc. (a)

242,600

5,584,652

Liberty Media Corp. - Interactive
Series A (a)

239,500

3,810,445

9,395,097

Leisure Equipment & Products - 1.4%

Eastman Kodak Co. (d)

534,300

10,648,599

Media - 2.1%

E.W. Scripps Co. Class A

181,100

7,374,392

Liberty Media Corp. - Capital
Series A (a)

74,900

8,060,738

15,435,130

Specialty Retail - 2.6%

Abercrombie & Fitch Co. Class A

96,200

7,666,178

AutoZone, Inc. (a)

48,200

5,826,416

RadioShack Corp.

136,800

2,373,480

TJX Companies, Inc.

125,600

3,963,936

19,830,010

Textiles, Apparel & Luxury Goods - 1.9%

Crocs, Inc. (a)(d)

219,600

7,639,884

Jones Apparel Group, Inc. (d)

399,000

6,703,200

14,343,084

TOTAL CONSUMER DISCRETIONARY

98,704,187

CONSUMER STAPLES - 7.1%

Beverages - 1.9%

Coca-Cola Enterprises, Inc.

311,600

7,188,612

Molson Coors Brewing Co. Class B

168,900

7,544,763

14,733,375

Food & Staples Retailing - 2.8%

BJ's Wholesale Club, Inc. (a)(d)

170,600

5,534,264

Safeway, Inc.

259,000

8,026,410

SUPERVALU, Inc.

239,600

7,202,376

20,763,050

Food Products - 2.2%

Corn Products International, Inc.

136,600

4,617,080

Dean Foods Co.

158,400

4,435,200

Del Monte Foods Co.

316,300

2,837,211

Tyson Foods, Inc. Class A

308,000

4,389,000

16,278,491

Shares

Value

Personal Products - 0.2%

Estee Lauder Companies, Inc. Class A

44,100

$ 1,861,020

TOTAL CONSUMER STAPLES

53,635,936

ENERGY - 8.1%

Energy Equipment & Services - 4.8%

Helmerich & Payne, Inc.

206,500

8,098,930

Nabors Industries Ltd. (a)

152,100

4,140,162

National Oilwell Varco, Inc. (a)

121,000

7,287,830

Patterson-UTI Energy, Inc. (d)

450,800

8,826,664

Superior Energy Services, Inc. (a)

74,400

2,982,696

Tidewater, Inc.

81,800

4,332,128

35,668,410

Oil, Gas & Consumable Fuels - 3.3%

Frontline Ltd. (NY Shares)

137,300

5,844,861

Hess Corp. (d)

211,800

19,237,794

25,082,655

TOTAL ENERGY

60,751,065

FINANCIALS - 30.0%

Capital Markets - 3.0%

GLG Partners, Inc. (a)(d)

756,200

8,915,598

Janus Capital Group, Inc. (d)

352,500

9,521,025

MF Global Ltd.

151,200

4,543,560

22,980,183

Commercial Banks - 3.5%

Associated Banc-Corp.

277,700

7,825,586

BOK Financial Corp.

71,195

3,878,704

East West Bancorp, Inc. (d)

188,900

4,544,934

Synovus Financial Corp.

219,900

2,904,879

Zions Bancorp

125,800

6,886,292

26,040,395

Diversified Financial Services - 0.6%

MSCI, Inc. Class A

136,800

4,510,296

Insurance - 13.9%

ACE Ltd.

158,000

9,217,720

Allied World Assurance Co. Holdings Ltd.

147,800

7,039,714

Assurant, Inc.

187,200

12,147,408

Axis Capital Holdings Ltd.

308,900

12,368,356

CNA Financial Corp.

189,800

6,451,302

Endurance Specialty Holdings Ltd.

286,800

11,621,136

Everest Re Group Ltd.

133,100

13,534,939

PartnerRe Ltd.

152,100

12,058,488

RenaissanceRe Holdings Ltd.

197,800

11,272,622

W.R. Berkley Corp.

303,200

9,174,832

104,886,517

Real Estate Investment Trusts - 7.0%

Annaly Capital Management, Inc.

658,200

12,979,704

Boston Properties, Inc.

138,600

12,740,112

Common Stocks - continued

Shares

Value

FINANCIALS - continued

Real Estate Investment Trusts - continued

Colonial Properties Trust (SBI)

97,800

$ 2,409,792

Kimco Realty Corp.

198,300

7,101,123

ProLogis Trust

155,700

9,240,795

Public Storage

73,700

5,767,025

Taubman Centers, Inc.

47,400

2,377,110

52,615,661

Thrifts & Mortgage Finance - 2.0%

Astoria Financial Corp.

262,500

7,134,750

Washington Federal, Inc.

323,700

7,904,754

15,039,504

TOTAL FINANCIALS

226,072,556

HEALTH CARE - 2.3%

Health Care Equipment & Supplies - 0.3%

Kinetic Concepts, Inc. (a)(d)

41,100

2,045,958

Health Care Providers & Services - 0.5%

Humana, Inc. (a)

42,500

3,412,750

Life Sciences Tools & Services - 1.2%

Charles River Laboratories International, Inc. (a)

74,200

4,607,820

Invitrogen Corp. (a)

54,600

4,677,582

9,285,402

Pharmaceuticals - 0.3%

Watson Pharmaceuticals, Inc. (a)

85,000

2,219,350

TOTAL HEALTH CARE

16,963,460

INDUSTRIALS - 10.4%

Aerospace & Defense - 1.7%

DRS Technologies, Inc.

88,400

4,744,428

L-3 Communications Holdings, Inc.

72,500

8,035,175

12,779,603

Airlines - 0.8%

Northwest Airlines Corp. (a)

152,600

2,855,146

UAL Corp.

75,800

2,876,610

5,731,756

Building Products - 0.6%

Lennox International, Inc.

126,800

4,711,888

Commercial Services & Supplies - 2.2%

Allied Waste Industries, Inc. (a)

700,300

6,897,955

R.R. Donnelley & Sons Co.

267,200

9,322,608

16,220,563

Construction & Engineering - 1.7%

KBR, Inc. (a)

309,000

9,761,310

URS Corp. (a)

69,800

3,064,220

12,825,530

Shares

Value

Industrial Conglomerates - 0.6%

McDermott International, Inc. (a)

94,500

$ 4,458,510

Machinery - 2.5%

AGCO Corp. (a)

106,000

6,383,320

Cummins, Inc.

117,300

5,663,244

Ingersoll-Rand Co. Ltd. Class A

168,400

6,655,168

18,701,732

Road & Rail - 0.3%

Hertz Global Holdings, Inc. (a)

160,200

2,390,184

TOTAL INDUSTRIALS

77,819,766

INFORMATION TECHNOLOGY - 6.4%

Communications Equipment - 0.7%

Juniper Networks, Inc. (a)

195,900

5,318,685

Computers & Peripherals - 2.2%

NCR Corp. (a)

422,600

9,077,448

Western Digital Corp. (a)

278,300

7,361,035

16,438,483

Electronic Equipment & Instruments - 1.4%

Avnet, Inc. (a)

152,500

5,430,525

Ingram Micro, Inc. Class A (a)

173,400

3,083,052

Jabil Circuit, Inc.

174,200

2,308,150

10,821,727

Semiconductors & Semiconductor Equipment - 1.6%

Atmel Corp. (a)

1,316,800

4,161,088

MEMC Electronic Materials, Inc. (a)

25,000

1,786,500

Novellus Systems, Inc. (a)

111,700

2,653,992

Varian Semiconductor Equipment Associates, Inc. (a)

109,100

3,514,111

12,115,691

Software - 0.5%

Compuware Corp. (a)

405,900

3,450,150

TOTAL INFORMATION TECHNOLOGY

48,144,736

MATERIALS - 6.8%

Chemicals - 3.1%

Celanese Corp. Class A

146,800

5,458,024

CF Industries Holdings, Inc.

57,100

6,105,703

Chemtura Corp. (d)

867,400

5,811,580

The Mosaic Co. (a)

71,200

6,479,912

23,855,219

Containers & Packaging - 1.9%

Ball Corp.

85,100

3,905,239

Owens-Illinois, Inc. (a)

91,500

4,611,600

Smurfit-Stone Container Corp. (a)(d)

603,500

5,727,215

14,244,054

Common Stocks - continued

Shares

Value

MATERIALS - continued

Metals & Mining - 1.8%

Cleveland-Cliffs, Inc. (d)

42,700

$ 4,348,568

United States Steel Corp.

88,000

8,985,680

13,334,248

TOTAL MATERIALS

51,433,521

TELECOMMUNICATION SERVICES - 1.5%

Diversified Telecommunication Services - 1.5%

Embarq Corp.

116,500

5,277,450

Qwest Communications International, Inc.

1,043,800

6,137,544

11,414,994

UTILITIES - 14.0%

Electric Utilities - 4.6%

Allegheny Energy, Inc.

123,300

6,755,607

Edison International

305,600

15,940,096

Reliant Energy, Inc. (a)

558,600

11,881,422

34,577,125

Gas Utilities - 3.3%

Energen Corp.

191,700

12,057,930

Questar Corp.

247,300

12,590,043

24,647,973

Independent Power Producers & Energy Traders - 4.2%

Constellation Energy Group, Inc.

103,700

9,743,652

Mirant Corp. (a)

315,100

11,608,284

NRG Energy, Inc. (a)

268,200

10,349,838

31,701,774

Multi-Utilities - 1.9%

CMS Energy Corp.

583,000

9,135,610

OGE Energy Corp.

162,500

5,318,625

14,454,235

TOTAL UTILITIES

105,381,107

TOTAL COMMON STOCKS

(Cost $760,656,463)

750,321,328

Money Market Funds - 10.6%

Shares

Value

Fidelity Cash Central Fund, 3.79% (b)

5,496,422

$ 5,496,422

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

74,160,825

74,160,825

TOTAL MONEY MARKET FUNDS

(Cost $79,657,247)

79,657,247

TOTAL INVESTMENT PORTFOLIO - 110.3%

(Cost $840,313,710)

829,978,575

NET OTHER ASSETS - (10.3)%

(77,171,615)

NET ASSETS - 100%

$ 752,806,960

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 394,997

Fidelity Securities Lending Cash Central Fund

169,409

Total

$ 564,406

Other Information

Distribution of investments by country of issue, as a percentage of total net assets, is as follows:

United States of America

86.9%

Bermuda

11.3%

Cayman Islands

1.2%

Others (individually less than 1%)

0.6%

100.0%

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $31,051,104 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Value Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $73,863,006) - See accompanying schedule:

Unaffiliated issuers (cost $760,656,463)

$ 750,321,328

Fidelity Central Funds (cost $79,657,247)

79,657,247

Total Investments (cost $840,313,710)

$ 829,978,575

Receivable for investments sold

20,806,082

Receivable for fund shares sold

2,652,227

Dividends receivable

223,102

Distributions receivable from Fidelity Central Funds

36,747

Prepaid expenses

2,661

Other receivables

46

Total assets

853,699,440

Liabilities

Payable for investments purchased

$ 20,180,467

Payable for fund shares redeemed

5,956,855

Accrued management fee

312,903

Distribution fees payable

5,417

Other affiliated payables

203,019

Other payables and accrued expenses

72,994

Collateral on securities loaned, at value

74,160,825

Total liabilities

100,892,480

Net Assets

$ 752,806,960

Net Assets consist of:

Paid in capital

$ 788,210,259

Undistributed net investment income

1,465,714

Accumulated undistributed net realized gain (loss) on investments

(26,533,878)

Net unrealized appreciation (depreciation) on investments

(10,335,135)

Net Assets

$ 752,806,960

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($7,444,673 ÷ 494,661 shares)

$ 15.05

Maximum offering price per share (100/94.25 of $15.05)

$ 15.97

Class T:
Net Asset Value
and redemption price per share ($3,714,323 ÷ 247,029 shares)

$ 15.04

Maximum offering price per share (100/96.50 of $15.04)

$ 15.59

Class B:
Net Asset Value
and offering price per share ($1,304,436 ÷ 87,007 shares)A

$ 14.99

Class C:
Net Asset Value
and offering price per share ($1,657,889 ÷ 110,680 shares)A

$ 14.98

Mid Cap Value:
Net Asset Value
, offering price and redemption price per share ($737,234,029 ÷ 48,847,921 shares)

$ 15.09

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($1,451,610 ÷ 96,411 shares)

$ 15.06

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Value Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 11,189,048

Interest

8,457

Income from Fidelity Central Funds (including $169,409 from security lending)

564,406

Total income

11,761,911

Expenses

Management fee
Basic fee

$ 5,084,079

Performance adjustment

(317,451)

Transfer agent fees

2,236,010

Distribution fees

53,691

Accounting and security lending fees

315,091

Custodian fees and expenses

28,074

Independent trustees' compensation

3,227

Registration fees

145,678

Audit

58,698

Legal

4,195

Interest

4,946

Miscellaneous

29,800

Total expenses before reductions

7,646,038

Expense reductions

(105,134)

7,540,904

Net investment income (loss)

4,221,007

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

(3,216,421)

Change in net unrealized appreciation (depreciation) on investment securities

(85,150,820)

Net gain (loss)

(88,367,241)

Net increase (decrease) in net assets resulting from operations

$ (84,146,234)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ 4,221,007

$ 2,688,685

Net realized gain (loss)

(3,216,421)

24,851,516

Change in net unrealized appreciation (depreciation)

(85,150,820)

43,131,851

Net increase (decrease) in net assets resulting from operations

(84,146,234)

70,672,052

Distributions to shareholders from net investment income

(3,037,015)

(2,949,768)

Distributions to shareholders from net realized gain

(37,140,454)

(13,997,322)

Total distributions

(40,177,469)

(16,947,090)

Share transactions - net increase (decrease)

198,288,449

259,218,977

Redemption fees

47,852

33,264

Total increase (decrease) in net assets

74,012,598

312,977,203

Net Assets

Beginning of period

678,794,362

365,817,159

End of period (including undistributed net investment income of $1,465,714 and undistributed net investment income
of $375,291, respectively)

$ 752,806,960

$ 678,794,362

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

.03

Net realized and unrealized gain (loss)

(1.78)

Total from investment operations

(1.75)

Distributions from net investment income

(.06)

Distributions from net realized gain

(.77)

Total distributions

(.83)

Net asset value, end of period

$ 15.05

Total Return B,C,D

(10.28)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.14% A

Expenses net of fee waivers, if any

1.14% A

Expenses net of all reductions

1.13% A

Net investment income (loss)

.16% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 7,445

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.02)

Net realized and unrealized gain (loss)

(1.76)

Total from investment operations

(1.78)

Distributions from net investment income

(.04)

Distributions from net realized gain

(.77)

Total distributions

(.81)

Net asset value, end of period

$ 15.04

Total Return B,C,D

(10.46)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.39% A

Expenses net of fee waivers, if any

1.39% A

Expenses net of all reductions

1.39% A

Net investment income (loss)

(.10)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 3,714

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.10)

Net realized and unrealized gain (loss)

(1.76)

Total from investment operations

(1.86)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.77)

Total distributions

(.78)

Net asset value, end of period

$ 14.99

Total Return B,C,D

(10.88)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.89% A

Expenses net of fee waivers, if any

1.89% A

Expenses net of all reductions

1.89% A

Net investment income (loss)

(.59)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,304

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.10)

Net realized and unrealized gain (loss)

(1.77)

Total from investment operations

(1.87)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.77)

Total distributions

(.78)

Net asset value, end of period

$ 14.98

Total Return B,C,D

(10.94)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.90% A

Expenses net of fee waivers, if any

1.90% A

Expenses net of all reductions

1.90% A

Net investment income (loss)

(.60)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,658

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Mid Cap Value

Years ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 17.18

$ 15.65

$ 14.14

$ 12.32

$ 8.85

Income from Investment Operations

Net investment income (loss) B

.08

.09

.16 E

.08

.06

Net realized and unrealized gain (loss)

(1.34)

1.98

2.59

2.10

3.45

Total from investment operations

(1.26)

2.07

2.75

2.18

3.51

Distributions from net investment income

(.06)

(.09)

(.10)

(.04)

(.04)

Distributions from net realized gain

(.77)

(.45)

(1.15)

(.32)

-

Total distributions

(.83)

(.54)

(1.24) H

(.36)

(.04)

Redemption fees added to paid in capital B,G

-

-

-

-

-

Net asset value, end of period

$ 15.09

$ 17.18

$ 15.65

$ 14.14

$ 12.32

Total Return A

(7.67)%

13.48%

19.97%

17.75%

39.69%

Ratios to Average Net Assets C,F

Expenses before reductions

.83%

.84%

.86%

.91%

1.07%

Expenses net of fee waivers, if any

.82%

.84%

.86%

.91%

1.07%

Expenses net of all reductions

.82%

.84%

.81%

.90%

1.05%

Net investment income (loss)

.47%

.56%

1.08% E

.59%

.55%

Supplemental Data

Net assets, end of period (000 omitted)

$ 737,234

$ 678,794

$ 365,817

$ 153,231

$ 95,797

Portfolio turnover rate D

264%

187%

207%

196%

97%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.04 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been .81%.

F Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

G Amount represents less than $.01 per share.

H Total distributions of $1.24 per share is comprised of distributions from net investment income of $0.95 and distributions from net realized gain of $1.145 per share.

Financial Highlights - Institutional Class

Years ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) D

.07

Net realized and unrealized gain (loss)

(1.78)

Total from investment operations

(1.71)

Distributions from net investment income

(.09)

Distributions from net realized gain

(.77)

Total distributions

(.86)

Net asset value, end of period

$ 15.06

Total Return B,C

(10.06)%

Ratios to Average Net Assets E,H

Expenses before reductions

.89% A

Expenses net of fee waivers, if any

.89% A

Expenses net of all reductions

.88% A

Net investment income (loss)

.41% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,452

Portfolio turnover rate F

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Growth Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the fund's dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table and the fund's returns do not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Large Cap Growth

-7.26%

10.76%

2.35%

A From November 15, 2001.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Large Cap Growth, a class of the fund, on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell 1000® Growth Index performed over the same period.



Annual Report

Fidelity Large Cap Growth Fund

Management's Discussion of Fund Performance

Comments from Edward Best, Portfolio Manager of Fidelity® Large Cap Growth Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the 12 months ending January 31, 2008, Large Cap Growth fell 7.26%, trailing the 0.51% return of the Russell 1000® Growth Index. Poor security selection in consumer discretionary, information technology and industrials accounted for the majority of the fund's underperformance compared with the index. Within consumer discretionary, the fund was hurt most by unproductive picks within the retailing and homebuilding segments. Consumer electronics retailer RadioShack and discount retailer Big Lots fell on investors' concerns about the companies' ability to weather recessionary concerns and a weak U.S. consumer. Pharmaceuticals company Sepracor suffered due to a change in Medicare reimbursement for one of its top-selling patented drugs. Airlines US Airways and AMR Corp. - parent company of American Airlines - underperformed due to increased pressure from rising energy prices. The fund's positions in homebuilders KB Home, Ryland Group, D.R. Horton and Lennar, all of which I sold when I took over the fund in June, also held back performance. While our overall stock picking within industrials detracted, stock selection within the sector's capital goods segment actually helped the fund. Specifically, Manitowoc, a major supplier of lifting equipment to the global construction industry, aided performance. Also within capital goods, defense contractor Lockheed Martin benefited from an increase in U.S. defense spending. Stock selection among health care names was positive as well. Pharmaceutical company Biogen Idec gained on news of the re-launch of its multiple sclerosis drug, Tysabri. While stock selection in technology hurt overall, there were some contributors. Apple advanced as the stock did well on the success of its iPod, iPhone and iMac products. Software developer Microsoft also aided performance. By the end of the period, I sold US Airways, AMR Corp., RadioShack, Big Lots, Sepracor, and Apple to take advantage of other opportunities in the market.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Large Cap Growth Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Microsoft Corp.

5.0

5.0

Exxon Mobil Corp.

4.4

1.6

Oracle Corp.

4.1

0.0

The Boeing Co.

3.9

0.0

Cisco Systems, Inc.

3.8

2.4

Bristol-Myers Squibb Co.

3.6

0.0

Accenture Ltd. Class A

3.3

3.3

NIKE, Inc. Class B

3.3

0.0

NYMEX Holdings, Inc.

3.1

0.0

Kroger Co.

3.0

3.1

37.5

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Information Technology

26.0

26.9

Health Care

16.8

16.2

Industrials

14.2

14.2

Consumer Discretionary

11.9

13.5

Consumer Staples

10.9

9.9

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.4%

Stocks 99.8%

Short-Term
Investments and
Net Other Assets 0.6%

Short-Term
Investments and
Net Other Assets 0.2%

* Foreign investments

4.4%

**Foreign
investments

5.5%

Annual Report

Fidelity Large Cap Growth Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.4%

Shares

Value

CONSUMER DISCRETIONARY - 11.9%

Auto Components - 1.6%

The Goodyear Tire & Rubber Co. (a)

99,300

$ 2,499,381

Diversified Consumer Services - 1.2%

Apollo Group, Inc. Class A (non-vtg.) (a)

22,600

1,802,124

Hotels, Restaurants & Leisure - 2.6%

McDonald's Corp.

74,200

3,973,410

Media - 2.3%

Regal Entertainment Group Class A (d)

189,300

3,509,622

Specialty Retail - 0.9%

Gamestop Corp. Class A (a)

27,400

1,417,402

Textiles, Apparel & Luxury Goods - 3.3%

NIKE, Inc. Class B

79,900

4,934,624

TOTAL CONSUMER DISCRETIONARY

18,136,563

CONSUMER STAPLES - 10.9%

Beverages - 5.0%

Pepsi Bottling Group, Inc.

114,200

3,979,870

PepsiCo, Inc.

52,500

3,579,975

7,559,845

Food & Staples Retailing - 3.0%

Kroger Co.

181,400

4,616,630

Food Products - 2.4%

Kellogg Co.

77,200

3,697,880

Household Products - 0.5%

Energizer Holdings, Inc. (a)

8,300

777,046

TOTAL CONSUMER STAPLES

16,651,401

ENERGY - 7.9%

Energy Equipment & Services - 0.9%

Baker Hughes, Inc.

5,700

370,101

Nabors Industries Ltd. (a)

24,400

664,168

Schlumberger Ltd. (NY Shares)

5,000

377,300

1,411,569

Oil, Gas & Consumable Fuels - 7.0%

Exxon Mobil Corp.

78,000

6,739,200

Sunoco, Inc.

13,000

808,600

Valero Energy Corp.

53,000

3,137,070

10,684,870

TOTAL ENERGY

12,096,439

FINANCIALS - 7.5%

Capital Markets - 4.4%

Charles Schwab Corp.

198,200

4,419,860

Northern Trust Corp.

30,900

2,266,824

6,686,684

Shares

Value

Diversified Financial Services - 3.1%

NYMEX Holdings, Inc. (d)

41,700

$ 4,795,500

TOTAL FINANCIALS

11,482,184

HEALTH CARE - 16.8%

Biotechnology - 1.2%

Biogen Idec, Inc. (a)

29,300

1,785,835

Health Care Equipment & Supplies - 2.8%

Baxter International, Inc.

69,700

4,233,578

Health Care Providers & Services - 9.2%

Aetna, Inc.

86,000

4,580,360

Express Scripts, Inc. (a)

11,400

769,386

Medco Health Solutions, Inc. (a)

35,200

1,762,816

UnitedHealth Group, Inc.

47,700

2,425,068

WellPoint, Inc. (a)

57,800

4,519,960

14,057,590

Pharmaceuticals - 3.6%

Bristol-Myers Squibb Co.

234,400

5,435,736

TOTAL HEALTH CARE

25,512,739

INDUSTRIALS - 14.2%

Aerospace & Defense - 5.6%

Lockheed Martin Corp.

5,900

636,728

Precision Castparts Corp.

16,900

1,923,220

The Boeing Co.

71,500

5,947,370

8,507,318

Commercial Services & Supplies - 3.9%

Allied Waste Industries, Inc. (a)

460,200

4,532,970

HNI Corp. (d)

15,400

518,364

Manpower, Inc.

15,300

860,778

5,912,112

Construction & Engineering - 2.7%

Fluor Corp.

7,500

912,525

Shaw Group, Inc. (a)

13,800

779,700

URS Corp. (a)

54,700

2,401,330

4,093,555

Industrial Conglomerates - 0.8%

McDermott International, Inc. (a)

24,500

1,155,910

Machinery - 1.2%

AGCO Corp. (a)

8,700

523,914

Cummins, Inc.

16,500

796,620

Manitowoc Co., Inc.

15,400

587,048

1,907,582

TOTAL INDUSTRIALS

21,576,477

INFORMATION TECHNOLOGY - 26.0%

Communications Equipment - 3.8%

Cisco Systems, Inc. (a)

236,300

5,789,350

Common Stocks - continued

Shares

Value

INFORMATION TECHNOLOGY - continued

Computers & Peripherals - 0.5%

Lexmark International, Inc. Class A (a)

1,400

$ 50,694

Western Digital Corp. (a)

29,000

767,050

817,744

Electronic Equipment & Instruments - 0.5%

National Instruments Corp.

27,400

735,964

IT Services - 6.2%

Accenture Ltd. Class A

143,600

4,971,432

Mastercard, Inc. Class A (d)

7,000

1,449,000

The Western Union Co.

135,200

3,028,480

9,448,912

Semiconductors & Semiconductor Equipment - 2.2%

MEMC Electronic Materials, Inc. (a)

12,500

893,250

NVIDIA Corp. (a)

82,500

2,028,675

Texas Instruments, Inc.

12,000

371,160

3,293,085

Software - 12.8%

Autodesk, Inc. (a)

76,100

3,131,515

BMC Software, Inc. (a)

74,200

2,377,368

Microsoft Corp.

233,200

7,602,320

Oracle Corp. (a)

306,900

6,306,795

19,417,998

TOTAL INFORMATION TECHNOLOGY

39,503,053

MATERIALS - 3.0%

Chemicals - 2.0%

Air Products & Chemicals, Inc.

4,200

378,084

E.I. du Pont de Nemours & Co.

10,100

456,318

Monsanto Co.

12,800

1,439,232

Praxair, Inc.

4,700

380,277

The Mosaic Co. (a)

4,000

364,040

3,017,951

Metals & Mining - 1.0%

Freeport-McMoRan Copper & Gold, Inc. Class B

8,400

747,852

Newmont Mining Corp.

7,100

385,814

Southern Copper Corp.

4,400

412,896

1,546,562

TOTAL MATERIALS

4,564,513

Shares

Value

TELECOMMUNICATION SERVICES - 0.3%

Wireless Telecommunication Services - 0.3%

Crown Castle International Corp. (a)

10,700

$ 387,233

UTILITIES - 0.9%

Electric Utilities - 0.9%

Exelon Corp.

18,300

1,394,277

TOTAL COMMON STOCKS

(Cost $156,064,347)

151,304,879

Money Market Funds - 6.4%

Fidelity Cash Central Fund, 3.79% (b)

1,837,290

1,837,290

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

7,887,500

7,887,500

TOTAL MONEY MARKET FUNDS

(Cost $9,724,790)

9,724,790

TOTAL INVESTMENT PORTFOLIO - 105.8%

(Cost $165,789,137)

161,029,669

NET OTHER ASSETS - (5.8)%

(8,892,495)

NET ASSETS - 100%

$ 152,137,174

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Includes investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 60,857

Fidelity Securities Lending Cash Central Fund

56,422

Total

$ 117,279

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $7,837,041 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Growth Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $7,883,516) - See accompanying schedule:

Unaffiliated issuers (cost $156,064,347)

$ 151,304,879

Fidelity Central Funds (cost $9,724,790)

9,724,790

Total Investments (cost $165,789,137)

$ 161,029,669

Receivable for investments sold

18,559,475

Receivable for fund shares sold

222,238

Dividends receivable

82,079

Distributions receivable from Fidelity Central Funds

4,743

Prepaid expenses

514

Receivable from investment adviser for expense reductions

25,900

Total assets

179,924,618

Liabilities

Payable for investments purchased

$ 19,486,946

Payable for fund shares redeemed

257,786

Accrued management fee

52,051

Distribution fees payable

1,918

Other affiliated payables

48,536

Other payables and accrued expenses

52,707

Collateral on securities loaned, at value

7,887,500

Total liabilities

27,787,444

Net Assets

$ 152,137,174

Net Assets consist of:

Paid in capital

$ 162,808,253

Accumulated undistributed net realized gain (loss) on investments

(5,911,611)

Net unrealized appreciation (depreciation) on investments

(4,759,468)

Net Assets

$ 152,137,174

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($1,302,425 ÷ 132,172 shares)

$ 9.85

Maximum offering price per share (100/94.25 of $9.85)

$ 10.45

Class T:
Net Asset Value
and redemption price per share ($1,097,491 ÷ 111,448 shares)

$ 9.85

Maximum offering price per share (100/96.50 of $9.85)

$ 10.21

Class B:
Net Asset Value
and offering price per share ($543,149 ÷ 55,237 shares)A

$ 9.83

Class C:
Net Asset Value
and offering price per share ($944,595 ÷ 96,233 shares)A

$ 9.82

Large Cap Growth:
Net Asset Value
, offering price and redemption price per share ($147,863,609 ÷ 14,947,307 shares)

$ 9.89

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($385,905 ÷ 39,068 shares)

$ 9.88

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Growth Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 1,475,584

Interest

922

Income from Fidelity Central Funds (including $56,422 from security lending)

117,279

Total income

1,593,785

Expenses

Management fee
Basic fee

$ 979,536

Performance adjustment

37,062

Transfer agent fees

534,768

Distribution fees

10,735

Accounting and security lending fees

70,624

Custodian fees and expenses

18,147

Independent trustees' compensation

638

Registration fees

103,527

Audit

55,247

Legal

928

Miscellaneous

11,842

Total expenses before reductions

1,823,054

Expense reductions

(97,947)

1,725,107

Net investment income (loss)

(131,322)

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

10,237,592

Change in net unrealized appreciation (depreciation) on investment securities

(21,747,840)

Net gain (loss)

(11,510,248)

Net increase (decrease) in net assets resulting from operations

$ (11,641,570)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ (131,322)

$ 26,323

Net realized gain (loss)

10,237,592

4,571,573

Change in net unrealized appreciation (depreciation)

(21,747,840)

(297,654)

Net increase (decrease) in net assets resulting from operations

(11,641,570)

4,300,242

Distributions to shareholders from net investment income

-

(137,207)

Distributions to shareholders from net realized gain

(18,572,404)

(3,644,497)

Total distributions

(18,572,404)

(3,781,704)

Share transactions - net increase (decrease)

(1,164,336)

25,478,221

Redemption fees

-

5,257

Total increase (decrease) in net assets

(31,378,310)

26,002,016

Net Assets

Beginning of period

183,515,484

157,513,468

End of period

$ 152,137,174

$ 183,515,484

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.03)

Net realized and unrealized gain (loss)

(.72)

Total from investment operations

(.75)

Distributions from net realized gain

(1.25)

Net asset value, end of period

$ 9.85

Total Return B,C,D

(6.99)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.20% A

Expenses net of fee waivers, if any

1.20% A

Expenses net of all reductions

1.20% A

Net investment income (loss)

(.29)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,302

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.06)

Net realized and unrealized gain (loss)

(.69)

Total from investment operations

(.75)

Distributions from net realized gain

(1.25)

Net asset value, end of period

$ 9.85

Total Return B,C,D

(7.05)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.47% A

Expenses net of fee waivers, if any

1.47% A

Expenses net of all reductions

1.47% A

Net investment income (loss)

(.56)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,097

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.12)

Net realized and unrealized gain (loss)

(.70)

Total from investment operations

(.82)

Distributions from net realized gain

(1.20)

Net asset value, end of period

$ 9.83

Total Return B,C,D

(7.62)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.99% A

Expenses net of fee waivers, if any

1.99% A

Expenses net of all reductions

1.99% A

Net investment income (loss)

(1.07)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 543

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.11)

Net realized and unrealized gain (loss)

(.70)

Total from investment operations

(.81)

Distributions from net realized gain

(1.22)

Net asset value, end of period

$ 9.82

Total Return B,C,D

(7.54)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.96% A

Expenses net of fee waivers, if any

1.96% A

Expenses net of all reductions

1.96% A

Net investment income (loss)

(1.05)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 945

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Large Cap Growth

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 11.92

$ 11.82

$ 10.17

$ 9.21

$ 6.93

Income from Investment Operations

Net investment income (loss) B

(.01)

- H

.02 E

(.01) F

(.01)

Net realized and unrealized gain (loss)

(.77)

.37

1.87

.97

2.29

Total from investment operations

(.78)

.37

1.89

.96

2.28

Distributions from net investment income

-

(.01)

-

-

-

Distributions from net realized gain

(1.25)

(.26)

(.24)

-

-

Total distributions

(1.25)

(.27)

(.24)

-

-

Redemption fees added to paid in capital B

-

- H,I

- H

- H

- H

Net asset value, end of period

$ 9.89

$ 11.92

$ 11.82

$ 10.17

$ 9.21

Total Return A

(7.26)%

3.20%

18.66%

10.42%

32.90%

Ratios to Average Net Assets C,G

Expenses before reductions

1.03%

1.10%

1.12%

1.30%

1.53%

Expenses net of fee waivers, if any

.99%

1.00%

1.00%

1.20%

1.20%

Expenses net of all reductions

.98%

.99%

.94%

1.13%

1.18%

Net investment income (loss)

(.07)%

.02%

.15% E

(.07)% F

(.15)%

Supplemental Data

Net assets, end of period (000 omitted)

$ 147,864

$ 183,515

$ 157,513

$ 49,453

$ 23,079

Portfolio turnover rate D

428%

189%

268%

274%

81%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.07)%.

F Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.32)%.

G Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

H Amount represents less than $.01 per share.

I The redemption fee was eliminated during the year ended January 31, 2007.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net realized and unrealized gain (loss)

(.70)

Distributions from net realized gain

(1.27)

Net asset value, end of period

$ 9.88

Total Return B,C

(6.64)%

Ratios to Average Net Assets D,E

Expenses before reductions

.88% A

Expenses net of fee waivers, if any

.88% A

Expenses net of all reductions

.88% A

Net investment income (loss)

.03% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 386

Portfolio turnover rate F

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Growth Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the fund's dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table and the fund's returns do not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Mid Cap Growth

-9.68%

13.19%

5.08%

A From November 15, 2001.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Mid Cap Growth, a class of the fund, on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell Midcap® Growth Index performed over the same period.



Annual Report

Fidelity Mid Cap Growth Fund

Management's Discussion of Fund Performance

Comments from Patrick Venanzi, who became Portfolio Manager of Fidelity® Mid Cap Growth Fund on January 4, 2008

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the 12 months ending January 31, 2008, the fund fell 9.68%, compared with a return of -1.03% for the Russell Midcap® Growth Index. Poor stock selection in consumer discretionary, industrials, materials and consumer staples hurt the most. While the fund struggled to keep pace with the index for most of the period, stock selection in energy and health care helped limit some of the downside. Within industrials, the fund's overweighting in transportation stocks, specifically in the airlines, detracted from performance. Our holdings in US Airways and AMR Corp., the parent company of American Airlines, suffered from rising energy costs. Overweighted holdings in the consumer durables and apparel segment - primarily the homebuilders - also took a bite out of fund performance. D.R. Horton, Ryland Group, KB Home and Lennar were particularly hard hit as the U.S. residential real estate market continued to slow down. I sold D.R. Horton, Ryland Group and Lennar. On the positive side, energy services company National Oilwell Varco was the fund's biggest contributor. Another contributor was the fund's out-of-index position in consumer electronics giant Apple, which continued to gain from strong sales of its iPod, iPhone and iMac product lines. I sold the fund's position in Apple to lock in profits. Heavy-lifting equipment and crane manufacturer Manitowoc advanced due to the continuing worldwide build-out of commercial and industrial infrastructure. I sold this position by the end of the period. BE Aerospace, which makes products for aircraft cabin interiors, also contributed, as investors expressed confidence in the company's new contracts to supply airline carriers that are upgrading their fleets. Lastly, MEMC Electronic Materials, which produces silicon wafers, helped as well.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Mid Cap Growth Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

PPL Corp.

1.4

0.7

Southwestern Energy Co.

1.4

0.3

Constellation Energy Group, Inc.

1.3

1.0

National Oilwell Varco, Inc.

1.3

3.9

Align Technology, Inc.

1.1

0.0

Centex Corp.

1.1

0.0

TranS1, Inc.

1.0

0.0

Polycom, Inc.

0.9

0.0

Intuitive Surgical, Inc.

0.9

0.0

Northwest Airlines Corp.

0.9

0.0

11.3

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Information Technology

19.8

19.0

Industrials

17.1

16.1

Health Care

16.6

12.9

Consumer Discretionary

14.1

17.6

Energy

11.6

10.9

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 100.2%

Stocks 97.8%

Short-Term
Investments and
Net Other Assets(dagger) (0.2)%

Short-Term
Investments and
Net Other Assets 2.2%

* Foreign
investments

11.7%

**Foreign investments

2.1%

(dagger) Short-Term Investments
and Net Other Assets
are not included in the pie chart.

Annual Report

Fidelity Mid Cap Growth Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 100.2%

Shares

Value

CONSUMER DISCRETIONARY - 14.1%

Automobiles - 0.3%

Renault SA

7,300

$ 833,337

Diversified Consumer Services - 1.1%

American Public Education, Inc.

7,200

283,752

Apollo Group, Inc. Class A (non-vtg.) (a)

8,200

653,868

DeVry, Inc.

19,000

1,048,610

Regis Corp.

56,400

1,428,612

3,414,842

Hotels, Restaurants & Leisure - 2.1%

Bally Technologies, Inc. (a)

20,200

962,328

Boyd Gaming Corp.

66,400

1,774,872

Buffalo Wild Wings, Inc. (a)

8,400

211,428

Burger King Holdings, Inc.

54,789

1,443,690

Gaylord Entertainment Co. (a)

29,700

866,943

Papa John's International, Inc. (a)

26,100

660,069

Peet's Coffee & Tea, Inc. (a)

7,000

153,580

Starwood Hotels & Resorts Worldwide, Inc.

5,400

244,350

6,317,260

Household Durables - 2.1%

Centex Corp. (d)

116,700

3,241,926

KB Home

100

2,750

Newell Rubbermaid, Inc.

15,100

364,212

Pulte Homes, Inc.

33,500

547,390

Whirlpool Corp.

25,700

2,187,327

6,343,605

Internet & Catalog Retail - 1.2%

Blue Nile, Inc. (a)

17,400

961,350

Expedia, Inc. (a)

10,600

244,012

Priceline.com, Inc. (a)

21,099

2,289,663

3,495,025

Leisure Equipment & Products - 0.7%

Hasbro, Inc.

12,400

322,028

Leapfrog Enterprises, Inc. Class A (a)

36,300

236,313

Mattel, Inc.

53,533

1,124,728

Nikon Corp.

14,000

381,830

2,064,899

Media - 2.7%

Cablevision Systems Corp. - NY Group Class A (a)

30,300

711,444

DISH Network Corp. Class A (a)

13,057

368,730

E.W. Scripps Co. Class A

35,800

1,457,776

Grupo Televisa SA de CV (CPO) sponsored ADR

94,000

2,095,260

Regal Entertainment Group Class A

87,100

1,614,834

SR Teleperformance SA

44,300

1,386,369

The Walt Disney Co.

11,700

350,181

Time Warner, Inc.

21,600

339,984

8,324,578

Shares

Value

Multiline Retail - 0.1%

Dollar Tree Stores, Inc. (a)

14,300

$ 400,543

Specialty Retail - 3.2%

Abercrombie & Fitch Co. Class A

5,100

406,419

Advance Auto Parts, Inc.

30,500

1,088,240

Asbury Automotive Group, Inc.

13,100

185,758

AutoZone, Inc. (a)

8,800

1,063,744

Gamestop Corp. Class A (a)

5,400

279,342

Group 1 Automotive, Inc.

8,700

230,028

Home Depot, Inc.

19,700

604,199

Lumber Liquidators, Inc.

3,400

30,260

OfficeMax, Inc.

49,800

1,233,546

Ross Stores, Inc.

36,300

1,058,145

Tween Brands, Inc. (a)(d)

44,700

1,431,741

Urban Outfitters, Inc. (a)

28,800

835,200

Williams-Sonoma, Inc.

49,500

1,330,560

Zumiez, Inc. (a)

3,500

67,305

9,844,487

Textiles, Apparel & Luxury Goods - 0.6%

Coach, Inc. (a)

13,000

416,650

Deckers Outdoor Corp. (a)

2,800

339,472

Lululemon Athletica, Inc.

15,700

532,073

Under Armour, Inc. Class A (sub. vtg.) (a)

16,000

644,000

1,932,195

TOTAL CONSUMER DISCRETIONARY

42,970,771

CONSUMER STAPLES - 3.4%

Beverages - 0.1%

Coca-Cola Femsa SA de CV sponsored ADR

7,200

338,040

Food & Staples Retailing - 1.3%

CVS Caremark Corp.

14,600

570,422

Performance Food Group Co. (a)

48,000

1,518,240

Safeway, Inc.

25,200

780,948

Sysco Corp.

42,100

1,223,005

4,092,615

Food Products - 1.5%

Campbell Soup Co.

17,300

546,853

Marine Harvest ASA (a)

2,600,000

1,400,493

McCormick & Co., Inc. (non-vtg.)

55,300

1,864,716

Seaboard Corp.

370

475,543

Wm. Wrigley Jr. Co.

3,000

172,290

4,459,895

Household Products - 0.4%

Energizer Holdings, Inc. (a)

13,300

1,245,146

Personal Products - 0.1%

Estee Lauder Companies, Inc. Class A

8,900

375,580

TOTAL CONSUMER STAPLES

10,511,276

Common Stocks - continued

Shares

Value

ENERGY - 11.6%

Energy Equipment & Services - 5.0%

Atwood Oceanics, Inc. (a)

21,000

$ 1,744,890

Cameron International Corp. (a)

31,700

1,276,242

Diamond Offshore Drilling, Inc.

16,700

1,885,931

Grey Wolf, Inc. (a)

297,600

1,773,696

Nabors Industries Ltd. (a)

35,000

952,700

National Oilwell Varco, Inc. (a)

65,120

3,922,178

North American Energy Partners, Inc. (a)

14,200

170,116

Ocean RIG ASA (a)(d)

114,500

787,337

Smith International, Inc.

23,900

1,295,619

Transocean, Inc. (a)

10,961

1,343,819

15,152,528

Oil, Gas & Consumable Fuels - 6.6%

Alpha Natural Resources, Inc. (a)

21,500

719,390

Arch Coal, Inc.

15,700

690,800

Boardwalk Pipeline Partners, LP

5,700

177,156

Canadian Natural Resources Ltd.

10,100

646,227

Chesapeake Energy Corp.

46,600

1,734,918

CONSOL Energy, Inc.

35,300

2,576,900

Copano Energy LLC

4,900

173,166

Denbury Resources, Inc. (a)

65,700

1,662,210

Energy Transfer Equity LP

5,200

171,652

Foundation Coal Holdings, Inc.

7,000

366,100

Frontier Oil Corp.

38,100

1,343,787

Holly Corp.

28,601

1,384,860

OPTI Canada, Inc. (a)

80,400

1,325,913

Petrobank Energy & Resources Ltd. (a)

7,000

351,761

Southwestern Energy Co. (a)

76,100

4,254,751

Tesoro Corp.

22,400

874,720

Valero Energy Corp.

100

5,919

Western Refining, Inc.

44,460

949,221

Williams Companies, Inc.

27,100

866,387

20,275,838

TOTAL ENERGY

35,428,366

FINANCIALS - 9.2%

Capital Markets - 2.6%

Franklin Resources, Inc.

23,600

2,459,828

KBW, Inc. (a)

6,300

187,173

Legg Mason, Inc.

14,500

1,044,000

MF Global Ltd.

73,200

2,199,660

SEI Investments Co.

67,400

1,866,980

T. Rowe Price Group, Inc.

3,300

166,947

7,924,588

Commercial Banks - 1.5%

Bank of Yokohama Ltd.

84,000

544,305

Chiba Bank Ltd.

52,000

384,877

Chuo Mitsui Trust Holdings, Inc.

55,000

377,598

Mitsubishi UFJ Financial Group, Inc.

70,000

691,600

Shinsei Bank Ltd.

117,000

534,769

Shares

Value

Shizuoka Bank Ltd.

40,000

$ 439,763

Sumitomo Mitsui Financial Group, Inc.

100

788,113

Tokyo Tomin Bank Ltd.

20,600

572,491

UMB Financial Corp.

7,600

320,188

4,653,704

Consumer Finance - 0.4%

Cash America International, Inc.

37,100

1,206,121

Diversified Financial Services - 1.5%

CME Group, Inc.

800

495,120

IntercontinentalExchange, Inc. (a)

9,400

1,315,624

MSCI, Inc. Class A

11,800

389,046

NYMEX Holdings, Inc.

18,400

2,116,000

Osaka Securities Exchange Co. Ltd.

64

346,092

4,661,882

Insurance - 0.9%

Assurant, Inc.

19,400

1,258,866

CNA Financial Corp.

11,480

390,205

CNinsure, Inc. ADR

44,000

472,560

LandAmerica Financial Group, Inc.

4,900

255,584

Philadelphia Consolidated Holdings Corp. (a)

9,600

343,680

Stewart Information Services Corp.

1,100

37,653

2,758,548

Real Estate Investment Trusts - 1.5%

Annaly Capital Management, Inc.

124,000

2,445,280

MFA Mortgage Investments, Inc.

189,300

1,930,860

4,376,140

Thrifts & Mortgage Finance - 0.8%

People's United Financial, Inc.

58,700

991,443

Washington Federal, Inc.

55,900

1,365,078

2,356,521

TOTAL FINANCIALS

27,937,504

HEALTH CARE - 16.6%

Biotechnology - 3.5%

Alexion Pharmaceuticals, Inc. (a)

18,500

1,208,420

Amylin Pharmaceuticals, Inc. (a)

74,607

2,212,098

Biogen Idec, Inc. (a)

26,400

1,609,080

Cephalon, Inc. (a)

11,300

741,619

Genentech, Inc. (a)

8,000

561,520

Gilead Sciences, Inc. (a)

41,362

1,889,830

Theravance, Inc. (a)(d)

127,038

2,506,460

10,729,027

Health Care Equipment & Supplies - 5.8%

Abiomed, Inc. (a)

25,000

377,500

Align Technology, Inc. (a)(d)

283,984

3,345,332

American Medical Systems Holdings, Inc. (a)

90,000

1,286,100

Conceptus, Inc. (a)

158,500

2,580,380

Hillenbrand Industries, Inc.

13,300

687,876

Hologic, Inc. (a)

14,000

901,040

Common Stocks - continued

Shares

Value

HEALTH CARE - continued

Health Care Equipment & Supplies - continued

Intuitive Surgical, Inc. (a)

11,000

$ 2,794,000

Inverness Medical Innovations, Inc. (a)

3,000

135,150

Mindray Medical International Ltd. sponsored ADR

10,000

341,000

NxStage Medical, Inc. (a)

14,100

171,174

Regeneration Technologies, Inc. (a)

215,000

1,720,000

TranS1, Inc. (d)

213,000

3,107,670

Varian Medical Systems, Inc. (a)

5,300

275,547

17,722,769

Health Care Providers & Services - 2.4%

athenahealth, Inc. (d)

49,400

1,551,160

Brookdale Senior Living, Inc. (d)

11,000

245,520

Coventry Health Care, Inc. (a)

5,500

311,190

Express Scripts, Inc. (a)

28,600

1,930,214

Health Net, Inc. (a)

10,300

478,847

Henry Schein, Inc. (a)

5,473

318,145

MWI Veterinary Supply, Inc. (a)

8,000

305,840

Pediatrix Medical Group, Inc. (a)

24,587

1,674,129

Psychiatric Solutions, Inc. (a)

5,900

178,003

VCA Antech, Inc. (a)

10,200

394,332

7,387,380

Health Care Technology - 1.0%

Cerner Corp. (a)

43,200

2,263,680

MedAssets, Inc.

7,900

159,185

TriZetto Group, Inc. (a)

38,900

759,328

3,182,193

Life Sciences Tools & Services - 2.0%

Affymetrix, Inc. (a)(d)

60,600

1,215,636

Applera Corp. - Applied Biosystems Group

30,000

945,900

Charles River Laboratories International, Inc. (a)

19,700

1,223,370

Covance, Inc. (a)

1,800

149,688

PAREXEL International Corp. (a)

10,000

544,100

Pharmaceutical Product Development, Inc.

30,200

1,309,472

Thermo Fisher Scientific, Inc. (a)

12,770

657,527

6,045,693

Pharmaceuticals - 1.9%

Allergan, Inc.

20,400

1,370,676

BioForm Medical, Inc. (d)

186,481

1,305,367

Perrigo Co.

15,700

484,188

Renovo Group PLC (a)

342,000

920,384

Teva Pharmaceutical Industries Ltd. sponsored ADR

22,300

1,026,692

XenoPort, Inc. (a)

8,600

527,696

5,635,003

TOTAL HEALTH CARE

50,702,065

Shares

Value

INDUSTRIALS - 17.1%

Aerospace & Defense - 1.0%

BE Aerospace, Inc. (a)

21,444

$ 827,953

Goodrich Corp.

19,900

1,244,745

Spirit AeroSystems Holdings, Inc.
Class A (a)

31,700

875,554

2,948,252

Air Freight & Logistics - 0.2%

United Parcel Service, Inc. Class B

6,400

468,224

Airlines - 2.3%

AMR Corp. (a)

103,000

1,435,820

Continental Airlines, Inc. Class B (a)

63,900

1,738,719

Northwest Airlines Corp. (a)

146,700

2,744,757

US Airways Group, Inc. (a)

89,200

1,231,852

7,151,148

Building Products - 0.3%

Ameron International Corp.

8,000

718,800

Universal Forest Products, Inc.

6,000

217,200

936,000

Commercial Services & Supplies - 5.1%

Advisory Board Co. (a)

2,600

165,672

American Reprographics Co. (a)

46,200

726,726

Copart, Inc. (a)

33,984

1,389,266

Corporate Executive Board Co.

9,400

540,970

Corrections Corp. of America (a)

94,182

2,499,590

Fuel Tech, Inc. (a)

35,600

677,824

GeoEye, Inc. (a)

15,000

524,400

IHS, Inc. Class A (a)

11,500

712,310

InnerWorkings, Inc. (a)

12,000

166,200

Manpower, Inc.

45,400

2,554,204

Republic Services, Inc.

56,600

1,698,000

The Geo Group, Inc. (a)

54,600

1,306,032

Waste Connections, Inc. (a)

45,600

1,329,696

Waste Management, Inc.

34,400

1,115,936

15,406,826

Construction & Engineering - 0.7%

Chicago Bridge & Iron Co. NV
(NY Shares)

4,900

218,001

Fluor Corp.

7,500

912,525

Grupo Acciona SA

2,000

509,808

Quanta Services, Inc. (a)

20,000

438,400

2,078,734

Electrical Equipment - 3.9%

Belden, Inc.

7,000

296,100

Energy Conversion Devices, Inc. (a)(d)

57,020

1,316,022

Evergreen Solar, Inc. (a)

154,800

1,887,012

First Solar, Inc. (a)

10,300

1,872,231

JA Solar Holdings Co. Ltd. ADR

16,000

813,280

Nexans SA

19,800

2,188,427

Q-Cells AG (a)

6,000

564,589

Sunpower Corp. Class A (a)(d)

35,400

2,445,786

Common Stocks - continued

Shares

Value

INDUSTRIALS - continued

Electrical Equipment - continued

Superior Essex, Inc. (a)

13,100

$ 315,055

Vestas Wind Systems AS (a)

2,800

271,999

11,970,501

Machinery - 1.3%

AGCO Corp. (a)

20,600

1,240,532

GEA Group AG (a)

32,000

988,991

Ingersoll-Rand Co. Ltd. Class A

18,000

711,360

Terex Corp. (a)

17,100

1,004,796

3,945,679

Marine - 0.4%

Ultrapetrol (Bahamas) Ltd. (a)

70,100

1,125,105

Road & Rail - 1.7%

J.B. Hunt Transport Services, Inc. (d)

56,100

1,744,710

Knight Transportation, Inc.

102,900

1,765,764

Landstar System, Inc.

36,500

1,826,095

5,336,569

Trading Companies & Distributors - 0.2%

Genesis Lease Ltd. ADR

31,600

635,160

TOTAL INDUSTRIALS

52,002,198

INFORMATION TECHNOLOGY - 19.8%

Communications Equipment - 2.0%

Ciena Corp. (a)

15,000

406,950

CommScope, Inc. (a)

11,700

518,895

Corning, Inc.

22,000

529,540

Juniper Networks, Inc. (a)

64,700

1,756,605

Polycom, Inc. (a)

111,000

2,802,750

6,014,740

Computers & Peripherals - 1.7%

Brocade Communications Systems, Inc. (a)

226,602

1,561,288

Diebold, Inc.

28,644

741,307

NCR Corp. (a)

61,600

1,323,168

Western Digital Corp. (a)

54,600

1,444,170

5,069,933

Electronic Equipment & Instruments - 1.6%

Agilent Technologies, Inc. (a)

8,700

295,017

Avnet, Inc. (a)

17,300

616,053

Bell Microproducts, Inc. (a)

29,818

156,843

Comverge, Inc.

6,100

113,704

Flextronics International Ltd. (a)

51,100

597,870

Ingram Micro, Inc. Class A (a)

30,700

545,846

Itron, Inc. (a)

24,000

1,977,600

Jabil Circuit, Inc.

47,300

626,725

4,929,658

Internet Software & Services - 2.8%

Akamai Technologies, Inc. (a)

16,800

507,360

DealerTrack Holdings, Inc. (a)

23,600

636,256

Shares

Value

Dice Holdings, Inc.

258,600

$ 1,649,868

DivX, Inc. (a)

50,000

712,500

Internet Brands, Inc. Class A

30,979

239,158

Omniture, Inc. (a)

85,700

2,118,504

Online Resources Corp. (a)

89,800

916,858

The Knot, Inc. (a)

49,700

727,608

ValueClick, Inc. (a)

44,100

962,703

8,470,815

IT Services - 1.1%

Cognizant Technology Solutions Corp. Class A (a)

72,700

2,028,330

CyberSource Corp. (a)

38,000

636,500

Mastercard, Inc. Class A

3,800

786,600

3,451,430

Semiconductors & Semiconductor Equipment - 7.4%

Altera Corp.

93,000

1,570,770

Applied Materials, Inc.

125,400

2,247,168

ARM Holdings PLC sponsored ADR

23,600

167,088

ASML Holding NV (NY Shares) (a)

20,000

531,800

Broadcom Corp. Class A (a)

66,500

1,468,320

Cypress Semiconductor Corp. (a)

62,200

1,321,750

FormFactor, Inc. (a)

21,063

510,146

Himax Technologies, Inc. sponsored ADR

285,100

1,405,543

Hittite Microwave Corp. (a)

12,400

493,768

Intersil Corp. Class A

31,400

723,142

Lam Research Corp. (a)

46,500

1,785,135

LDK Solar Co. Ltd. Sponsored ADR (d)

19,800

697,950

Marvell Technology Group Ltd. (a)

122,180

1,450,277

MEMC Electronic Materials, Inc. (a)

18,500

1,322,010

Microchip Technology, Inc.

35,500

1,132,805

Miraial Co. Ltd.

15,900

386,547

National Semiconductor Corp.

53,000

976,790

Skyworks Solutions, Inc. (a)(d)

124,700

1,003,835

Teradyne, Inc. (a)

5,400

59,238

Varian Semiconductor Equipment Associates, Inc. (a)

63,800

2,054,998

Xilinx, Inc.

61,200

1,338,444

22,647,524

Software - 3.2%

Activision, Inc. (a)

25,100

649,337

Citrix Systems, Inc. (a)

12,000

415,440

CompuGROUP Holding AG (a)

33,000

622,352

Concur Technologies, Inc. (a)

53,300

1,868,698

FactSet Research Systems, Inc.

6,000

335,580

Jack Henry & Associates, Inc.

53,300

1,310,114

Microsoft Corp.

20,800

678,080

Nintendo Co. Ltd.

400

197,600

Nuance Communications, Inc. (a)

87,000

1,382,430

Shanda Interactive Entertainment Ltd. sponsored ADR (a)

17,900

492,071

SPSS, Inc. (a)

9,000

297,450

Common Stocks - continued

Shares

Value

INFORMATION TECHNOLOGY - continued

Software - continued

Take-Two Interactive Software, Inc. (a)(d)

61,650

$ 1,013,526

THQ, Inc. (a)

33,800

608,738

9,871,416

TOTAL INFORMATION TECHNOLOGY

60,455,516

MATERIALS - 3.1%

Chemicals - 0.2%

FMC Corp.

11,300

600,708

Construction Materials - 0.5%

Martin Marietta Materials, Inc. (d)

13,800

1,693,536

Containers & Packaging - 0.4%

Smurfit-Stone Container Corp. (a)

39,600

375,804

Temple-Inland, Inc.

39,600

742,500

1,118,304

Metals & Mining - 1.9%

Allegheny Technologies, Inc.

19,542

1,375,757

Barrick Gold Corp.

6,900

356,224

Century Aluminum Co. (a)

14,000

727,860

Newmont Mining Corp.

10,000

543,400

Nucor Corp.

22,200

1,283,160

Randgold Resources Ltd. sponsored ADR

5,800

276,718

RTI International Metals, Inc. (a)

23,140

1,278,485

5,841,604

Paper & Forest Products - 0.1%

Glatfelter

12,400

179,304

TOTAL MATERIALS

9,433,456

TELECOMMUNICATION SERVICES - 1.3%

Wireless Telecommunication Services - 1.3%

American Tower Corp. Class A (a)

52,705

1,978,019

NII Holdings, Inc. (a)

48,691

2,077,158

4,055,177

UTILITIES - 4.0%

Electric Utilities - 2.1%

Allegheny Energy, Inc.

17,600

964,304

Enernoc, Inc.

14,300

504,075

Shares

Value

Entergy Corp.

4,500

$ 486,810

PPL Corp.

89,800

4,393,015

6,348,204

Independent Power Producers & Energy Traders - 1.9%

AES Corp. (a)

79,400

1,514,952

Clipper Windpower PLC (a)

2,400

28,693

Constellation Energy Group, Inc.

42,800

4,021,488

Ormat Technologies, Inc.

3,800

165,186

5,730,319

TOTAL UTILITIES

12,078,523

TOTAL COMMON STOCKS

(Cost $305,398,397)

305,574,852

Money Market Funds - 4.3%

Fidelity Cash Central Fund, 3.79% (b)

2,288,816

2,288,816

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

10,764,675

10,764,675

TOTAL MONEY MARKET FUNDS

(Cost $13,053,491)

13,053,491

TOTAL INVESTMENT PORTFOLIO - 104.5%

(Cost $318,451,888)

318,628,343

NET OTHER ASSETS - (4.5)%

(13,583,264)

NET ASSETS - 100%

$ 305,045,079

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 442,525

Fidelity Securities Lending Cash Central Fund

313,679

Total

$ 756,204

Other Information

Distribution of investments by country of issue, as a percentage of total net assets, is as follows:

United States of America

88.3%

Japan

1.8%

Bermuda

1.7%

France

1.5%

Cayman Islands

1.3%

Others (individually less than 1%)

5.4%

100.0%

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $18,706,577 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Growth Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $10,830,743) - See accompanying schedule:

Unaffiliated issuers (cost $305,398,397)

$ 305,574,852

Fidelity Central Funds (cost $13,053,491)

13,053,491

Total Investments (cost $318,451,888)

$ 318,628,343

Cash

944,763

Receivable for investments sold

60,992,695

Receivable for fund shares sold

386,427

Dividends receivable

50,720

Distributions receivable from Fidelity Central Funds

53,069

Prepaid expenses

1,182

Other receivables

2,063

Total assets

381,059,262

Liabilities

Payable for investments purchased

$ 23,187,980

Payable for fund shares redeemed

41,783,947

Accrued management fee

108,332

Distribution fees payable

1,571

Other affiliated payables

110,841

Other payables and accrued expenses

56,837

Collateral on securities loaned, at value

10,764,675

Total liabilities

76,014,183

Net Assets

$ 305,045,079

Net Assets consist of:

Paid in capital

$ 319,016,951

Accumulated undistributed net realized gain (loss) on investments and foreign currency transactions

(14,150,236)

Net unrealized appreciation (depreciation) on investments and assets and liabilities in foreign currencies

178,364

Net Assets

$ 305,045,079

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($1,936,249 ÷ 158,895 shares)

$ 12.19

Maximum offering price per share (100/94.25 of $12.19)

$ 12.93

Class T:
Net Asset Value
and redemption price per share ($591,188 ÷ 48,688 shares)

$ 12.14

Maximum offering price per share (100/96.50 of $12.14)

$ 12.58

Class B:
Net Asset Value
and offering price per share ($413,537 ÷ 34,216 shares)A

$ 12.09

Class C:
Net Asset Value
and offering price per share ($697,298 ÷ 57,672 shares)A

$ 12.09

Mid Cap Growth:
Net Asset Value
, offering price and redemption price per share ($301,224,741 ÷ 24,671,114 shares)

$ 12.21

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($182,066 ÷ 14,896 shares)

$ 12.22

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Growth Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 2,094,532

Interest

386

Income from Fidelity Central Funds (including $313,679 from security lending)

756,204

Total income

2,851,122

Expenses

Management fee
Basic fee

$ 2,332,314

Performance adjustment

(430,070)

Transfer agent fees

1,202,314

Distribution fees

10,363

Accounting and security lending fees

170,995

Custodian fees and expenses

18,214

Independent trustees' compensation

1,531

Registration fees

100,229

Audit

56,662

Legal

2,290

Miscellaneous

25,324

Total expenses before reductions

3,490,166

Expense reductions

(113,488)

3,376,678

Net investment income (loss)

(525,556)

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

13,581,162

Foreign currency transactions

16,150

Total net realized gain (loss)

13,597,312

Change in net unrealized appreciation (depreciation) on:

Investment securities

(50,635,596)

Assets and liabilities in foreign currencies

1,909

Total change in net unrealized appreciation (depreciation)

(50,633,687)

Net gain (loss)

(37,036,375)

Net increase (decrease) in net assets resulting from operations

$ (37,561,931)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ (525,556)

$ (1,458,396)

Net realized gain (loss)

13,597,312

(4,757,238)

Change in net unrealized appreciation (depreciation)

(50,633,687)

5,161,873

Net increase (decrease) in net assets resulting from operations

(37,561,931)

(1,053,761)

Distributions to shareholders from net realized gain

(21,909,497)

(5,089,264)

Share transactions - net increase (decrease)

(76,821,412)

97,426,141

Redemption fees

26,012

46,437

Total increase (decrease) in net assets

(136,266,828)

91,329,553

Net Assets

Beginning of period

441,311,907

349,982,354

End of period

$ 305,045,079

$ 441,311,907

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.05)

Net realized and unrealized gain (loss)

(1.30)

Total from investment operations

(1.35)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.19

Total Return B,C,D

(9.95)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.10% A

Expenses net of fee waivers, if any

1.10% A

Expenses net of all reductions

1.10% A

Net investment income (loss)

(.41)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,936

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.09)

Net realized and unrealized gain (loss)

(1.31)

Total from investment operations

(1.40)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.14

Total Return B,C,D

(10.30)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.36% A

Expenses net of fee waivers, if any

1.36% A

Expenses net of all reductions

1.36% A

Net investment income (loss)

(.68)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 591

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.16)

Net realized and unrealized gain (loss)

(1.29)

Total from investment operations

(1.45)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.09

Total Return B,C,D

(10.65)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.85% A

Expenses net of fee waivers, if any

1.85% A

Expenses net of all reductions

1.85% A

Net investment income (loss)

(1.16)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 414

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.15)

Net realized and unrealized gain (loss)

(1.30)

Total from investment operations

(1.45)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.09

Total Return B,C,D

(10.65)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.85% A

Expenses net of fee waivers, if any

1.85% A

Expenses net of all reductions

1.85% A

Net investment income (loss)

(1.16)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 697

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Mid Cap Growth

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 14.31

$ 14.38

$ 11.58

$ 10.63

$ 7.32

Income from Investment Operations

Net investment income (loss) B

(.02)

(.04)

.01 E

(.03) F

(.07)

Net realized and unrealized gain (loss)

(1.29)

.15

3.09

1.15

3.38

Total from investment operations

(1.31)

.11

3.10

1.12

3.31

Distributions from net realized gain

(.79)

(.18)

(.30)

(.17)

-

Redemption fees added to paid in capital B,H

-

-

-

-

-

Net asset value, end of period

$ 12.21

$ 14.31

$ 14.38

$ 11.58

$ 10.63

Total Return A

(9.68)%

.80%

27.15%

10.55%

45.22%

Ratios to Average Net Assets C,G

Expenses before reductions

.83%

1.02%

1.04%

1.02%

1.25%

Expenses net of fee waivers, if any

.81%

1.00%

1.00%

1.02%

1.20%

Expenses net of all reductions

.81%

.99%

.95%

.99%

1.16%

Net investment income (loss)

(.12)%

(.33)%

.07% E

(.31)% F

(.77)%

Supplemental Data

Net assets, end of period (000 omitted)

$ 301,225

$ 441,312

$ 349,982

$ 77,658

$ 60,660

Portfolio turnover rate D

245%

178%

173%

220%

94%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.03 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.18)%.

F Investment income per share reflects a special dividend which amounted to $.01 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.44)%.

G Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

H Amount represents less than $.01 per share.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net realized and unrealized gain (loss) D

(1.32)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.22

Total Return B,C

(9.74)%

Ratios to Average Net Assets E,H

Expenses before reductions

.72% A

Expenses net of fee waivers, if any

.72% A

Expenses net of all reductions

.72% A

Net investment income (loss)

(.03)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 182

Portfolio turnover rate F

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Notes to Financial Statements

For the period ended January 31, 2008

1. Organization.

Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund(the Funds) are funds of Fidelity Devonshire Trust(the trust) and are authorized to issue an unlimited number of shares. The trust is registered under the Investment Company Act of 1940, as amended(the 1940 Act), as an open-end management company organized as a Massachusetts business trust. The Funds offer Class A, Class T, Class B, Class C, Institutional Class and Large Cap Value, Mid Cap Value, Large Cap Growth and Mid Cap Growth shares, respectively, each of which has equal rights as to assets and voting privileges. Each class has exclusive voting rights with respect to matters that affect that class. The Funds commenced sale of Class A, Class T, Class B, Class C and Institutional Class shares and the existing class of each Fund was designated Large Cap Value, Mid Cap Value, Large Cap Growth and Mid Cap Growth, respectively, on February 13, 2007. Class B shares will automatically convert to Class A shares after a holding period of seven years from the initial date of purchase. Investment income, realized and unrealized capital gains and losses, the common expenses of the Fund, and certain fund-level expense reductions, if any, are allocated on a pro rata basis to each class based on the relative net assets of each class to the total net assets of the Fund. Each class differs with respect to transfer agent and distribution and service plan fees incurred. Certain expense reductions also differ by class. Certain Funds investments in emerging markets can be subject to social, economic, regulatory, and political uncertainties and can be extremely volatile.

2. Investments in Fidelity Central Funds.

The Funds may invest in Fidelity Central Funds, which are open-end investment companies available only to other investment companies and accounts managed by Fidelity Management & Research Company(FMR) and its affiliates. The Funds' Schedules of Investments list each of the Fidelity Central Funds as an investment of each Fund, but does not include the underlying holdings of each Fidelity Central Fund. As an Investing Fund, the Funds indirectly bear their proportionate share of the expenses of the underlying Fidelity Central Funds.

The Money Market Central Funds seek preservation of capital and current income and are managed by Fidelity Investments Money Management, Inc.(FIMM), an affiliate of FMR.

A complete list of holdings for each Fidelity Central Fund is available upon request or at the SEC's web site at www.sec.gov. In addition, the financial statements of the Fidelity Central Funds, which are not covered by the Funds' Report of Independent Registered Public Accounting Firm, are available on the SEC's web site or upon request.

3. Significant Accounting Policies.

The financial statements have been prepared in conformity with accounting principles generally accepted in the United States of America, which require management to make certain estimates and assumptions at the date of the financial statements. Actual results could differ from those estimates. The following summarizes the significant accounting policies of the Funds:

Security Valuation. Investments are valued and net asset value(NAV) per share is calculated(NAV calculation) as of the close of business of the New York Stock Exchange(NYSE), normally 4:00 p.m. Eastern time. Wherever possible, each Fund uses independent pricing services approved by the Board of Trustees to value their investments.

Equity securities, including restricted securities, for which market quotations are readily available, are valued at the last reported sale price or official closing price as reported by an independent pricing service on the primary market or exchange on which they are traded. In the event there were no sales during the day or closing prices are not available, securities are valued at the last quoted bid price. Investments in open-end mutual funds, including the Fidelity Central Funds, are valued at their closing net asset value each business day. Short-term securities with remaining maturities of sixty days or less for which quotations are not readily available are valued at amortized cost, which approximates value.

When current market prices or quotations are not readily available or do not accurately reflect fair value, valuations may be determined in accordance with procedures adopted by the Board of Trustees. For example, when developments occur between the close of a market and the close of the NYSE that may materially affect the value of some or all of the securities, or when trading in a security is halted, those securities may be fair valued. Factors used in the determination of fair value may include monitoring news to identify significant market or security specific events such as changes in the value of U.S. securities markets, reviewing developments in foreign markets and evaluating the performance of ADRs, futures contracts and exchange-traded funds. Because each Fund's utilization of fair value pricing depends on market activity, the frequency with which fair value pricing is used cannot be predicted and may be utilized to a significant extent. The value of securities used for NAV calculation under fair value pricing may differ from published prices for the same securities.

Foreign Currency. Certain Funds may use foreign currency contracts to facilitate transactions in foreign-denominated securities. Losses from these transactions may arise from changes in the value of the foreign currency or if the counterparties do not perform under the contracts' terms.

Annual Report

Notes to Financial Statements - continued

3. Significant Accounting Policies - continued

Foreign Currency - continued

Foreign-denominated assets, including investment securities, and liabilities are translated into U.S. dollars at the exchange rate at period end. Purchases and sales of investment securities, income and dividends received and expenses denominated in foreign currencies are translated into U.S. dollars at the exchange rate in effect on the transaction date.

The effects of exchange rate fluctuations on investments are included with the net realized and unrealized gain(loss) on investment securities. Other foreign currency transactions resulting in realized and unrealized gain(loss) are disclosed separately.

Investment Transactions and Income. For financial reporting purposes, the Funds' investment holdings and NAV include trades executed through the end of the last business day of the period. The NAV for processing shareholder transactions includes trades executed through the end of the prior business day. Gains and losses on securities sold are determined on the basis of identified cost and may include proceeds received from litigation. Dividend income is recorded on the ex-dividend date, except for certain dividends from foreign securities where the ex-dividend date may have passed, which are recorded as soon as the Funds are informed of the ex-dividend date. Non-cash dividends included in dividend income, if any, are recorded at the fair market value of the securities received. Distributions received on securities that represent a return of capital or capital gain are recorded as a reduction of cost of investments and/or as a realized gain. The Funds estimate the components of distributions received that may be considered return of capital distributions or capital gain distributions. Interest income and distributions from the Fidelity Central Funds are accrued as earned. Interest income includes coupon interest and amortization of premium and accretion of discount on debt securities.

Expenses. Most expenses of the trust can be directly attributed to a fund. Expenses which cannot be directly attributed are apportioned among each Fund in the trust. Expense estimates are accrued in the period to which they relate and adjustments are made when actual amounts are known.

Income Tax Information and Distributions to Shareholders. Each year, each Fund intends to qualify as a regulated investment company by distributing substantially all of its taxable income and realized gains under Subchapter M of the Internal Revenue Code and filing its U.S. federal tax return. As a result, no provision for income taxes is required. Each Fund adopted the provisions of FASB Interpretation No. 48, Accounting for Uncertainties in Income Taxes(FIN 48), on June 29, 2007. FIN 48 sets forth a minimum threshold for financial statement recognition of the benefit of a tax position taken or expected to be taken in a tax return. The implementation of FIN 48 did not result in any unrecognized tax benefits in the accompanying financial statements. Each of the Funds' federal tax returns for the prior three fiscal years remains subject to examination by the Internal Revenue Service. Foreign taxes are provided for based on each Fund's understanding of the tax rules and rates that exist in the foreign markets in which it invests.

Distributions are recorded on the ex-dividend date. Income dividends and capital gain distributions are declared separately for each class. Income and capital gain distributions are determined in accordance with income tax regulations, which may differ from generally accepted accounting principles. In addition, certain Funds will claim a portion of the payment made to redeeming shareholders as a distribution for income tax purposes.

Capital accounts within the financial statements are adjusted for permanent book-tax differences. These adjustments have no impact on net assets or the results of operations. Temporary book-tax differences will reverse in a subsequent period.

Book-tax differences are primarily due to foreign currency transactions, passive foreign investment companies (PFIC), partnerships, capital loss carryforwards and losses deferred due to wash sales and excise tax regulations.

The tax-basis components of distributable earnings and the federal tax cost as of period end were as follows for each Fund:

Cost for Federal
Income Tax
Purposes

Unrealized
Appreciation

Unrealized
Depreciation

Net Unrealized
Appreciation/
(Depreciation)

Fidelity Large Cap Value Fund

$ 1,571,171,429

$ 98,475,075

$ (92,166,547)

$ 6,308,528

Fidelity Mid Cap Value Fund

844,773,929

42,574,804

(57,370,158)

(14,795,354)

Fidelity Large Cap Growth Fund

166,610,940

4,786,892

(10,368,163)

(5,581,271)

Fidelity Mid Cap Growth Fund

319,883,846

21,520,033

(22,775,536)

(1,255,503)

Annual Report

3. Significant Accounting Policies - continued

Income Tax Information and Distributions to Shareholders - continued

The tax character of distributions paid was as follows:

January 31, 2008

Ordinary
Income

Long-term
Capital Gains

Total

Fidelity Large Cap Value Fund

$ 37,304,898

$ 67,715,379

$ 105,020,277

Fidelity Mid Cap Value Fund

10,958,373

29,219,096

40,177,469

Fidelity Large Cap Growth Fund

6,193,151

12,379,253

18,572,404

Fidelity Mid Cap Growth Fund

-

21,909,497

21,909,497

January 31, 2007

Ordinary Income

Long-term
Capital Gains

Total

Fidelity Large Cap Value Fund

$ 17,006,124

$ 7,416,999

$ 24,423,123

Fidelity Mid Cap Value Fund

7,722,049

9,225,041

16,947,090

Fidelity Large Cap Growth Fund

1,920,898

1,860,806

3,781,704

Fidelity Mid Cap Growth Fund

2,544,632

2,544,632

5,089,264

Short-Term Trading (Redemption) Fees. Shares held in the Fidelity Mid Cap Value Fund and Fidelity Mid Cap Growth Fund less than 30 days are subject to a redemption fee equal to 0.75% of the proceeds of the redeemed shares. All redemption fees, including any estimated redemption fees paid by FMR, are retained by the Funds and accounted for as an addition to paid in capital.

New Accounting Pronouncement. In September 2006, Statement of Financial Accounting Standards No. 157, Fair Value Measurements(SFAS 157), was issued and is effective for fiscal years beginning after November 15, 2007. SFAS 157 defines fair value, establishes a framework for measuring fair value and expands disclosures about fair value measurements. Management is currently evaluating the impact the adoption of SFAS 157 will have on the Funds' financial statement disclosures.

4. Operating Policies.

Repurchase Agreements. FMR has received an Exemptive Order from the Securities and Exchange Commission(the SEC) which permits certain Funds and other affiliated entities of FMR to transfer uninvested cash balances into joint trading accounts which are then invested in repurchase agreements. Certain Funds may also invest directly with institutions in repurchase agreements. Repurchase agreements are collateralized by government or non-government securities. Upon settlement date, collateral is held in segregated accounts with custodian banks and may be obtained in the event of a default of the counterparty. Each applicable Fund monitors, on a daily basis, the value of the collateral to ensure it is at least equal to the principal amount of the repurchase agreement(including accrued interest). In the event of a default by the counterparty, realization of the collateral proceeds could be delayed, during which time the value of the collateral may decline.

5. Purchases and Sales of Investments.

Purchases and sales of securities, other than short-term securities, are noted in the table below.

Purchases($)

Sales($)

Fidelity Large Cap Value Fund

3,485,833,978

3,246,788,832

Fidelity Mid Cap Value Fund

2,520,812,788

2,356,094,981

Fidelity Large Cap Growth Fund

745,952,270

764,990,241

Fidelity Mid Cap Growth Fund

995,464,036

1,090,139,044

6. Fees and Other Transactions with Affiliates.

Management Fee. FMR and its affiliates provide the Funds with investment management related services for which the Funds pay a monthly management fee. The management fee is the sum of an individual fund fee rate and a group fee rate. The individual fund fee rate is applied to each Fund's average net assets. The group fee rate is based upon the average net assets of all the mutual funds advised by FMR. The group fee rate decreases as assets under management increase and increases as assets under management decrease. In addition, the management fee for the funds is subject to a performance adjustment(up to a maximum ± .20% of each applicable Fund's average net assets over a 36 month performance period). The upward or downward adjustment to the management fee is based on each applicable Fund's relative investment performance of the asset-weighted return of all

Annual Report

Notes to Financial Statements - continued

6. Fees and Other Transactions with Affiliates - continued

Management Fee - continued

classes as compared to an appropriate benchmark index. For the period, each Fund's annual management fee rate expressed as a percentage of each Fund's average net assets, including the performance adjustment, if applicable was as follows:

Individual Rate

Group Rate

Total

Fidelity Large Cap Value Fund

.30%

.26%

.58%

Fidelity Mid Cap Value Fund

.30%

.26%

.52%

Fidelity Large Cap Growth Fund

.30%

.26%

.58%

Fidelity Mid Cap Growth Fund

.30%

.26%

.46%

Distribution and Service Plan. In accordance with Rule 12b-1 of the 1940 Act, the Fund has adopted separate Distribution and Service Plans for each class of shares. Certain classes pay Fidelity Distributors Corporation(FDC), an affiliate of FMR, separate Distribution and Service Fees, each of which is based on an annual percentage of each class' average net assets. In addition, FDC may pay financial intermediaries for selling shares of the Fund and providing shareholder support services. For the period, the Distribution and Service Fee rates and the total amounts paid to and retained by FDC were as follows:

Fidelity Large Cap Value Fund

Distribution
Fee

Service
Fee

Paid to
FDC

Retained
by FDC

Class A

0%

.25%

$ 14,716

$ 3,419

Class T

.25%

.25%

29,520

209

Class B

.75%

.25%

20,952

15,912

Class C

.75%

.25%

8,540

4,769

$ 73,728

$ 24,309

Fidelity Mid Cap Value Fund

Class A

0%

.25%

$ 9,176

$ 3,141

Class T

.25%

.25%

10,994

355

Class B

.75%

.25%

17,435

13,285

Class C

.75%

.25%

16,086

12,118

$ 53,691

$ 28,899

Fidelity Large Cap Growth Fund

Class A

0%

.25%

$ 1,502

$ 246

Class T

.25%

.25%

2,248

460

Class B

.75%

.25%

2,797

2,331

Class C

.75%

.25%

4,188

3,019

$ 10,735

$ 6,056

Fidelity Mid Cap Growth Fund

Class A

0%

.25%

$ 1,756

$ 245

Class T

.25%

.25%

1,796

463

Class B

.75%

.25%

2,959

2,452

Class C

.75%

.25%

3,852

3,025

$ 10,363

$ 6,185

Sales Load. FDC receives a front-end sales charge of up to 5.75% for selling Class A shares, and 3.50% for selling Class T shares, some of which is paid to financial intermediaries for selling shares of the Fund. FDC receives the proceeds of contingent deferred sales charges levied on Class A, Class T, Class B, and Class C redemptions. These charges depend on the holding period. The deferred sales charges range from 5% to 1% for Class B, 1% for Class C, 1.00% to .50% for certain purchases of Class A shares and .25% for certain purchases of Class T shares.

For the period, sales charge amounts retained by FDC were as follows:

Fidelity Large Cap Value Fund

Retained
by FDC

Class A

$ 18,265

Class T

4,391

Class B *

1,096

Class C *

11

$ 22,763

Annual Report

6. Fees and Other Transactions with Affiliates - continued

Sales Load - continued

For the period, sales charge amounts retained by FDC were as follows: - continued

Fidelity Mid Cap Value Fund

Retained
by FDC

Class A

$ 22,621

Class T

5,951

Class B*

2,448

Class C*

137

$ 31,157

Fidelity Large Cap Growth Fund

Class A

$ 4,515

Class T

1,123

Class B*

509

Class C*

62

$ 6,209

Fidelity Mid Cap Growth Fund

Class A

$ 6,603

Class T

1,049

Class B*

962

Class C*

165

$ 8,779

* When Class B and Class C shares are initially sold, FDC pays commissions from its own resources to financial intermediaries through which the sales are made.

Transfer Agent Fees. Fidelity Investments Institutional Operations Company, Inc.(FIIOC), an affiliate of FMR, is the transfer, dividend disbursing and shareholder servicing agent for each class of the Funds. FIIOC receives account fees and asset-based fees that vary according to the account size and type of account of the shareholders of the respective classes of each Fund. FIIOC pays for typesetting, printing and mailing of shareholder reports, except proxy statements. Prior to January 1, 2008, Fidelity Service Company, Inc. (FSC), also an affiliate of FMR, was the Fund's transfer agent. For the period, the total transfer agent fees paid by each class to FIIOC or FSC, were as follows:

Fidelity Large Cap Value Fund

Amount

% of
Average
Net Assets
*

Class A

$ 20,036

.34

Class T

20,697

.35

Class B

7,804

.37

Class C

2,694

.31

Large Cap Value

3,716,520

.23

Institutional Class

944

.23

$ 3,768,695

Fidelity Mid Cap Value Fund

Class A

$ 10,910

.30

Class T

6,730

.31

Class B

5,278

.30

Class C

5,017

.31

Mid Cap Value

2,206,272

.24

Institutional Class

1,803

.30

$ 2,236,010

Fidelity Large Cap Growth Fund

Class A

$ 1,787

.30

Class T

1,353

.31

Class B

833

.30

Class C

1,224

.29

Large Cap Growth

529,155

.31

Institutional Class

416

.20

$ 534,768

Annual Report

Notes to Financial Statements - continued

6. Fees and Other Transactions with Affiliates - continued

Transfer Agent Fees - continued

Fidelity Mid Cap Growth Fund

Amount

% of
Average
Net Assets
*

Class A

$ 2,094

.30

Class T

1,136

.32

Class B

899

.30

Class C

1,151

.30

Mid Cap Growth

1,196,809

.29

Institutional Class

225

.17

$ 1,202,314

* Annualized

Accounting and Security Lending Fees. Fidelity Service Company, Inc.(FSC), an affiliate of FMR, maintains each Fund's accounting records. The accounting fee is based on the level of average net assets for the month. Under a separate contract, FSC administers the security lending program. The security lending fee is based on the number and duration of lending transactions.

Brokerage Commissions. Certain Funds placed a portion of their portfolio transactions with brokerage firms which are affiliates of the investment adviser. The commissions paid to these affiliated firms were as follows:

Amount

Fidelity Large Cap Value Fund

$ 18,780

Fidelity Mid Cap Value Fund

12,752

Fidelity Large Cap Growth Fund

13,318

Fidelity Mid Cap Growth Fund

10,529

Interfund Lending Program. Pursuant to an Exemptive Order issued by the SEC, the Funds, along with other registered investment companies having management contracts with FMR, may participate in an interfund lending program. This program provides an alternative credit facility allowing the funds to borrow from, or lend money to, other participating affiliated funds. At period end, there were no interfund loans outstanding. Each applicable fund's activity in this program during the period for which loans were outstanding was as follows:

Borrower or
Lender

Average Daily Loan Balance

Weighted
Average Interest Rate

Interest
Expense

Fidelity Large Cap Value Fund

Borrower

$ 6,183,857

4.35%

$ 5,226

Fidelity Mid Cap Value Fund

Borrower

$ 5,637,000

5.27%

$ 4,946

7. Committed Line of Credit.

Certain Funds participate with other funds managed by FMR in a $4.2 billion credit facility(the "line of credit") to be utilized for temporary or emergency purposes to fund shareholder redemptions or for other short-term liquidity purposes. The participating funds have agreed to pay commitment fees on their pro rata portion of the line of credit, which is reflected in Miscellaneous Expense on the Statement of Operations, and is as follows:

Fidelity Large Cap Value Fund

$ 3,040

Fidelity Mid Cap Value Fund

1,696

Fidelity Large Cap Growth Fund

343

Fidelity Mid Cap Growth Fund

824

During the period, there were no borrowings on this line of credit.

8. Security Lending.

Certain Funds lend portfolio securities from time to time in order to earn additional income. On the settlement date of the loan, each applicable Fund receives collateral(in the form of U.S. Treasury obligations, letters of credit and/or cash) against the loaned securities and maintains collateral in an amount not less than 100% of the market value of the loaned securities during the period of the loan. The market value of the loaned securities is determined at the close of business of the Funds and any additional required collateral is delivered to the Funds on the next business day. If the borrower defaults on its obligation to return the securities loaned because of insolvency or other reasons, a fund could experience delays and costs in recovering the securities loaned or in gaining access to the collateral. Any cash collateral received is invested in the Fidelity Securities Lending Cash

Annual Report

8. Security Lending - continued

Central Fund. The value of loaned securities and cash collateral at period end are disclosed on each applicable Fund's Statement of Assets and Liabilities. Security lending income represents the income earned on investing cash collateral, less fees and expenses associated with the loan, plus any premium payments that may be received on the loan of certain types of securities. Security lending income is presented on each applicable Fund's Statement of Operations as a component of income from Fidelity Central Funds.

9. Expense Reductions.

FMR voluntarily agreed to reimburse funds to the extent annual operating expenses exceeded certain levels of average net assets as noted in the table below. Some expenses, for example interest expense, including commitment fees, are excluded from this reimbursement.

The following classes of each applicable Fund were in reimbursement during the period:

Expense
Limitations

Reimbursement
from adviser

Fidelity Large Cap Growth Fund

Large Cap Growth

1.00%

$ 58,552

In addition, FMR voluntarily agreed to reimburse a portion of certain fund's existing class' operating expenses. During the period, this reimbursement reduced certain fund's existing class' expenses by the following amounts:

Reimbursement
from adviser

Fidelity Large Cap Value Fund

Large Cap Value

$ 85,974

Fidelity Mid Cap Value Fund

Mid Cap Value

$ 85,078

Fidelity Large Cap Growth Fund

Large Cap Growth

$ 25,900

Fidelity Mid Cap Growth Fund

Mid Cap Growth

$ 84,552

In addition, through arrangements with each applicable Fund's custodian and each class' transfer agent, credits realized as a result of uninvested cash balances were used to reduce the Fund's expenses. All of the applicable expense reductions are noted in the table below.

Brokerage Service Arrangements

Transfer
Agent
expense
reduction

Fidelity Large Cap Value Fund

$ -

$ -

Large Cap Value

-

50,587

Institutional Class

-

50

Fidelity Mid Cap Value Fund

-

-

Class A

-

31

Class B

-

9

Mid Cap Value

-

19,810

Institutional Class

-

70

Fidelity Large Cap Growth Fund

-

-

Large Cap Growth

-

13,456

Fidelity Mid Cap Growth Fund

1,543

-

Mid Cap Growth

-

27,293

10. Other.

The Funds' organizational documents provide former and current trustees and officers with a limited indemnification against liabilities arising in connection with the performance of their duties to the Funds. In the normal course of business, the Funds may also enter into contracts that provide general indemnifications. The Funds' maximum exposure under these arrangements is unknown as this would be dependent on future claims that may be made against the Funds. The risk of material loss from such claims is considered remote.

Annual Report

Notes to Financial Statements - continued

10. Other - continued

In December 2006, the Independent Trustees, with the assistance of independent counsel, completed an investigation regarding gifts, gratuities and business entertainment provided by certain brokers to certain individuals who were employed on FMR's domestic equity trading desk during the period 2002 to 2004. The Independent Trustees and FMR agreed that, despite the absence of proof that the Fidelity mutual funds experienced diminished execution quality as a result of the improper receipt of gifts and business entertainment, the conduct at issue was serious and was worthy of redress. Accordingly, the Independent Trustees requested, and FMR agreed to make a payment of $42 million plus accrued interest, which equaled approximately $7.3 million, to certain Fidelity mutual funds. FMR also reimbursed the related legal expenses, which are recorded in the accompanying Statement of Operations as an expense reduction.

Subsequent to period end, the Trustees approved a method for allocating this payment among the funds and, in total, FMR paid each fund the following amounts:

Fidelity Large Cap Value Fund

$ 12,127

Fidelity Mid Cap Value Fund

$ 23,107

Fidelity Large Cap Growth Fund

$ 14,058

Fidelity Mid Cap Growth Fund

$ 16,394

In a related administrative order dated March 5, 2008, the U.S. Securities and Exchange Commission ("SEC") announced a settlement with FMR and FMR Co., Inc. (an affiliate of FMR) involving the SEC's regulatory rules for investment advisers and the improper receipt of gifts, gratuities and business entertainment. Without admitting or denying the SEC's findings, FMR agreed to pay an $8 million civil penalty to the United States Treasury.

11. Distributions to Shareholders.

Distributions to shareholders of each class were as follows:

Years ended January 31,

2008 A

2007

Fidelity Large Cap Value Fund

From net investment income

Class A

$ 75,483

$ -

Class T

44,031

-

Class B

824

-

Class C

4,115

-

Large Cap Value

14,152,017

10,401,399

Institutional Class

10,129

-

Total

$ 14,286,599

$ 10,401,399

From net realized gain

Class A

$ 372,920

$ -

Class T

317,190

-

Class B

97,442

-

Class C

51,607

-

Large Cap Value

89,854,195

14,021,724

Institutional Class

40,324

-

Total

$ 90,733,678

$ 14,021,724

Fidelity Mid Cap Value Fund

From net investment income

Class A

$ 18,691

$ -

Class T

5,836

-

Class B

259

-

Class C

66

-

Mid Cap Value

3,005,000

2,949,768

Institutional Class

7,163

-

Total

$ 3,037,015

$ 2,949,768

From net realized gain

Class A

$ 176,774

$ -

Class T

106,263

-

Class B

46,244

-

Class C

67,330

-

Mid Cap Value

36,700,729

13,997,322

Institutional Class

43,114

-

Total

$ 37,140,454

$ 13,997,322

Annual Report

11. Distributions to Shareholders - continued

Distributions to shareholders of each class were as follows: - continued

Years ended January 31,

2008 A

2007

Fidelity Large Cap Growth Fund

From net investment income

Large Cap Growth

$ -

$ 137,207

From net realized gain

Class A

$ 123,133

$ -

Class T

101,048

-

Class B

40,430

-

Class C

75,734

-

Large Cap Growth

18,196,705

3,644,497

Institutional Class

35,354

-

Total

$ 18,572,404

$ 3,644,497

Fidelity Mid Cap Growth Fund

From net realized gain

Class A

$ 100,774

$ -

Class T

34,957

-

Class B

25,358

-

Class C

40,118

-

Mid Cap Growth

21,697,261

5,089,264

Institutional Class

11,029

-

Total

$ 21,909,497

$ 5,089,264

A Distributions for Class A,T,B,C and Institutional are for the period February 13, 2007(commencement of sale of shares) to January 31, 2008.

12. Share Transactions.

Transactions for each class of shares were as follows:

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Fidelity Large Cap Value Fund

Class A

Shares sold

1,045,290

-

$ 16,232,942

$ -

Reinvestment of distributions

29,393

-

423,574

-

Shares redeemed

(352,749)

-

(5,365,826)

-

Net increase (decrease)

721,934

-

$ 11,290,690

$ -

Class T

Shares sold

935,752

-

$ 14,740,653

$ -

Reinvestment of distributions

24,935

-

359,076

-

Shares redeemed

(518,994)

-

(7,932,251)

-

Net increase (decrease)

441,693

-

$ 7,167,478

$ -

Class B

Shares sold

315,386

-

$ 4,951,665

$ -

Reinvestment of distributions

6,739

-

97,191

-

Shares redeemed

(184,721)

-

(2,835,906)

-

Net increase (decrease)

137,404

-

$ 2,212,950

$ -

Class C

Shares sold

138,185

-

$ 2,136,503

$ -

Reinvestment of distributions

3,506

-

50,496

-

Shares redeemed

(52,323)

-

(788,391)

-

Net increase (decrease)

89,368

-

$ 1,398,608

$ -

Annual Report

Notes to Financial Statements - continued

12. Share Transactions - continued

Transactions for each class of shares were as follows: - continued

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Fidelity Large Cap Value Fund - continued

Large Cap Value

Shares sold

52,779,386

72,550,724

$ 813,954,360

$ 1,029,577,278

Reinvestment of distributions

7,056,478

1,672,345

102,030,422

23,955,243

Shares redeemed

(40,898,915)

(25,616,795)

(619,081,485)

(368,491,524)

Net increase (decrease)

18,936,949

48,606,274

$ 296,903,297

$ 685,040,997

Institutional Class

Shares sold

92,735

-

$ 1,397,015

$ -

Reinvestment of distributions

3,477

-

50,127

-

Shares redeemed

(17,951)

-

(257,652)

-

Net increase (decrease)

78,261

-

$ 1,189,490

$ -

Fidelity Mid Cap Value Fund

Class A

Shares sold

615,995

-

$ 10,540,942

$ -

Reinvestment of distributions

11,024

-

178,558

-

Shares redeemed

(132,358)

-

(2,195,988)

-

Net increase (decrease)

494,661

-

$ 8,523,512

$ -

Class T

Shares sold

325,063

-

$ 5,643,056

$ -

Reinvestment of distributions

6,694

-

108,412

-

Shares redeemed

(84,728)

-

(1,422,111)

-

Net increase (decrease)

247,029

-

$ 4,329,357

$ -

Class B

Shares sold

204,574

-

$ 3,623,364

$ -

Reinvestment of distributions

2,791

-

45,364

-

Shares redeemed

(120,358)

-

(2,068,797)

-

Net increase (decrease)

87,007

-

$ 1,599,931

$ -

Class C

Shares sold

168,571

-

$ 2,987,730

$ -

Reinvestment of distributions

3,136

-

50,659

-

Shares redeemed

(61,027)

-

(991,710)

-

Net increase (decrease)

110,680

-

$ 2,046,679

$ -

Mid Cap Value

Shares sold

34,123,012

26,022,900

$ 603,742,926

$ 415,931,492

Reinvestment of distributions

2,338,406

1,003,867

38,409,538

16,337,499

Shares redeemed

(27,133,004)

(10,884,934)

(462,055,715)

(173,050,014)

Net increase (decrease)

9,328,414

16,141,833

$ 180,096,749

$ 259,218,977

Institutional Class

Shares sold

102,472

-

$ 1,790,525

$ -

Reinvestment of distributions

3,101

-

50,277

-

Shares redeemed

(9,162)

-

(148,581)

-

Net increase (decrease)

96,411

-

$ 1,692,221

$ -

Fidelity Large Cap Growth Fund

Class A

Shares sold

165,474

-

$ 1,900,337

$ -

Reinvestment of distributions

10,979

-

116,895

-

Shares redeemed

(44,281)

-

(486,246)

-

Net increase (decrease)

132,172

-

$ 1,530,986

$ -

Class T

Shares sold

167,227

-

$ 1,920,600

$ -

Reinvestment of distributions

9,490

-

101,048

-

Shares redeemed

(65,269)

-

(790,917)

-

Net increase (decrease)

111,448

-

$ 1,230,731

$ -

Annual Report

12. Share Transactions - continued

Transactions for each class of shares were as follows: - continued

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Class B

Shares sold

55,847

-

$ 636,287

$ -

Reinvestment of distributions

3,795

-

40,430

-

Shares redeemed

(4,405)

-

(49,003)

-

Net increase (decrease)

55,237

-

$ 627,714

$ -

Class C

Shares sold

106,310

-

$ 1,230,146

$ -

Reinvestment of distributions

6,439

-

68,402

-

Shares redeemed

(16,516)

-

(191,737)

-

Net increase (decrease)

96,233

-

$ 1,106,811

$ -

Large Cap Growth

Shares sold

5,794,783

10,973,541

$ 66,845,422

$ 124,315,764

Reinvestment of distributions

1,665,396

318,299

17,914,862

3,701,456

Shares redeemed

(7,903,937)

(9,222,108)

(90,877,571)

(102,538,999)

Net increase (decrease)

(443,758)

2,069,732

$ (6,117,287)

$ 25,478,221

Institutional Class

Shares sold

38,550

-

$ 453,557

$ -

Reinvestment of distributions

3,006

-

32,121

-

Shares redeemed

(2,488)

-

(28,969)

-

Net increase (decrease)

39,068

-

$ 456,709

$ -

Fidelity Mid Cap Growth Fund

Class A

Shares sold

165,234

-

$ 2,326,735

$ -

Reinvestment of distributions

7,218

-

97,305

-

Shares redeemed

(13,557)

-

(186,849)

-

Net increase (decrease)

158,895

-

$ 2,237,191

$ -

Class T

Shares sold

54,813

-

$ 796,065

$ -

Reinvestment of distributions

2,601

-

34,957

-

Shares redeemed

(8,726)

-

(121,838)

-

Net increase (decrease)

48,688

-

$ 709,184

$ -

Class B

Shares sold

35,917

-

$ 516,488

$ -

Reinvestment of distributions

1,882

-

25,182

-

Shares redeemed

(3,583)

-

(47,283)

-

Net increase (decrease)

34,216

-

$ 494,387

$ -

Class C

Shares sold

59,795

-

$ 849,458

$ -

Reinvestment of distributions

2,811

-

37,643

-

Shares redeemed

(4,934)

-

(66,440)

-

Net increase (decrease)

57,672

-

$ 820,661

$ -

Mid Cap Growth

Shares sold

7,994,956

24,363,804

$ 114,388,509

$ 336,430,868

Reinvestment of distributions

1,580,341

357,726

21,334,599

4,986,704

Shares redeemed

(15,751,578)

(18,209,467)

(217,016,915)

(243,991,431)

Net increase (decrease)

(6,176,281)

6,512,063

$ (81,293,807)

$ 97,426,141

Institutional Class

Shares sold

14,080

-

$ 199,943

$ -

Reinvestment of distributions

816

-

11,029

-

Net increase (decrease)

14,896

-

$ 210,972

$ -

A Share transactions for class A,T,B,C and Institutional are for the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

Annual Report

Report of Independent Registered Public Accounting Firm

To the Trustees of Fidelity Devonshire Trust and the Shareholders of Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund:

In our opinion, the accompanying statements of assets and liabilities, including the schedules of investments, and the related statements of operations and of changes in net assets and the financial highlights present fairly, in all material respects, the financial position of Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund (funds of Fidelity Devonshire Trust) at January 31, 2008, the results of each of their operations for the year then ended, the changes in each of their net assets for each of the two years in the period then ended and the financial highlights for each of the periods indicated, in conformity with accounting principles generally accepted in the United States of America. These financial statements and financial highlights (hereafter referred to as "financial statements") are the responsibility of the Fidelity Devonshire Trust's management. Our responsibility is to express an opinion on these financial statements based on our audits. We conducted our audits of these financial statements in accordance with the standards of the Public Company Accounting Oversight Board (United States). Those standards require that we plan and perform the audit to obtain reasonable assurance about whether the financial statements are free of material misstatement. An audit includes examining, on a test basis, evidence supporting the amounts and disclosures in the financial statements, assessing the accounting principles used and significant estimates made by management, and evaluating the overall financial statement presentation. We believe that our audits, which included confirmation of securities at January 31, 2008 by correspondence with the custodian and brokers, provide a reasonable basis for our opinion.

/s/ PricewaterhouseCoopers LLP

PricewaterhouseCoopers LLP

Boston, Massachusetts
March 24, 2008

Annual Report

Trustees and Officers

The Trustees, Members of the Advisory Board, and executive officers of the trust and funds, as applicable, are listed below. The Board of Trustees governs each fund and is responsible for protecting the interests of shareholders. The Trustees are experienced executives who meet periodically throughout the year to oversee each fund's activities, review contractual arrangements with companies that provide services to each fund, and review each fund's performance. Except for James C. Curvey, each of the Trustees oversees 374 funds advised by FMR or an affiliate. Mr. Curvey oversees 369 funds advised by FMR or an affiliate.

The Trustees hold office without limit in time except that (a) any Trustee may resign; (b) any Trustee may be removed by written instrument, signed by at least two-thirds of the number of Trustees prior to such removal; (c) any Trustee who requests to be retired or who has become incapacitated by illness or injury may be retired by written instrument signed by a majority of the other Trustees; and (d) any Trustee may be removed at any special meeting of shareholders by a two-thirds vote of the outstanding voting securities of the trust. Each Trustee who is not an interested person (as defined in the 1940 Act) (Independent Trustee), shall retire not later than the last day of the calendar year in which his or her 72nd birthday occurs. The Independent Trustees may waive this mandatory retirement age policy with respect to individual Trustees. The executive officers and Advisory Board Members hold office without limit in time, except that any officer and Advisory Board Member may resign or may be removed by a vote of a majority of the Trustees at any regular meeting or any special meeting of the Trustees. Except as indicated, each individual has held the office shown or other offices in the same company for the past five years.

The funds' Statement of Additional Information (SAI) includes more information about the Trustees. To request a free copy, call Fidelity at 1-800-544-8544.

Interested Trustees*:

Correspondence intended for each Trustee who is an interested person may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Edward C. Johnson 3d (77)

Year of Election or Appointment: 1985

Mr. Johnson is Chairman of the Board of Trustees. Mr. Johnson serves as President (2006-present), Chief Executive Officer, Chairman, and a Director of FMR LLC; Chairman and a Director of FMR; Chairman and a Director of Fidelity Research & Analysis Company (FRAC); Chairman and a Director of Fidelity Investments Money Management, Inc.; and Chairman (2001-present) and a Director of FMR Co., Inc. In addition, Mr. Johnson serves as Chairman and Director of FIL Limited. Mr. Edward C. Johnson 3d and Mr. Arthur E. Johnson are not related.

James C. Curvey (72)

Year of Election or Appointment: 2007

Trustee of Fidelity Devonshire Trust. Mr. Curvey also serves as Trustee (2007-present) or Member of the Advisory Board (2007-present) of other investment companies advised by FMR. Mr. Curvey is a Director of FMR and FMR Co., Inc. (2007-present). Mr. Curvey is also Vice Chairman (2006-present) and Director of FMR LLC. Mr. Curvey joined Fidelity in 1982 and served in numerous senior management positions, including President and Chief Operating Officer of FMR LLC (1997-2000) and President of Fidelity Strategic Investments (2000-2002). In addition, he serves as a member of the Board of Directors of Geerlings & Wade, Inc. (wine distribution).

* Trustees have been determined to be "Interested Trustees" by virtue of, among other things, their affiliation with the trust or various entities under common control with FMR. FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Independent Trustees:

Correspondence intended for each Independent Trustee (that is, the Trustees other than the Interested Trustees) may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235.

Name, Age; Principal Occupation

Dennis J. Dirks (59)

Year of Election or Appointment: 2005

Prior to his retirement in May 2003, Mr. Dirks was Chief Operating Officer and a member of the Board of The Depository Trust & Clearing Corporation (DTCC) (1999-2003). He also served as President, Chief Operating Officer, and Board member of The Depository Trust Company (DTC) (1999-2003) and President and Board member of the National Securities Clearing Corporation (NSCC) (1999-2003). In addition, Mr. Dirks served as Chief Executive Officer and Board member of the Government Securities Clearing Corporation (2001-2003) and Chief Executive Officer and Board member of the Mortgage-Backed Securities Clearing Corporation (2001-2003). Mr. Dirks also serves as a Trustee and a member of the Finance Committee of Manhattan College (2005-present) and a Trustee and a member of the Finance Committee of AHRC of Nassau County (2006-present).

Albert R. Gamper, Jr. (65)

Year of Election or Appointment: 2006

Prior to his retirement in December 2004, Mr. Gamper served as Chairman of the Board of CIT Group Inc. (commercial finance). During his tenure with CIT Group Inc. Mr. Gamper served in numerous senior management positions, including Chairman (1987-1989; 1999-2001; 2002-2004), Chief Executive Officer (1987-2004), and President (1989-2002). He currently serves as a member of the Board of Directors of Public Service Enterprise Group (utilities, 2001-present), Chairman of the Board of Governors, Rutgers University (2004-present), and Chairman of the Board of Saint Barnabas Health Care System.

George H. Heilmeier (71)

Year of Election or Appointment: 2004

Dr. Heilmeier is Chairman Emeritus of Telcordia Technologies (communication software and systems), where prior to his retirement, he served as company Chairman and Chief Executive Officer. He currently serves on the Boards of Directors of The Mitre Corporation (systems engineering and information technology support for the government), and HRL Laboratories (private research and development, 2004-present). He is Chairman of the General Motors Science & Technology Advisory Board and a Life Fellow of the Institute of Electrical and Electronics Engineers (IEEE). Dr. Heilmeier is a member of the Defense Science Board and the National Security Agency Advisory Board. He is also a member of the National Academy of Engineering, the American Academy of Arts and Sciences, and the Board of Overseers of the School of Engineering and Applied Science of the University of Pennsylvania. Previously, Dr. Heilmeier served as a Director of TRW Inc. (automotive, space, defense, and information technology, 1992-2002), Compaq (1994-2002), Automatic Data Processing, Inc. (ADP) (technology-based business outsourcing, 1995-2002), INET Technologies Inc. (telecommunications network surveillance, 2001-2004), and Teletech Holdings (customer management services). He is the recipient of the 2005 Kyoto Prize in Advanced Technology for his invention of the liquid crystal display, and a member of the Consumer Electronics Hall of Fame.

James H. Keyes (67)

Year of Election or Appointment: 2007

Prior to his retirement in 2003, Mr. Keyes was Chairman, President, and Chief Executive Officer of Johnson Controls, Inc. (automotive supplier, 1993-2003). He currently serves as a member of the boards of LSI Logic Corporation (semiconductor technologies), Navistar International Corporation (manufacture and sale of trucks, buses, and diesel engines, 2002-present), and Pitney Bowes, Inc. (integrated mail, messaging, and document management solutions).

Marie L. Knowles (61)

Year of Election or Appointment: 2001

Prior to Ms. Knowles' retirement in June 2000, she served as Executive Vice President and Chief Financial Officer of Atlantic Richfield Company (ARCO) (diversified energy, 1996-2000). From 1993 to 1996, she was a Senior Vice President of ARCO and President of ARCO Transportation Company. She served as a Director of ARCO from 1996 to 1998. She currently serves as a Director of Phelps Dodge Corporation (copper mining and manufacturing) and McKesson Corporation (healthcare service, 2002-present). Ms. Knowles is a Trustee of the Brookings Institution and the Catalina Island Conservancy and also serves as a member of the Advisory Board for the School of Engineering of the University of Southern California.

Ned C. Lautenbach (63)

Year of Election or Appointment: 2000

Mr. Lautenbach is Chairman of the Independent Trustees (2006-present). Mr. Lautenbach has been a partner of Clayton, Dubilier & Rice, Inc. (private equity investment firm) since September 1998. Previously, Mr. Lautenbach was with the International Business Machines Corporation (IBM) from 1968 until his retirement in 1998. Mr. Lautenbach serves as a Director of Sony Corporation (2006-present) and Eaton Corporation (diversified industrial) as well as the Philharmonic Center for the Arts in Naples, Florida. He also is a member of the Board of Trustees of Fairfield University (2005-present), as well as a member of the Council on Foreign Relations.

Cornelia M. Small (63)

Year of Election or Appointment: 2005

Ms. Small is a member (2000-present) and Chairperson (2002-present) of the Investment Committee, and a member (2002-present) of the Board of Trustees of Smith College. Previously, she served as Chief Investment Officer (1999-2000), Director of Global Equity Investments (1996-1999), and a member of the Board of Directors of Scudder, Stevens & Clark (1990-1997) and Scudder Kemper Investments (1997-1999). In addition, Ms. Small served as Co-Chair (2000-2003) of the Annual Fund for the Fletcher School of Law and Diplomacy.

William S. Stavropoulos (68)

Year of Election or Appointment: 2002

Mr. Stavropoulos is Chairman Emeritus of the Board of Directors of The Dow Chemical Company. Since joining The Dow Chemical Company in 1967, Mr. Stavropoulos served in numerous senior management positions, including President (1993-2000; 2002-2003), CEO (1995-2000; 2002-2004), and Chairman of the Executive Committee (2000-2004). Currently, he is a Director of NCR Corporation (data warehousing and technology solutions), Chemical Financial Corporation, Maersk Inc. (industrial conglomerate, 2002-present), Tyco International, Inc. (multinational manufacturing and services, 2007-present), and a member of the Advisory Board for Metalmark Capital (private equity investment firm, 2005-present). He is a special advisor to Clayton, Dubilier & Rice, Inc., a private equity investment firm. He also serves as a member of the Board of Trustees of the American Enterprise Institute for Public Policy Research. In addition, Mr. Stavropoulos is a member of The Business Council, J.P. Morgan International Council and the University of Notre Dame Advisory Council for the College of Science.

Kenneth L. Wolfe (68)

Year of Election or Appointment: 2005

Mr. Wolfe is Chairman and a Director of Hershey Foods Corporation (2007-present), where prior to his retirement in 2001, he was Chairman and Chief Executive Officer. Mr. Wolfe currently serves as a member of the board of Revlon Inc. (2004-present). Previously, Mr. Wolfe served as a member of the boards of Adelphia Communications Corporation (2003-2006) and Bausch & Lomb, Inc. (1993-2007).

Advisory Board Members and Executive Officers**:

Correspondence intended for Mr. Mauriello, Mr. Thomas, Mr. Wiley, Mr. Lacy, and Mr. Arthur Johnson may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235. Correspondence intended for each executive officer and Mr. Lynch may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Arthur E. Johnson (61)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Johnson serves as Senior Vice President of Corporate Strategic Development of Lockheed Martin Corporation (defense contractor). In addition, Mr. Johnson serves as a member of the Board of Directors of AGL Resources, Inc. (holding company, 2002-present), and IKON Office Solutions, Inc. (document management systems and services). Mr. Arthur E. Johnson and Mr. Edward C. Johnson 3d are not related.

Alan J. Lacy (54)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lacy serves as Senior Adviser (2007-present) of Oak Hill Capital Partners, L.P. (a private equity firm). Mr. Lacy also served as Vice Chairman and Chief Executive Officer of Sears Holdings Corporation and Sears, Roebuck and Co. (retail, 2005-2006; 2000-2005). In addition, Mr. Lacy serves as a member of the Board of Directors of The Western Union Company (global money transfer, 2006-present) and Bristol-Myers Squibb (global pharmaceuticals, 2007-present). Mr. Lacy is a Trustee of the National Parks Conservation Association and The Field Museum of Natural History.

Peter S. Lynch (64)

Year of Election or Appointment: 2003

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lynch is Vice Chairman and a Director of FMR, and Vice Chairman (2001-present) and a Director of FMR Co., Inc. Previously, Mr. Lynch served as a Trustee of the Fidelity funds (1990-2003). In addition, he serves as a Trustee of Boston College and as the Chairman of the Inner-City Scholarship Fund.

Joseph Mauriello (63)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Prior to his retirement in January 2006, Mr. Mauriello served in numerous senior management positions including Deputy Chairman and Chief Operating Officer (2004-2005), and Vice Chairman of Financial Services (2002-2004) of KPMG LLP US (professional services firm, 1965-2005). Mr. Mauriello currently serves as a member of the Board of Directors of XL Capital Ltd. (global insurance and re-insurance company, 2006-present) and of Arcadia Resources Inc. (health care services and products, 2007-present). He also served as a Director of the Hamilton Funds of the Bank of New York (2006-2007).

David M. Thomas (58)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Previously, Mr. Thomas served as Executive Chairman (2005-2006) and Chairman and Chief Executive Officer (2000-2005) of IMS Health, Inc. (pharmaceutical and healthcare information solutions). In addition, Mr. Thomas serves as a member of the Board of Directors of Fortune Brands, Inc. (consumer products holding company), and Interpublic Group of Companies, Inc. (marketing communication, 2004-present).

Michael E. Wiley (57)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Wiley also serves as a member of the Board of Trustees of the University of Tulsa (2000-2006; 2007-present). He serves as a Director of Tesoro Corporation (independent oil refiner and marketer, 2005-present), and a Director of Bill Barrett Corporation (exploration and production company, 2005-present). In addition, he also serves as a Director of Post Oak Bank (privately-held bank, 2004-present). Previously, Mr. Wiley served as a Sr. Energy Advisor of Katzenbach Partners, LLC (consulting firm, 2006-2007), as an Advisory Director of Riverstone Holdings (private investment firm), Chairman, President, and CEO of Baker Hughes, Inc. (oilfield services company, 2000-2004), and as Director of Spinnaker Exploration Company (exploration and production company, 2001-2005).

Kimberley H. Monasterio (44)

Year of Election or Appointment: 2007

President and Treasurer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Ms. Monasterio also serves as President and Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2004-present). Previously, Ms. Monasterio served as Deputy Treasurer of the Fidelity funds (2004-2006). Before joining Fidelity Investments, Ms. Monasterio served as Treasurer (2000-2004) and Chief Financial Officer (2002-2004) of the Franklin Templeton Funds and Senior Vice President of Franklin Templeton Services, LLC (2000-2004).

Walter C. Donovan (45)

Year of Election or Appointment: 2007

Vice President of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Donovan also serves as Vice President of Fidelity's Equity Funds (2007-present). Mr. Donovan also serves as Executive Vice President of FMR (2005-present) and FMR Co., Inc. (2005-present). Previously, Mr. Donovan served as Vice President of Fidelity's High Income Funds (2005-2007), Fixed-Income Funds (2005-2006), certain Asset Allocation Funds (2005-2006), certain Balanced Funds (2005-2006), and as Vice President and Director of Fidelity's International Equity Trading group (1998-2005).

Bruce T. Herring (42)

Year of Election or Appointment: 2006

Vice President of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Herring also serves as Vice President of certain Equity Funds (2006-present). Mr. Herring is Senior Vice President of FMR (2006-present) and Vice President of FMR Co., Inc. (2001-present). Previously, Mr. Herring served as a portfolio manager for Fidelity U.S. Equity Funds (2001-2005).

Brian B. Hogan (43)

Year of Election or Appointment: 2007

Vice President of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Hogan also serves as Vice President of Sector Funds (2007-present). Mr. Hogan is Senior Vice President of Equity Research (2006-present). Mr. Hogan also serves as Vice President of FMR and FMR Co., Inc. Previously, Mr. Hogan served as a portfolio manager.

Eric D. Roiter (59)

Year of Election or Appointment: 2001

Secretary of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. He also serves as Secretary of other Fidelity funds; Vice President, General Counsel, and Secretary of FMR Co., Inc. (2001-present) and FMR; Assistant Secretary of Fidelity Management & Research (U.K.) Inc. (2001-present), Fidelity Research & Analysis Company (2001-present), and Fidelity Investments Money Management, Inc. (2001-present). Mr. Roiter is an Adjunct Member, Faculty of Law, at Boston College Law School (2003-present). Previously, Mr. Roiter served as Vice President and Secretary of Fidelity Distributors Corporation (FDC) (1998-2005).

John B. McGinty, Jr. (45)

Year of Election or Appointment: 2008

Assistant Secretary of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. McGinty also serves as Assistant Secretary of other Fidelity funds (2008-present) and is an employee of FMR LLC (2004-present). Mr. McGinty also serves as Senior Vice President, Secretary, and Chief Legal Officer of FDC (2007-present). Before joining Fidelity Investments, Mr. McGinty practiced law at Ropes & Gray, LLP.

R. Stephen Ganis (41)

Year of Election or Appointment: 2006

Anti-Money Laundering (AML) officer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Ganis also serves as AML officer of other Fidelity funds (2006-present) and FMR LLC (2003-present). Before joining Fidelity Investments, Mr. Ganis practiced law at Goodwin Procter, LLP (2000-2002).

Joseph B. Hollis (59)

Year of Election or Appointment: 2006

Chief Financial Officer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Hollis also serves as Chief Financial Officer of other Fidelity funds. Mr. Hollis is President of Fidelity Pricing and Cash Management Services (FPCMS) (2005-present). Mr. Hollis also serves as President and Director of Fidelity Service Company, Inc. (2006-present). Previously, Mr. Hollis served as Senior Vice President of Cash Management Services (1999-2002) and Investment Management Operations (2002-2005).

Kenneth A. Rathgeber (60)

Year of Election or Appointment: 2004

Chief Compliance Officer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Rathgeber also serves as Chief Compliance Officer of other Fidelity funds (2004-present) and Executive Vice President of Risk Oversight for Fidelity Investments (2002-present). He is Chief Compliance Officer of FMR (2005-present), FMR Co., Inc. (2005-present), Fidelity Management & Research (U.K.) Inc. (2005-present), Fidelity Research & Analysis Company (2005-present), Fidelity Investments Money Management, Inc. (2005-present), and Strategic Advisers, Inc. (2005-present). Previously, Mr. Rathgeber served as Executive Vice President and Chief Operating Officer for Fidelity Investments Institutional Services Company, Inc. (1998-2002).

Bryan A. Mehrmann (46)

Year of Election or Appointment: 2005

Deputy Treasurer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Mehrmann also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR. Previously, Mr. Mehrmann served as Vice President of Fidelity Investments Institutional Services Group (FIIS)/Fidelity Investments Institutional Operations Corporation, Inc. (FIIOC) Client Services (1998-2004).

Kenneth B. Robins (38)

Year of Election or Appointment:2005

Deputy Treasurer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Robins also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2004-present). Before joining Fidelity Investments, Mr. Robins worked at KPMG LLP, where he was a partner in KPMG's department of professional practice (2002-2004) and a Senior Manager (1999-2000). In addition, Mr. Robins served as Assistant Chief Accountant, United States Securities and Exchange Commission (2000-2002).

Robert G. Byrnes (41)

Year of Election or Appointment: 2005

Assistant Treasurer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Byrnes also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Byrnes served as Vice President of FPCMS (2003-2005). Before joining Fidelity Investments, Mr. Byrnes worked at Deutsche Asset Management where he served as Vice President of the Investment Operations Group (2000-2003).

Peter L. Lydecker (54)

Year of Election or Appointment: 2004

Assistant Treasurer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Lydecker also serves as Assistant Treasurer of other Fidelity funds (2004) and is an employee of FMR.

Paul M. Murphy (60)

Year of Election or Appointment: 2007

Assistant Treasurer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Murphy also serves as Assistant Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2007-present). Previously, Mr. Murphy served as Chief Financial Officer of the Fidelity Funds (2005-2006), Vice President and Associate General Counsel of FMR (2007), and Senior Vice President of Fidelity Pricing and Cash Management Services Group (FPCMS) (1994-2007).

Name, Age; Principal Occupation

Gary W. Ryan (49)

Year of Election or Appointment: 2005

Assistant Treasurer of Large Cap Growth, Large Cap Value, Mid Cap Growth, and Mid Cap Value. Mr. Ryan also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Ryan served as Vice President of Fund Reporting in FPCMS (1999-2005).

** FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Annual Report

Distributions

The funds hereby designate as capital gain dividend the amounts noted below for the taxable year ended January 31, 2008, or, if subsequently determined to be different, the net capital gain of such year.

Fidelity Large Cap Value Fund

$53,592,103

Fidelity Mid Cap Value Fund

$30,917,128

Fidelity Large Cap Growth Fund

$9,534,777

Fidelity Mid Cap Growth Fund

$25,157,607

A percentage of the dividends distributed during the fiscal year for the following funds qualifies for the dividends-received deduction for corporate shareholders:

March,
2007

December, 2007

Fidelity Large Cap Value Fund

45%

100%

Fidelity Mid Cap Value Fund

26%

100%

Fidelity Large Cap Growth Fund

91%

18%

A percentage of the dividends distributed during the fiscal year for the following funds may be taken into account as a dividend for purposes of the maximum rate under section 1(h)(11) of the Internal Revenue Code.

March,
2007

December, 2007

Fidelity Large Cap Value Fund

45%

100%

Fidelity Mid Cap Value Fund

26%

100%

Fidelity Large Cap Growth Fund

92%

20%

The fund will notify shareholders in January 2009 of amounts for use in preparing 2008 income tax returns.

Annual Report

Managing Your Investments

Fidelity offers several ways to conveniently manage your personal investments via your telephone or PC. You can access your account information, conduct trades and research your investments 24 hours a day.

By Phone

Fidelity Automated Service Telephone provides a single toll-free number to access account balances, positions, quotes and trading. It's easy to navigate the service, and on your first call, the system will help you create a personal identification number (PIN) for security.

(phone_graphic)

Fidelity Automated Service Telephone (FAST®)
1-800-544-5555

Press

1   For mutual fund and brokerage trading.

2   For quotes.*

3   For account balances and holdings.

4   To review orders and mutual fund activity.

5   To change your PIN.

*0   To speak to a Fidelity representative.

By PC

Fidelity's web site on the Internet provides a wide range of information, including daily financial news, fund performance, interactive planning tools and news about Fidelity products and services.

(computer_graphic)

Fidelity's Web Site
www.fidelity.com

* When you call the quotes line, please remember that a fund's yield and return will vary and, except for money market funds, share price will also vary. This means that you may have a gain or loss when you sell your shares. There is no assurance that money market funds will be able to maintain a stable $1 share price; an investment in a money market fund is not insured or guaranteed by the U.S. government. Total returns are historical and include changes in share price, reinvestment of dividends and capital gains, and the effects of any sales charges.

Annual Report

To Write Fidelity

We'll give your correspondence immediate attention and send you written confirmation upon completion of your request.

(letter_graphic)

Making Changes
To Your Account

(such as changing name, address, bank, etc.)

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0002

(letter_graphic)

For Non-Retirement
Accounts

Buying shares

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0003

Overnight Express
Fidelity Investments
Attn: Distribution Services
100 Crosby Parkway - KC1H
Covington, KY 41015

Selling shares

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0035

Overnight Express
Fidelity Investments
Attn: Distribution Services
100 Crosby Parkway - KC1H
Covington, KY 41015

General Correspondence

Fidelity Investments
P.O. Box 500
Merrimack, NH 03054-0500

(letter_graphic)

For Retirement
Accounts

Buying shares

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0003

Selling shares

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0035

Overnight Express
Fidelity Investments
Attn: Distribution Services
100 Crosby Parkway - KC1H
Covington, KY 41015

General Correspondence

Fidelity Investments
P.O. Box 500
Merrimack, NH 03054-0500

Annual Report

Investment Adviser

Fidelity Management & Research Company
Boston, MA

Investment Sub-Adviser

FMR Co., Inc.

Fidelity Management & Research
(U.K.) Inc.

Fidelity Research & Analysis Company

Fidelity Investments Japan Limited

Fidelity International Investment Advisors

Fidelity International Investment Advisors
(U.K.) Limited

General Distributor

Fidelity Distributors Corporation
Boston, MA

Transfer and Service Agents

Fidelity Investments Institutional Operations
Company, Inc.
Boston, MA

Fidelity Service Company, Inc.
Boston, MA

Custodian

Citibank, N.A.
New York, NY

The Fidelity Telephone Connection

Mutual Fund 24-Hour Service

Exchanges/Redemptions
and Account Assistance 1-800-544-6666

Product Information 1-800-544-6666

Retirement Accounts 1-800-544-4774
(8 a.m. - 9 p.m.)

TDD Service 1-800-544-0118
for the deaf and hearing impaired
(9 a.m. - 9 p.m. Eastern time)

Fidelity Automated Service
Telephone (FAST®) (automated graphic)    1-800-544-5555

(automated graphic)    Automated line for quickest service

(Fidelity Investment logo)(registered trademark)
Corporate Headquarters
82 Devonshire St., Boston, MA 02109
www.fidelity.com

LMC-UANN-0308
1.789259.105

Fidelity Advisor Large Cap Value

Class A, Class T, Class B, and Class C

Fidelity Advisor Mid Cap Value

Class A, Class T, Class B, and Class C

Fidelity Advisor Large Cap Growth

Class A, Class T, Class B, and Class C

Fidelity Advisor Mid Cap Growth

Class A, Class T, Class B, and Class C

Funds

Annual Report

January 31, 2008

(2_fidelity_logos) (Registered_Trademark)

Each Class A, Class T, Class B, and Class C
are classes of
Fidelity® Large Cap Value,
Fidelity Mid Cap Value,
Fidelity Large Cap Growth, and
Fidelity Mid Cap Growth
Funds

Contents

Chairman's Message

<Click Here>

Ned Johnson's message to shareholders.

Shareholder Expense Example

<Click Here>

An example of shareholder expenses.

Fidelity Advisor Large Cap Value Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Advisor Mid Cap Value Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Advisor Large Cap Growth Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Advisor Mid Cap Growth Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Notes

<Click Here>

Notes to the Financial Statements

Report of Independent Registered Public Accounting Firm

<Click Here>

Trustees and Officers

<Click Here>

Distributions

<Click Here>

To view a fund's proxy voting guidelines and proxy voting record for the 12-month period ended June 30, visit http://www.fidelity.com (search for "proxy voting guidelines") or visit the Securities and Exchange Commission's (SEC) web site at http://www.sec.gov. You may also call 1-800-544-8544 to request a free copy of the proxy voting guidelines.

Standard & Poor's, S&P and S&P 500 are registered service marks of The McGraw-Hill Companies, Inc. and have been licensed for use by Fidelity Distributors Corporation.

Other third party marks appearing herein are the property of their respective owners.

All other marks appearing herein are registered or unregistered trademarks or service marks of FMR LLC or an affiliated company.

This report and the financial statements contained herein are submitted for the general information of the shareholders of the funds. This report is not authorized for distribution to prospective investors in the funds unless preceded or accompanied by an effective prospectus.

A fund files its complete schedule of portfolio holdings with the SEC for the first and third quarters of each fiscal year on Form N-Q. Forms N-Q are available on the SEC's web site at http://www.sec.gov. A fund's Forms N-Q may be reviewed and copied at the SEC's Public Reference Room in Washington, DC. Information regarding the operation of the SEC's Public Reference Room may be obtained by calling 1-800-SEC-0330. For a complete list of a fund's portfolio holdings, view the most recent holdings listing, semiannual report, or annual report on Fidelity's web site at http://www.fidelity.com or http://www.advisor.fidelity.com, as applicable.

NOT FDIC INSURED · MAY LOSE VALUE · NO BANK GUARANTEE

Neither the funds nor Fidelity Distributors Corporation is a bank.

Annual Report

Chairman's Message

(Photograph of Edward C. Johnson 3d.)

Dear Shareholder:

Stocks got off to a poor start in 2008, while investment-grade bonds and money markets showed positive returns, once again underscoring the importance of a diversified portfolio. Financial markets are always unpredictable, but there are a number of time-tested principles that can put the historical odds in your favor.

One of the basic tenets is to invest for the long term. Over time, riding out the markets' inevitable ups and downs has proven much more effective than selling into panic or chasing the hottest trend. Even missing only a few of the markets' best days can significantly diminish investor returns. Patience also affords the benefits of compounding - of earning interest on additional income or reinvested dividends and capital gains. There are tax advantages and cost benefits to consider as well. The more you sell, the more taxes you pay, and the more you trade, the higher the costs. While staying the course doesn't eliminate risk, it can considerably lessen the effect of short-term declines.

You can further manage your investing risk through diversification. And today, more than ever, geographic diversification should be taken into account. Studies

indicate that asset allocation is the single most important determinant of a portfolio's long-term success. The right mix of stocks, bonds and cash - aligned to your particular risk tolerance and investment objective - is very important. Age-appropriate rebalancing is also an essential aspect of asset allocation. For younger investors, an emphasis on equities - which historically have been the best-performing asset class over time - is encouraged. As investors near their specific goal, such as retirement or sending a child to college, consideration may be given to replacing volatile assets (e.g. common stocks) with more-stable fixed investments (bonds or savings plans).

A third investment principle - investing regularly - can help lower the average cost of your purchases. Investing a certain amount of money each month or quarter helps ensure you won't pay for all your shares at market highs. This strategy - known as dollar cost averaging - also reduces unconstructive "emotion" from investing, helping shareholders avoid selling weak performers just prior to an upswing, or chasing a hot performer just before a correction.

We invite you to contact us via the Internet, through our Investor Centers or over the phone. It is our privilege to provide you the information you need to make the investments that are right for you.

Sincerely,

/s/Edward C. Johnson 3d

Edward C. Johnson 3d

Annual Report

Shareholder Expense Example

As a shareholder of a Fund, you incur two types of costs: (1) transaction costs, including sales charges (loads) on purchase payments or redemption proceeds, redemption fees, and (2) ongoing costs, including management fees, distribution and/or service (12b-1) fees and other Fund expenses. This Example is intended to help you understand your ongoing costs (in dollars) of investing in the Funds and to compare these costs with the ongoing costs of investing in other mutual funds.

The Example is based on an investment of $1,000 invested at the beginning of the period and held for the entire period (August 1, 2007 to January 31, 2008).

Actual Expenses

The first line of the accompanying table for each class of each fund provides information about actual account values and actual expenses. You may use the information in this line, together with the amount you invested, to estimate the expenses that you paid over the period. Simply divide your account value by $1,000.00 (for example, an $8,600 account value divided by $1,000.00 = 8.6), then multiply the result by the number in the first line for a class of the fund under the heading entitled "Expenses Paid During Period" to estimate the expenses you paid on your account during this period. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, each Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Hypothetical Example for Comparison Purposes

The second line of the accompanying table for each class of each fund provides information about hypothetical account values and hypothetical expenses based on a Class' actual expense ratio and an assumed rate of return of 5% per year before expenses, which is not the Class' actual return. The hypothetical account values and expenses may not be used to estimate the actual ending account balance or expenses you paid for the period. You may use this information to compare the ongoing costs of investing in the Fund and other funds. To do so, compare this 5% hypothetical example with the 5% hypothetical examples that appear in the shareholder reports of the other funds. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, each Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Please note that the expenses shown in the table are meant to highlight your ongoing costs only and do not reflect any transaction costs. Therefore, the second line of the table is useful in comparing ongoing costs only, and will not help you determine the relative total costs of owning different funds. In addition, if these transactional costs were included, your costs would have been higher.

Beginning
Account Value
August 1, 2007

Ending
Account Value
January 31, 2008

Expenses Paid
During Period
*
August 1, 2007
to January 31, 2008

Fidelity Large Cap Value Fund

Class A

Actual

$ 1,000.00

$ 946.40

$ 6.13

HypotheticalA

$ 1,000.00

$ 1,018.90

$ 6.36

Class T

Actual

$ 1,000.00

$ 944.70

$ 7.35

HypotheticalA

$ 1,000.00

$ 1,017.64

$ 7.63

Class B

Actual

$ 1,000.00

$ 943.10

$ 9.80

HypotheticalA

$ 1,000.00

$ 1,015.12

$ 10.16

Class C

Actual

$ 1,000.00

$ 943.20

$ 9.65

HypotheticalA

$ 1,000.00

$ 1,015.27

$ 10.01

Large Cap Value

Actual

$ 1,000.00

$ 948.00

$ 4.27

HypotheticalA

$ 1,000.00

$ 1,020.82

$ 4.43

Institutional Class

Actual

$ 1,000.00

$ 948.00

$ 4.22

HypotheticalA

$ 1,000.00

$ 1,020.87

$ 4.38

Fidelity Mid Cap Value Fund

Class A

Actual

$ 1,000.00

$ 912.50

$ 5.54

HypotheticalA

$ 1,000.00

$ 1,019.41

$ 5.85

Beginning
Account Value
August 1, 2007

Ending
Account Value
January 31, 2008

Expenses Paid
During Period
*
August 1, 2007
to January 31, 2008

Class T

Actual

$ 1,000.00

$ 911.80

$ 6.79

HypotheticalA

$ 1,000.00

$ 1,018.10

$ 7.17

Class B

Actual

$ 1,000.00

$ 909.00

$ 9.09

HypotheticalA

$ 1,000.00

$ 1,015.68

$ 9.60

Class C

Actual

$ 1,000.00

$ 909.00

$ 9.24

HypotheticalA

$ 1,000.00

$ 1,015.53

$ 9.75

Mid Cap Value

Actual

$ 1,000.00

$ 913.40

$ 4.05

HypotheticalA

$ 1,000.00

$ 1,020.97

$ 4.28

Institutional Class

Actual

$ 1,000.00

$ 913.70

$ 4.34

HypotheticalA

$ 1,000.00

$ 1,020.67

$ 4.58

Fidelity Large Cap Growth Fund

Class A

Actual

$ 1,000.00

$ 955.40

$ 5.91

HypotheticalA

$ 1,000.00

$ 1,019.16

$ 6.11

Class T

Actual

$ 1,000.00

$ 955.60

$ 7.30

HypotheticalA

$ 1,000.00

$ 1,017.74

$ 7.53

Class B

Actual

$ 1,000.00

$ 952.20

$ 9.74

HypotheticalA

$ 1,000.00

$ 1,015.22

$ 10.06

Class C

Actual

$ 1,000.00

$ 953.00

$ 9.65

HypotheticalA

$ 1,000.00

$ 1,015.32

$ 9.96

Large Cap Growth

Actual

$ 1,000.00

$ 956.60

$ 4.78

HypotheticalA

$ 1,000.00

$ 1,020.32

$ 4.94

Institutional Class

Actual

$ 1,000.00

$ 958.20

$ 4.10

HypotheticalA

$ 1,000.00

$ 1,021.02

$ 4.23

Fidelity Mid Cap Growth Fund

Class A

Actual

$ 1,000.00

$ 909.40

$ 5.29

HypotheticalA

$ 1,000.00

$ 1,019.66

$ 5.60

Class T

Actual

$ 1,000.00

$ 905.80

$ 6.58

HypotheticalA

$ 1,000.00

$ 1,018.30

$ 6.97

Class B

Actual

$ 1,000.00

$ 904.90

$ 8.79

HypotheticalA

$ 1,000.00

$ 1,015.98

$ 9.30

Class C

Actual

$ 1,000.00

$ 904.20

$ 8.88

HypotheticalA

$ 1,000.00

$ 1,015.88

$ 9.40

Mid Cap Growth

Actual

$ 1,000.00

$ 909.50

$ 3.75

HypotheticalA

$ 1,000.00

$ 1,021.27

$ 3.97

Institutional Class

Actual

$ 1,000.00

$ 909.60

$ 3.32

HypotheticalA

$ 1,000.00

$ 1,021.73

$ 3.52

A 5% return per year before expenses

* Expenses are equal to each Class' annualized expense ratio (shown in the table below); multiplied by the average account value over the period, multiplied by 184/ 365 (to reflect the one-half year period).

Annual Report

Annualized
Expense Ratio

Fidelity Large Cap Value Fund

Class A

1.25%

Class T

1.50%

Class B

2.00%

Class C

1.97%

Large Cap Value

.87%

Institutional Class

.86%

Fidelity Mid Cap Value Fund

Class A

1.15%

Class T

1.41%

Class B

1.89%

Class C

1.92%

Mid Cap Value

.84%

Institutional Class

.90%

Fidelity Large Cap Growth Fund

Class A

1.20%

Class T

1.48%

Class B

1.98%

Class C

1.96%

Large Cap Growth

.97%

Institutional Class

.83%

Fidelity Mid Cap Growth Fund

Class A

1.10%

Class T

1.37%

Class B

1.83%

Class C

1.85%

Mid Cap Growth

.78%

Institutional Class

.69%

Annual Report

Fidelity Advisor Large Cap Value Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the class' dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table does not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Class A (incl. 5.75% sales charge) B

-10.16%

12.69%

6.75%

Class T (incl. 3.50% sales charge) C

-8.31%

13.16%

7.10%

Class B (incl. contingent deferred sales charge) D

-9.85%

13.63%

7.64%

Class C (incl. contingent deferred sales charge) E

-6.15%

13.90%

7.66%

A From November 15, 2001.

B Class A shares bear a 0.25% 12b-1 fee. The initial offering of Class A shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Value, the original class of the fund, which has no 12b-1 fee. Had Class A shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower.

C Class T shares bear a 0.50% 12b-1 fee. The initial offering of Class T shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Value, the original class of the fund, which has no 12b-1 fee. Had Class T shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower.

D Class B shares bear a 1.00% 12b-1 fee. The initial offering of Class B shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Value, the original class of the fund, which has no 12b-1 fee. Had Class B shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower. Class B shares' contingent deferred sales charge included in the past one year, past five years, and life of fund total return figures are 5%, 2%, and 0%, respectively.

E Class C shares bear a 1.00% 12b-1 fee. The initial offering of Class C shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Value, the original class of the fund, which has no 12b-1 fee. Had Class C shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower. Class C shares' contingent deferred sales charge included in the past one year, past five years, and life of fund total return figures are 1%, 0%, and 0%, respectively.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Fidelity Advisor Large Cap Value Fund - Class A on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell 1000® Value Index performed over the same period. The initial offering of Class A took place on February 13, 2007. See above for additional information regarding the performance of Class A.



Annual Report

Fidelity Advisor Large Cap Value Fund

Management's Discussion of Fund Performance

Comments from Bruce Dirks, Portfolio Manager of Fidelity Advisor Large Cap Value Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the year, Advisor Large Cap Value's Class A, Class T, Class B and Class C shares (excluding sales charges) all finished about even with or slightly ahead of the Russell 1000® Value Index, which was off 5.38%. (For specific class-level results, please refer to the performance section of this report.) During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among banks and real estate investment trusts (REITs). Favorable industry weightings within the weak financials and consumer discretionary sectors also added value, as did good stock picking in health care and pockets of information technology. Underweighting diversified financials giant Citigroup and domestic banking firm Wachovia added to relative performance, as both stocks fared poorly within an environment of deteriorating business fundamentals for financials. Among other strong contributors were Cummins, a diesel engine manufacturer, and CF Industries, an out-of-index company that produces fertilizer products. On the flip side, relative results were held back by unfavorable stock picks in consumer staples - especially among food retailers - energy and the transportation industry. Among the biggest detractors were Tyson Foods, a large processor of poultry and meat products, and YRC Worldwide, a transportation company. Underweighting such strong-performing energy stocks as Occidental Petroleum also hurt. Tyson, YRC and Occidental were no longer in the portfolio at period end.

For the year, Advisor Large Cap Value's Institutional Class shares finished ahead of the Russell 1000® Value Index, which was off 5.38%. (For specific class-level results, please refer to the performance section of this report.) During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among banks and real estate investment trusts (REITs). Favorable industry weightings within the weak financials and consumer discretionary sectors also added value, as did good stock picking in health care and pockets of information technology. Underweighting diversified financials giant Citigroup and domestic banking firm Wachovia added to relative performance, as both stocks fared poorly within an environment of deteriorating business fundamentals for financials. Among other strong contributors were Cummins, a diesel engine manufacturer, and CF Industries, an out-of-index company that produces fertilizer products. On the flip side, relative results were held back by unfavorable stock picks in consumer staples - especially among food retailers - energy and the transportation industry. Among the biggest detractors were Tyson Foods, a large processor of poultry and meat products, and YRC Worldwide, a transportation company. Underweighting such strong-performing energy stocks as Occidental Petroleum also hurt. Tyson, YRC and Occidental were no longer in the portfolio at period end.

The views expressed above reflect those of the portfolio manager only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Large Cap Value Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Exxon Mobil Corp.

4.3

4.5

AT&T, Inc.

3.8

3.8

General Electric Co.

3.4

3.0

ConocoPhillips

3.1

0.0

JPMorgan Chase & Co.

3.0

3.0

American International Group, Inc.

2.7

2.3

Chevron Corp.

2.7

2.5

Bank of America Corp.

2.4

2.4

Altria Group, Inc.

2.3

1.8

Verizon Communications, Inc.

2.2

2.3

29.9

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Financials

30.7

31.7

Energy

15.6

14.0

Industrials

10.6

10.7

Consumer Staples

8.5

7.3

Health Care

7.6

7.6

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.7%

Stocks 99.7%

Short-Term
Investments and
Net Other Assets 0.3%

Short-Term
Investments and
Net Other Assets 0.3%

* Foreign investments

5.3%

**Foreign investments

2.0%

Annual Report

Fidelity Large Cap Value Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.7%

Shares

Value

CONSUMER DISCRETIONARY - 7.3%

Automobiles - 0.5%

General Motors Corp. (d)

283,401

$ 8,023,082

Hotels, Restaurants & Leisure - 2.0%

McDonald's Corp.

557,500

29,854,125

Household Durables - 0.3%

Whirlpool Corp.

56,000

4,766,160

Internet & Catalog Retail - 0.3%

Expedia, Inc. (a)

198,600

4,571,772

Leisure Equipment & Products - 0.4%

Eastman Kodak Co.

261,400

5,209,702

Media - 1.2%

Liberty Media Corp. - Capital
Series A (a)

64,300

6,919,966

Viacom, Inc. Class B (non-vtg.) (a)

296,500

11,492,340

18,412,306

Multiline Retail - 0.7%

Macy's, Inc.

387,800

10,718,792

Specialty Retail - 0.8%

Home Depot, Inc.

414,300

12,706,581

Textiles, Apparel & Luxury Goods - 1.1%

NIKE, Inc. Class B (d)

255,700

15,792,032

TOTAL CONSUMER DISCRETIONARY

110,054,552

CONSUMER STAPLES - 8.5%

Beverages - 1.1%

Molson Coors Brewing Co. Class B

392,400

17,528,508

Food & Staples Retailing - 2.9%

BJ's Wholesale Club, Inc. (a)

147,100

4,771,924

Kroger Co.

621,500

15,817,175

Safeway, Inc.

267,700

8,296,023

SUPERVALU, Inc.

484,600

14,567,076

43,452,198

Food Products - 0.3%

Del Monte Foods Co.

459,500

4,121,715

Household Products - 1.9%

Procter & Gamble Co.

433,300

28,576,135

Tobacco - 2.3%

Altria Group, Inc.

450,000

34,119,000

TOTAL CONSUMER STAPLES

127,797,556

ENERGY - 15.6%

Energy Equipment & Services - 1.9%

National Oilwell Varco, Inc. (a)

232,500

14,003,475

Transocean, Inc. (a)

112,200

13,755,720

27,759,195

Oil, Gas & Consumable Fuels - 13.7%

Chevron Corp.

482,300

40,754,350

ConocoPhillips

574,900

46,175,968

Shares

Value

Exxon Mobil Corp.

744,700

$ 64,342,081

Hess Corp.

265,100

24,079,033

Marathon Oil Corp.

308,900

14,471,965

Valero Energy Corp.

274,801

16,265,471

206,088,868

TOTAL ENERGY

233,848,063

FINANCIALS - 30.7%

Capital Markets - 6.5%

GLG Partners, Inc. (a)(d)

1,247,500

14,708,025

Goldman Sachs Group, Inc.

143,200

28,750,264

Janus Capital Group, Inc. (d)

575,300

15,538,853

Lehman Brothers Holdings, Inc. (d)

459,300

29,473,281

State Street Corp.

120,500

9,895,460

98,365,883

Commercial Banks - 3.8%

Associated Banc-Corp.

181,900

5,125,942

PNC Financial Services Group, Inc.

113,200

7,428,184

Regions Financial Corp. (d)

222,500

5,615,900

Wachovia Corp.

507,800

19,768,654

Wells Fargo & Co.

359,100

12,212,991

Zions Bancorp

132,200

7,236,628

57,388,299

Consumer Finance - 0.9%

Capital One Financial Corp. (d)

252,700

13,850,487

Diversified Financial Services - 6.7%

Bank of America Corp.

813,700

36,087,595

Citigroup, Inc.

546,460

15,421,101

JPMorgan Chase & Co.

947,400

45,048,870

MSCI, Inc. Class A

115,197

3,798,045

100,355,611

Insurance - 10.5%

ACE Ltd.

281,000

16,393,540

American International Group, Inc.

742,300

40,945,268

Assurant, Inc.

142,600

9,253,314

Axis Capital Holdings Ltd.

226,400

9,065,056

Endurance Specialty Holdings Ltd.

252,800

10,243,456

Loews Corp.

357,400

16,687,006

PartnerRe Ltd.

157,400

12,478,672

Prudential Financial, Inc.

255,600

21,564,972

RenaissanceRe Holdings Ltd.

215,900

12,304,141

W.R. Berkley Corp.

284,000

8,593,840

157,529,265

Real Estate Investment Trusts - 2.0%

Annaly Capital Management, Inc.

716,400

14,127,408

Boston Properties, Inc.

77,000

7,077,840

ProLogis Trust

91,600

5,436,460

Public Storage

46,700

3,654,275

30,295,983

Common Stocks - continued

Shares

Value

FINANCIALS - continued

Thrifts & Mortgage Finance - 0.3%

Astoria Financial Corp.

167,400

$ 4,549,932

TOTAL FINANCIALS

462,335,460

HEALTH CARE - 7.6%

Biotechnology - 1.0%

Biogen Idec, Inc. (a)

239,600

14,603,620

Health Care Equipment & Supplies - 0.5%

Baxter International, Inc.

118,200

7,179,468

Health Care Providers & Services - 1.8%

Humana, Inc. (a)

169,900

13,642,970

Medco Health Solutions, Inc. (a)

275,400

13,792,032

27,435,002

Life Sciences Tools & Services - 1.4%

Charles River Laboratories International, Inc. (a)

160,000

9,936,000

Invitrogen Corp. (a)

127,800

10,948,626

20,884,626

Pharmaceuticals - 2.9%

Johnson & Johnson

349,000

22,077,740

Pfizer, Inc.

960,700

22,470,773

44,548,513

TOTAL HEALTH CARE

114,651,229

INDUSTRIALS - 10.6%

Aerospace & Defense - 3.2%

Honeywell International, Inc.

94,300

5,570,301

Northrop Grumman Corp.

165,400

13,126,144

Raytheon Co.

170,600

11,112,884

The Boeing Co.

212,800

17,700,704

47,510,033

Airlines - 0.5%

Northwest Airlines Corp. (a)

193,200

3,614,772

UAL Corp.

121,400

4,607,130

8,221,902

Commercial Services & Supplies - 0.6%

Allied Waste Industries, Inc. (a)

920,000

9,062,000

Construction & Engineering - 1.4%

KBR, Inc. (a)

488,600

15,434,874

URS Corp. (a)

133,200

5,847,480

21,282,354

Industrial Conglomerates - 3.4%

General Electric Co.

1,424,000

50,423,840

Machinery - 1.2%

AGCO Corp. (a)

127,000

7,647,940

Cummins, Inc.

220,900

10,665,052

18,312,992

Shares

Value

Road & Rail - 0.3%

Hertz Global Holdings, Inc. (a)

266,933

$ 3,982,640

TOTAL INDUSTRIALS

158,795,761

INFORMATION TECHNOLOGY - 3.1%

Computers & Peripherals - 1.6%

Hewlett-Packard Co.

147,600

6,457,500

International Business Machines Corp.

124,200

13,331,628

Western Digital Corp. (a)

176,300

4,663,135

24,452,263

Electronic Equipment & Instruments - 0.8%

Tyco Electronics Ltd.

337,100

11,397,351

IT Services - 0.5%

Accenture Ltd. Class A

215,500

7,460,610

Software - 0.2%

Oracle Corp. (a)

172,000

3,534,600

TOTAL INFORMATION TECHNOLOGY

46,844,824

MATERIALS - 4.1%

Chemicals - 2.2%

Celanese Corp. Class A

164,400

6,112,392

CF Industries Holdings, Inc.

76,500

8,180,145

Chemtura Corp.

1,255,300

8,410,510

The Mosaic Co. (a)

108,600

9,883,686

32,586,733

Containers & Packaging - 0.6%

Ball Corp.

91,600

4,203,524

Owens-Illinois, Inc. (a)

94,300

4,752,720

8,956,244

Metals & Mining - 1.3%

Freeport-McMoRan Copper & Gold, Inc. Class B

104,800

9,330,344

United States Steel Corp.

99,300

10,139,523

19,469,867

TOTAL MATERIALS

61,012,844

TELECOMMUNICATION SERVICES - 6.0%

Diversified Telecommunication Services - 6.0%

AT&T, Inc.

1,480,362

56,979,133

Verizon Communications, Inc.

842,100

32,707,164

89,686,297

UTILITIES - 6.2%

Electric Utilities - 1.7%

Entergy Corp.

112,000

12,116,160

PPL Corp.

179,000

8,756,680

Reliant Energy, Inc. (a)

255,100

5,425,977

26,298,817

Gas Utilities - 1.0%

Energen Corp.

244,700

15,391,630

Common Stocks - continued

Shares

Value

UTILITIES - continued

Independent Power Producers & Energy Traders - 2.4%

Constellation Energy Group, Inc.

92,200

$ 8,663,112

Mirant Corp. (a)

324,700

11,961,948

NRG Energy, Inc. (a)

382,800

14,772,252

35,397,312

Multi-Utilities - 1.1%

Public Service Enterprise Group, Inc.

170,100

16,329,600

TOTAL UTILITIES

93,417,359

TOTAL COMMON STOCKS

(Cost $1,476,386,384)

1,498,443,945

Money Market Funds - 5.2%

Fidelity Cash Central Fund, 3.79% (b)

9,464,505

9,464,505

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

69,571,507

69,571,507

TOTAL MONEY MARKET FUNDS

(Cost $79,036,012)

79,036,012

TOTAL INVESTMENT PORTFOLIO - 104.9%

(Cost $1,555,422,396)

1,577,479,957

NET OTHER ASSETS - (4.9)%

(74,027,771)

NET ASSETS - 100%

$ 1,503,452,186

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 625,964

Fidelity Securities Lending Cash Central Fund

89,576

Total

$ 715,540

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $42,711,239 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Value Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $69,644,477) - See accompanying schedule:

Unaffiliated issuers (cost $1,476,386,384)

$ 1,498,443,945

Fidelity Central Funds (cost $79,036,012)

79,036,012

Total Investments (cost $1,555,422,396)

$ 1,577,479,957

Receivable for investments sold

23,766,119

Receivable for fund shares sold

2,565,058

Dividends receivable

1,696,869

Distributions receivable from Fidelity Central Funds

33,811

Prepaid expenses

4,921

Total assets

1,605,546,735

Liabilities

Payable for investments purchased

$ 23,494,925

Payable for fund shares redeemed

7,847,405

Accrued management fee

720,873

Distribution fees payable

6,898

Other affiliated payables

363,497

Other payables and accrued expenses

89,444

Collateral on securities loaned, at value

69,571,507

Total liabilities

102,094,549

Net Assets

$ 1,503,452,186

Net Assets consist of:

Paid in capital

$ 1,523,198,617

Undistributed net investment income

4,864,259

Accumulated undistributed net realized gain (loss) on investments

(46,668,251)

Net unrealized appreciation (depreciation) on investments

22,057,561

Net Assets

$ 1,503,452,186

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering
Price

Class A:
Net Asset Value
and redemption price per share ($9,774,232 ÷ 721,934 shares)

$ 13.54

Maximum offering price per share (100/94.25 of $13.54)

$ 14.37

Class T:
Net Asset Value
and redemption price per share ($5,975,654 ÷ 441,693 shares)

$ 13.53

Maximum offering price per share (100/96.50 of $13.53)

$ 14.02

Class B:
Net Asset Value
and offering price per share ($1,859,809 ÷ 137,404 shares)A

$ 13.54

Class C:
Net Asset Value
and offering price per share ($1,208,181 ÷ 89,368 shares)A

$ 13.52

Large Cap Value:
Net Asset Value
, offering price and redemption price per share ($1,483,574,281 ÷ 109,324,284 shares)

$ 13.57

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($1,060,029 ÷ 78,261 shares)

$ 13.54

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Value Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 32,039,820

Interest

4,832

Income from Fidelity Central Funds (including $89,576 from security lending)

715,540

Total income

32,760,192

Expenses

Management fee
Basic fee

$ 9,024,987

Performance adjustment

288,595

Transfer agent fees

3,768,695

Distribution fees

73,728

Accounting and security lending fees

507,630

Custodian fees and expenses

37,007

Independent trustees' compensation

5,753

Registration fees

122,860

Audit

60,620

Legal

7,461

Interest

5,226

Miscellaneous

53,563

Total expenses before reductions

13,956,125

Expense reductions

(136,896)

13,819,229

Net investment income (loss)

18,940,963

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

18,026,031

Change in net unrealized appreciation (depreciation) on investment securities

(121,407,791)

Net gain (loss)

(103,381,760)

Net increase (decrease) in net assets resulting from operations

$ (84,440,797)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ 18,940,963

$ 9,973,926

Net realized gain (loss)

18,026,031

34,353,479

Change in net unrealized appreciation (depreciation)

(121,407,791)

98,674,962

Net increase (decrease) in net assets resulting from operations

(84,440,797)

143,002,367

Distributions to shareholders from net investment income

(14,286,599)

(10,401,399)

Distributions to shareholders from net realized gain

(90,733,678)

(14,021,724)

Total distributions

(105,020,277)

(24,423,123)

Share transactions - net increase (decrease)

320,162,513

685,040,997

Redemption fees

-

21,396

Total increase (decrease) in net assets

130,701,439

803,641,637

Net Assets

Beginning of period

1,372,750,747

569,109,110

End of period (including undistributed net investment income of $4,864,259 and undistributed net investment income
of $274,346, respectively)

$ 1,503,452,186

$ 1,372,750,747

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.12

Net realized and unrealized gain (loss)

(1.00)

Total from investment operations

(.88)

Distributions from net investment income

(.12)

Distributions from net realized gain

(.87)

Total distributions

(.99)

Net asset value, end of period

$ 13.54

Total Return B,C

(6.04)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.22% A

Expenses net of fee waivers, if any

1.22% A

Expenses net of all reductions

1.22% A

Net investment income (loss)

.81% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 9,774

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the sales charges.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.08

Net realized and unrealized gain (loss)

(1.01)

Total from investment operations

(.93)

Distributions from net investment income

(.08)

Distributions from net realized gain

(.87)

Total distributions

(.95)

Net asset value, end of period

$ 13.53

Total Return B,C

(6.34)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.47% A

Expenses net of fee waivers, if any

1.47% A

Expenses net of all reductions

1.47% A

Net investment income (loss)

.56% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 5,976

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the sales charges.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.01

Net realized and unrealized gain (loss)

(1.00)

Total from investment operations

(.99)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.87)

Total distributions

(.88)

Net asset value, end of period

$ 13.54

Total Return B,C

(6.74)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.99% A

Expenses net of fee waivers, if any

1.99% A

Expenses net of all reductions

1.99% A

Net investment income (loss)

.04% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,860

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the contingent deferred sales charge.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.01

Net realized and unrealized gain (loss)

(.98)

Total from investment operations

(.97)

Distributions from net investment income

(.05)

Distributions from net realized gain

(.87)

Total distributions

(.92)

Net asset value, end of period

$ 13.52

Total Return B,C

(6.61)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.94% A

Expenses net of fee waivers, if any

1.94% A

Expenses net of all reductions

1.94% A

Net investment income (loss)

.09% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,208

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the contingent deferred sales charge.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Large Cap Value

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 15.19

$ 13.62

$ 12.04

$ 10.64

$ 8.17

Income from Investment Operations

Net investment income (loss) B

.18

.16

.17

.09 E

.09

Net realized and unrealized gain (loss)

(.80)

1.80

1.87

1.47

2.47

Total from investment operations

(.62)

1.96

2.04

1.56

2.56

Distributions from net investment income

(.13)

(.13)

(.11)

(.05)

(.09)

Distributions from net realized gain

(.87)

(.26)

(.35)

(.11)

-

Total distributions

(1.00)

(.39)

(.46)

(.16)

(.09)

Redemption fees added to paid in capital B

-

- G,H

- G

- G

- G

Net asset value, end of period

$ 13.57

$ 15.19

$ 13.62

$ 12.04

$ 10.64

Total Return A

(4.39)%

14.63%

17.09%

14.68%

31.44%

Ratios to Average Net Assets C,F

Expenses before reductions

.86%

.89%

.89%

1.07%

1.45%

Expenses net of fee waivers, if any

.85%

.89%

.89%

1.07%

1.20%

Expenses net of all reductions

.85%

.89%

.84%

1.05%

1.18%

Net investment income (loss)

1.18%

1.10%

1.32%

.79% E

.99%

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,483,574

$ 1,372,751

$ 569,109

$ 177,004

$ 25,168

Portfolio turnover rate D

204%

164%

175%

170%

72%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been .59%.

F Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

G Amount represents less than $.01 per share.

H The redemption fee was eliminated during the year January 31, 2007.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.17

Net realized and unrealized gain (loss)

(1.02)

Total from investment operations

(.85)

Distributions from net investment income

(.15)

Distributions from net realized gain

(.87)

Total distributions

(1.02)

Net asset value, end of period

$ 13.54

Total Return B,C

(5.82)%

Ratios to Average Net Assets E,H

Expenses before reductions

.85% A

Expenses net of fee waivers, if any

.85% A

Expenses net of all reductions

.84% A

Net investment income (loss)

1.19% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,060

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Advisor Mid Cap Value Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the class' dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table does not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Class A (incl. 5.75% sales charge) B

-13.22%

14.21%

9.28%

Class T (incl. 3.50% sales charge) C

-11.33%

14.70%

9.66%

Class B (incl. contingent deferred sales charge) D

-12.91%

15.19%

10.21%

Class C (incl. contingent deferred sales charge) E

-9.48%

15.40%

10.20%

A From November 15, 2001.

B Class A shares bear a 0.25% 12b-1 fee. The initial offering of Class A shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Value, the original class of the fund, which has no 12b-1 fee. Had Class A shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower.

C Class T shares bear a 0.50% 12b-1 fee. The initial offering of Class T shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Value, the original class of the fund, which has no 12b-1 fee. Had Class T shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower.

D Class B shares bear a 1.00% 12b-1 fee. The initial offering of Class B shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Value, the original class of the fund, which has no 12b-1 fee. Had Class B shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower. Class B shares' contingent deferred sales charge included in the past one year, past five years, and life of fund total return figures are 5%, 2%, and 0%, respectively.

E Class C shares bear a 1.00% 12b-1 fee. The initial offering of Class C shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Value, the original class of the fund, which has no 12b-1 fee. Had Class C shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower. Class C shares' contingent deferred sales charge included in the past one year, past five years, and life of fund total return figures are 1%, 0%, and 0%, respectively.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Fidelity Advisor Mid Cap Value Fund - Class A on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell Midcap® Value Index performed over the same period. The initial offering of Class A took place on February 13, 2007. See above for additional information regarding performance of Class A.



Annual Report

Fidelity Advisor Mid Cap Value Fund

Management's Discussion of Fund Performance

Comments from Bruce Dirks, Portfolio Manager of Fidelity Advisor Mid Cap Value Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the year, Advisor Mid Cap Value's Class A, Class T, Class B and Class C shares (excluding sales charges) all finished slightly ahead of the benchmark Russell Midcap® Value Index, which declined 8.73%. (For specific class-level results, please refer to the performance section of this report.) During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among bank and asset management companies. Favorable industry weightings within the weak financials and consumer discretionary sectors also added value. Good stock picking in materials, energy and the equipment/services group of health care also helped. Among the fund's strongest contributors for the period were CF Industries, an out-of-index company that produces fertilizer products; Cummins, a diesel engine manufacturer; Annaly Capital Management, a real estate investment trust that owned AAA-rated mortgages; and National Oilwell Varco, an energy services contractor. Conversely, relative results were held back by some unfavorable stock selection decisions in the utilities, consumer staples and consumer discretionary sectors. The fund's poorest performers included transportation firm YRC Worldwide; original equipment manufacturer TRW Automotive; and photographic imaging company Eastman Kodak. YRC and TRW were no longer held at period end.

For the year, Advisor Mid Cap Value Fund's Institutional Class shares finished slightly ahead of the benchmark Russell Midcap® Value Index, which declined 8.73%. (For specific class-level results, please refer to the performance section of this report.) During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among bank and asset management companies. Favorable industry weightings within the weak financials and consumer discretionary sectors also added value. Good stock picking in materials, energy and the equipment/services group of health care also helped. Among the fund's strongest contributors for the period were CF Industries, an out-of-index company that produces fertilizer products; Cummins, a diesel engine manufacturer; Annaly Capital Management, a real estate investment trust that owned AAA-rated mortgages; and National Oilwell Varco, an energy services contractor. Conversely, relative results were held back by some unfavorable stock selection decisions in the utilities, consumer staples and consumer discretionary sectors. The fund's poorest performers included transportation firm YRC Worldwide; original equipment manufacturer TRW Automotive; and photographic imaging company Eastman Kodak. YRC and TRW were no longer held at period end.

The views expressed above reflect those of the portfolio manager only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Mid Cap Value Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Hess Corp.

2.5

2.3

Edison International

2.1

1.5

Everest Re Group Ltd.

1.8

0.0

Annaly Capital Management, Inc.

1.7

0.8

Boston Properties, Inc.

1.7

0.0

Questar Corp.

1.7

0.0

Axis Capital Holdings Ltd.

1.6

0.8

Assurant, Inc.

1.6

1.2

PartnerRe Ltd.

1.6

1.2

Energen Corp.

1.6

1.8

17.9

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Financials

30.0

29.5

Utilities

14.0

12.5

Consumer Discretionary

13.1

13.1

Industrials

10.4

10.7

Energy

8.1

7.0

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.7%

Stocks 99.4%

Short-Term
Investments and
Net Other Assets 0.3%

Short-Term
Investments and
Net Other Assets 0.6%

* Foreign investments

13.1%

**Foreign investments

4.2%

Annual Report

Fidelity Mid Cap Value Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.7%

Shares

Value

CONSUMER DISCRETIONARY - 13.1%

Diversified Consumer Services - 0.5%

Apollo Group, Inc. Class A (non-vtg.) (a)

50,800

$ 4,050,792

Hotels, Restaurants & Leisure - 1.1%

Burger King Holdings, Inc.

303,600

7,999,860

Household Durables - 2.3%

Leggett & Platt, Inc. (d)

296,500

5,639,430

Whirlpool Corp. (d)

133,500

11,362,185

17,001,615

Internet & Catalog Retail - 1.2%

Expedia, Inc. (a)

242,600

5,584,652

Liberty Media Corp. - Interactive
Series A (a)

239,500

3,810,445

9,395,097

Leisure Equipment & Products - 1.4%

Eastman Kodak Co. (d)

534,300

10,648,599

Media - 2.1%

E.W. Scripps Co. Class A

181,100

7,374,392

Liberty Media Corp. - Capital
Series A (a)

74,900

8,060,738

15,435,130

Specialty Retail - 2.6%

Abercrombie & Fitch Co. Class A

96,200

7,666,178

AutoZone, Inc. (a)

48,200

5,826,416

RadioShack Corp.

136,800

2,373,480

TJX Companies, Inc.

125,600

3,963,936

19,830,010

Textiles, Apparel & Luxury Goods - 1.9%

Crocs, Inc. (a)(d)

219,600

7,639,884

Jones Apparel Group, Inc. (d)

399,000

6,703,200

14,343,084

TOTAL CONSUMER DISCRETIONARY

98,704,187

CONSUMER STAPLES - 7.1%

Beverages - 1.9%

Coca-Cola Enterprises, Inc.

311,600

7,188,612

Molson Coors Brewing Co. Class B

168,900

7,544,763

14,733,375

Food & Staples Retailing - 2.8%

BJ's Wholesale Club, Inc. (a)(d)

170,600

5,534,264

Safeway, Inc.

259,000

8,026,410

SUPERVALU, Inc.

239,600

7,202,376

20,763,050

Food Products - 2.2%

Corn Products International, Inc.

136,600

4,617,080

Dean Foods Co.

158,400

4,435,200

Del Monte Foods Co.

316,300

2,837,211

Tyson Foods, Inc. Class A

308,000

4,389,000

16,278,491

Shares

Value

Personal Products - 0.2%

Estee Lauder Companies, Inc. Class A

44,100

$ 1,861,020

TOTAL CONSUMER STAPLES

53,635,936

ENERGY - 8.1%

Energy Equipment & Services - 4.8%

Helmerich & Payne, Inc.

206,500

8,098,930

Nabors Industries Ltd. (a)

152,100

4,140,162

National Oilwell Varco, Inc. (a)

121,000

7,287,830

Patterson-UTI Energy, Inc. (d)

450,800

8,826,664

Superior Energy Services, Inc. (a)

74,400

2,982,696

Tidewater, Inc.

81,800

4,332,128

35,668,410

Oil, Gas & Consumable Fuels - 3.3%

Frontline Ltd. (NY Shares)

137,300

5,844,861

Hess Corp. (d)

211,800

19,237,794

25,082,655

TOTAL ENERGY

60,751,065

FINANCIALS - 30.0%

Capital Markets - 3.0%

GLG Partners, Inc. (a)(d)

756,200

8,915,598

Janus Capital Group, Inc. (d)

352,500

9,521,025

MF Global Ltd.

151,200

4,543,560

22,980,183

Commercial Banks - 3.5%

Associated Banc-Corp.

277,700

7,825,586

BOK Financial Corp.

71,195

3,878,704

East West Bancorp, Inc. (d)

188,900

4,544,934

Synovus Financial Corp.

219,900

2,904,879

Zions Bancorp

125,800

6,886,292

26,040,395

Diversified Financial Services - 0.6%

MSCI, Inc. Class A

136,800

4,510,296

Insurance - 13.9%

ACE Ltd.

158,000

9,217,720

Allied World Assurance Co. Holdings Ltd.

147,800

7,039,714

Assurant, Inc.

187,200

12,147,408

Axis Capital Holdings Ltd.

308,900

12,368,356

CNA Financial Corp.

189,800

6,451,302

Endurance Specialty Holdings Ltd.

286,800

11,621,136

Everest Re Group Ltd.

133,100

13,534,939

PartnerRe Ltd.

152,100

12,058,488

RenaissanceRe Holdings Ltd.

197,800

11,272,622

W.R. Berkley Corp.

303,200

9,174,832

104,886,517

Real Estate Investment Trusts - 7.0%

Annaly Capital Management, Inc.

658,200

12,979,704

Boston Properties, Inc.

138,600

12,740,112

Common Stocks - continued

Shares

Value

FINANCIALS - continued

Real Estate Investment Trusts - continued

Colonial Properties Trust (SBI)

97,800

$ 2,409,792

Kimco Realty Corp.

198,300

7,101,123

ProLogis Trust

155,700

9,240,795

Public Storage

73,700

5,767,025

Taubman Centers, Inc.

47,400

2,377,110

52,615,661

Thrifts & Mortgage Finance - 2.0%

Astoria Financial Corp.

262,500

7,134,750

Washington Federal, Inc.

323,700

7,904,754

15,039,504

TOTAL FINANCIALS

226,072,556

HEALTH CARE - 2.3%

Health Care Equipment & Supplies - 0.3%

Kinetic Concepts, Inc. (a)(d)

41,100

2,045,958

Health Care Providers & Services - 0.5%

Humana, Inc. (a)

42,500

3,412,750

Life Sciences Tools & Services - 1.2%

Charles River Laboratories International, Inc. (a)

74,200

4,607,820

Invitrogen Corp. (a)

54,600

4,677,582

9,285,402

Pharmaceuticals - 0.3%

Watson Pharmaceuticals, Inc. (a)

85,000

2,219,350

TOTAL HEALTH CARE

16,963,460

INDUSTRIALS - 10.4%

Aerospace & Defense - 1.7%

DRS Technologies, Inc.

88,400

4,744,428

L-3 Communications Holdings, Inc.

72,500

8,035,175

12,779,603

Airlines - 0.8%

Northwest Airlines Corp. (a)

152,600

2,855,146

UAL Corp.

75,800

2,876,610

5,731,756

Building Products - 0.6%

Lennox International, Inc.

126,800

4,711,888

Commercial Services & Supplies - 2.2%

Allied Waste Industries, Inc. (a)

700,300

6,897,955

R.R. Donnelley & Sons Co.

267,200

9,322,608

16,220,563

Construction & Engineering - 1.7%

KBR, Inc. (a)

309,000

9,761,310

URS Corp. (a)

69,800

3,064,220

12,825,530

Shares

Value

Industrial Conglomerates - 0.6%

McDermott International, Inc. (a)

94,500

$ 4,458,510

Machinery - 2.5%

AGCO Corp. (a)

106,000

6,383,320

Cummins, Inc.

117,300

5,663,244

Ingersoll-Rand Co. Ltd. Class A

168,400

6,655,168

18,701,732

Road & Rail - 0.3%

Hertz Global Holdings, Inc. (a)

160,200

2,390,184

TOTAL INDUSTRIALS

77,819,766

INFORMATION TECHNOLOGY - 6.4%

Communications Equipment - 0.7%

Juniper Networks, Inc. (a)

195,900

5,318,685

Computers & Peripherals - 2.2%

NCR Corp. (a)

422,600

9,077,448

Western Digital Corp. (a)

278,300

7,361,035

16,438,483

Electronic Equipment & Instruments - 1.4%

Avnet, Inc. (a)

152,500

5,430,525

Ingram Micro, Inc. Class A (a)

173,400

3,083,052

Jabil Circuit, Inc.

174,200

2,308,150

10,821,727

Semiconductors & Semiconductor Equipment - 1.6%

Atmel Corp. (a)

1,316,800

4,161,088

MEMC Electronic Materials, Inc. (a)

25,000

1,786,500

Novellus Systems, Inc. (a)

111,700

2,653,992

Varian Semiconductor Equipment Associates, Inc. (a)

109,100

3,514,111

12,115,691

Software - 0.5%

Compuware Corp. (a)

405,900

3,450,150

TOTAL INFORMATION TECHNOLOGY

48,144,736

MATERIALS - 6.8%

Chemicals - 3.1%

Celanese Corp. Class A

146,800

5,458,024

CF Industries Holdings, Inc.

57,100

6,105,703

Chemtura Corp. (d)

867,400

5,811,580

The Mosaic Co. (a)

71,200

6,479,912

23,855,219

Containers & Packaging - 1.9%

Ball Corp.

85,100

3,905,239

Owens-Illinois, Inc. (a)

91,500

4,611,600

Smurfit-Stone Container Corp. (a)(d)

603,500

5,727,215

14,244,054

Common Stocks - continued

Shares

Value

MATERIALS - continued

Metals & Mining - 1.8%

Cleveland-Cliffs, Inc. (d)

42,700

$ 4,348,568

United States Steel Corp.

88,000

8,985,680

13,334,248

TOTAL MATERIALS

51,433,521

TELECOMMUNICATION SERVICES - 1.5%

Diversified Telecommunication Services - 1.5%

Embarq Corp.

116,500

5,277,450

Qwest Communications International, Inc.

1,043,800

6,137,544

11,414,994

UTILITIES - 14.0%

Electric Utilities - 4.6%

Allegheny Energy, Inc.

123,300

6,755,607

Edison International

305,600

15,940,096

Reliant Energy, Inc. (a)

558,600

11,881,422

34,577,125

Gas Utilities - 3.3%

Energen Corp.

191,700

12,057,930

Questar Corp.

247,300

12,590,043

24,647,973

Independent Power Producers & Energy Traders - 4.2%

Constellation Energy Group, Inc.

103,700

9,743,652

Mirant Corp. (a)

315,100

11,608,284

NRG Energy, Inc. (a)

268,200

10,349,838

31,701,774

Multi-Utilities - 1.9%

CMS Energy Corp.

583,000

9,135,610

OGE Energy Corp.

162,500

5,318,625

14,454,235

TOTAL UTILITIES

105,381,107

TOTAL COMMON STOCKS

(Cost $760,656,463)

750,321,328

Money Market Funds - 10.6%

Shares

Value

Fidelity Cash Central Fund, 3.79% (b)

5,496,422

$ 5,496,422

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

74,160,825

74,160,825

TOTAL MONEY MARKET FUNDS

(Cost $79,657,247)

79,657,247

TOTAL INVESTMENT PORTFOLIO - 110.3%

(Cost $840,313,710)

829,978,575

NET OTHER ASSETS - (10.3)%

(77,171,615)

NET ASSETS - 100%

$ 752,806,960

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 394,997

Fidelity Securities Lending Cash Central Fund

169,409

Total

$ 564,406

Other Information

Distribution of investments by country of issue, as a percentage of total net assets, is as follows:

United States of America

86.9%

Bermuda

11.3%

Cayman Islands

1.2%

Others (individually less than 1%)

0.6%

100.0%

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $31,051,104 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Value Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $73,863,006) - See accompanying schedule:

Unaffiliated issuers (cost $760,656,463)

$ 750,321,328

Fidelity Central Funds (cost $79,657,247)

79,657,247

Total Investments (cost $840,313,710)

$ 829,978,575

Receivable for investments sold

20,806,082

Receivable for fund shares sold

2,652,227

Dividends receivable

223,102

Distributions receivable from Fidelity Central Funds

36,747

Prepaid expenses

2,661

Other receivables

46

Total assets

853,699,440

Liabilities

Payable for investments purchased

$ 20,180,467

Payable for fund shares redeemed

5,956,855

Accrued management fee

312,903

Distribution fees payable

5,417

Other affiliated payables

203,019

Other payables and accrued expenses

72,994

Collateral on securities loaned, at value

74,160,825

Total liabilities

100,892,480

Net Assets

$ 752,806,960

Net Assets consist of:

Paid in capital

$ 788,210,259

Undistributed net investment income

1,465,714

Accumulated undistributed net realized gain (loss) on investments

(26,533,878)

Net unrealized appreciation (depreciation) on investments

(10,335,135)

Net Assets

$ 752,806,960

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($7,444,673 ÷ 494,661 shares)

$ 15.05

Maximum offering price per share (100/94.25 of $15.05)

$ 15.97

Class T:
Net Asset Value
and redemption price per share ($3,714,323 ÷ 247,029 shares)

$ 15.04

Maximum offering price per share (100/96.50 of $15.04)

$ 15.59

Class B:
Net Asset Value
and offering price per share ($1,304,436 ÷ 87,007 shares)A

$ 14.99

Class C:
Net Asset Value
and offering price per share ($1,657,889 ÷ 110,680 shares)A

$ 14.98

Mid Cap Value:
Net Asset Value
, offering price and redemption price per share ($737,234,029 ÷ 48,847,921 shares)

$ 15.09

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($1,451,610 ÷ 96,411 shares)

$ 15.06

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Value Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 11,189,048

Interest

8,457

Income from Fidelity Central Funds (including $169,409 from security lending)

564,406

Total income

11,761,911

Expenses

Management fee
Basic fee

$ 5,084,079

Performance adjustment

(317,451)

Transfer agent fees

2,236,010

Distribution fees

53,691

Accounting and security lending fees

315,091

Custodian fees and expenses

28,074

Independent trustees' compensation

3,227

Registration fees

145,678

Audit

58,698

Legal

4,195

Interest

4,946

Miscellaneous

29,800

Total expenses before reductions

7,646,038

Expense reductions

(105,134)

7,540,904

Net investment income (loss)

4,221,007

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

(3,216,421)

Change in net unrealized appreciation (depreciation) on investment securities

(85,150,820)

Net gain (loss)

(88,367,241)

Net increase (decrease) in net assets resulting from operations

$ (84,146,234)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ 4,221,007

$ 2,688,685

Net realized gain (loss)

(3,216,421)

24,851,516

Change in net unrealized appreciation (depreciation)

(85,150,820)

43,131,851

Net increase (decrease) in net assets resulting from operations

(84,146,234)

70,672,052

Distributions to shareholders from net investment income

(3,037,015)

(2,949,768)

Distributions to shareholders from net realized gain

(37,140,454)

(13,997,322)

Total distributions

(40,177,469)

(16,947,090)

Share transactions - net increase (decrease)

198,288,449

259,218,977

Redemption fees

47,852

33,264

Total increase (decrease) in net assets

74,012,598

312,977,203

Net Assets

Beginning of period

678,794,362

365,817,159

End of period (including undistributed net investment income of $1,465,714 and undistributed net investment income
of $375,291, respectively)

$ 752,806,960

$ 678,794,362

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

.03

Net realized and unrealized gain (loss)

(1.78)

Total from investment operations

(1.75)

Distributions from net investment income

(.06)

Distributions from net realized gain

(.77)

Total distributions

(.83)

Net asset value, end of period

$ 15.05

Total Return B,C,D

(10.28)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.14% A

Expenses net of fee waivers, if any

1.14% A

Expenses net of all reductions

1.13% A

Net investment income (loss)

.16% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 7,445

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.02)

Net realized and unrealized gain (loss)

(1.76)

Total from investment operations

(1.78)

Distributions from net investment income

(.04)

Distributions from net realized gain

(.77)

Total distributions

(.81)

Net asset value, end of period

$ 15.04

Total Return B,C,D

(10.46)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.39% A

Expenses net of fee waivers, if any

1.39% A

Expenses net of all reductions

1.39% A

Net investment income (loss)

(.10)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 3,714

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.10)

Net realized and unrealized gain (loss)

(1.76)

Total from investment operations

(1.86)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.77)

Total distributions

(.78)

Net asset value, end of period

$ 14.99

Total Return B,C,D

(10.88)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.89% A

Expenses net of fee waivers, if any

1.89% A

Expenses net of all reductions

1.89% A

Net investment income (loss)

(.59)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,304

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.10)

Net realized and unrealized gain (loss)

(1.77)

Total from investment operations

(1.87)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.77)

Total distributions

(.78)

Net asset value, end of period

$ 14.98

Total Return B,C,D

(10.94)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.90% A

Expenses net of fee waivers, if any

1.90% A

Expenses net of all reductions

1.90% A

Net investment income (loss)

(.60)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,658

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Mid Cap Value

Years ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 17.18

$ 15.65

$ 14.14

$ 12.32

$ 8.85

Income from Investment Operations

Net investment income (loss) B

.08

.09

.16 E

.08

.06

Net realized and unrealized gain (loss)

(1.34)

1.98

2.59

2.10

3.45

Total from investment operations

(1.26)

2.07

2.75

2.18

3.51

Distributions from net investment income

(.06)

(.09)

(.10)

(.04)

(.04)

Distributions from net realized gain

(.77)

(.45)

(1.15)

(.32)

-

Total distributions

(.83)

(.54)

(1.24) H

(.36)

(.04)

Redemption fees added to paid in capital B,G

-

-

-

-

-

Net asset value, end of period

$ 15.09

$ 17.18

$ 15.65

$ 14.14

$ 12.32

Total Return A

(7.67)%

13.48%

19.97%

17.75%

39.69%

Ratios to Average Net Assets C,F

Expenses before reductions

.83%

.84%

.86%

.91%

1.07%

Expenses net of fee waivers, if any

.82%

.84%

.86%

.91%

1.07%

Expenses net of all reductions

.82%

.84%

.81%

.90%

1.05%

Net investment income (loss)

.47%

.56%

1.08% E

.59%

.55%

Supplemental Data

Net assets, end of period (000 omitted)

$ 737,234

$ 678,794

$ 365,817

$ 153,231

$ 95,797

Portfolio turnover rate D

264%

187%

207%

196%

97%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.04 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been .81%.

F Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

G Amount represents less than $.01 per share.

H Total distributions of $1.24 per share is comprised of distributions from net investment income of $0.95 and distributions from net realized gain of $1.145 per share.

Financial Highlights - Institutional Class

Years ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) D

.07

Net realized and unrealized gain (loss)

(1.78)

Total from investment operations

(1.71)

Distributions from net investment income

(.09)

Distributions from net realized gain

(.77)

Total distributions

(.86)

Net asset value, end of period

$ 15.06

Total Return B,C

(10.06)%

Ratios to Average Net Assets E,H

Expenses before reductions

.89% A

Expenses net of fee waivers, if any

.89% A

Expenses net of all reductions

.88% A

Net investment income (loss)

.41% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,452

Portfolio turnover rate F

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Advisor Large Cap Growth Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the class' dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table does not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Class A (incl. 5.75% sales charge) B

-12.85%

9.39%

1.33%

Class T (incl. 3.50% sales charge) C

-10.83%

9.89%

1.70%

Class B (incl. contingent deferred sales charge) D

-12.29%

10.27%

2.18%

Class C (incl. contingent deferred sales charge) E

-8.91%

10.56%

2.20%

A From November 15, 2001.

B Class A shares bear a 0.25% 12b-1 fee. The initial offering of Class A shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Growth, the original class of the fund, which has no 12b-1 fee. Had Class A shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower.

C Class T shares bear a 0.50% 12b-1 fee. The initial offering of Class T shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Growth, the original class of the fund, which has no 12b-1 fee. Had Class T shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower.

D Class B shares bear a 1.00% 12b-1 fee. The initial offering of Class B shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Growth, the original class of the fund, which has no 12b-1 fee. Had Class B shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower. Class B shares' contingent deferred sales charge included in the past one year, past five years, and life of fund total return figures are 5%, 2%, and 0%, respectively.

E Class C shares bear a 1.00% 12b-1 fee. The initial offering of Class C shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Growth, the original class of the fund, which has no 12b-1 fee. Had Class C shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower. Class C shares' contingent deferred sales charge included in the past one year, past five years, and life of fund total return figures are 1%, 0%, and 0%, respectively.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Fidelity Advisor Large Cap Growth Fund - Class A on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell 1000® Growth Index performed over the same period. The initial offering of Class A took place on February 13, 2007. See above for additional information regarding performance of Class A.



Annual Report

Fidelity Advisor Large Cap Growth Fund

Management's Discussion of Fund Performance

Comments from Edward Best, Portfolio Manager of Fidelity Advisor Large Cap Growth Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the 12 months ending January 31, 2008, the fund's Class A, Class T, Class B and Class C shares (excluding sales charges) trailed the 0.51% gain of the Russell 1000® Growth Index. (For specific class-level results, please refer to the performance section of this report.) Poor security selection in consumer discretionary, information technology and industrials accounted for the majority of the fund's underperformance compared with the index. Within consumer discretionary, the fund was hurt most by unproductive picks within the retailing and homebuilding segments. Consumer electronics retailer RadioShack and discount retailer Big Lots fell on investors' concerns about the companies' ability to weather recessionary concerns and a weak U.S. consumer. Pharmaceuticals company Sepracor suffered due to a change in Medicare reimbursement for one of its top-selling patented drugs. Airlines US Airways and AMR Corp. - parent company of American Airlines - underperformed due to increased pressure from rising energy prices. The fund's positions in homebuilders KB Home, Ryland Group, D.R. Horton and Lennar, all of which I sold when I took over the fund in June, also held back performance. While our overall stock picking within industrials detracted, stock selection within the sector's capital goods segment actually helped the fund. Specifically, Manitowoc, a major supplier of lifting equipment to the global construction industry, aided performance. Also within capital goods, defense contractor Lockheed Martin benefited from an increase in U.S. defense spending. Stock selection among health care names was positive as well. Pharmaceutical company Biogen Idec gained on news of the re-launch of its multiple sclerosis drug, Tysabri. While stock selection in technology hurt overall, there were some contributors. Apple advanced as the stock did well on the success of its iPod, iPhone and iMac products. Software developer Microsoft also aided performance. By the end of the period, I sold US Airways, AMR Corp., RadioShack, Big Lots, Sepracor and Apple to take advantage of other opportunities in the market.

For the 12 months ending January 31, 2008, the fund's Institutional Class shares trailed the 0.51% gain of the Russell 1000® Growth Index. (For specific class-level results, please refer to the performance section of this report.) Poor security selection in consumer discretionary, information technology and industrials accounted for the majority of the fund's underperformance compared with the index. Within consumer discretionary, the fund was hurt most by unproductive picks within the retailing and homebuilding segments. Consumer electronics retailer RadioShack and discount retailer Big Lots fell on investors' concerns about the companies' ability to weather recessionary concerns and a weak U.S. consumer. Pharmaceuticals company Sepracor suffered due to a change in Medicare reimbursement for one of its top-selling patented drugs. Airlines US Airways and AMR Corp. - parent company of American Airlines - underperformed due to increased pressure from rising energy prices. The fund's positions in homebuilders KB Home, Ryland Group, D.R. Horton and Lennar, all of which I sold when I took over the fund in June, also held back performance. While our overall stock picking within industrials detracted, stock selection within the sector's capital goods segment actually helped the fund. Specifically, Manitowoc, a major supplier of lifting equipment to the global construction industry, aided performance. Also within capital goods, defense contractor Lockheed Martin benefited from an increase in U.S. defense spending. Stock selection among health care names also was positive. Pharmaceutical company Biogen Idec gained on news of the re-launch of its multiple sclerosis drug, Tysabri. While stock selection in technology hurt overall, there were some contributors. Apple advanced as the stock did well on the success of its iPod, iPhone and iMac products. Software developer Microsoft also aided performance. By the end of the period, I sold US Airways, AMR Corp., RadioShack, Big Lots, Sepracor, and Apple to take advantage of other opportunities in the market.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Large Cap Growth Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Microsoft Corp.

5.0

5.0

Exxon Mobil Corp.

4.4

1.6

Oracle Corp.

4.1

0.0

The Boeing Co.

3.9

0.0

Cisco Systems, Inc.

3.8

2.4

Bristol-Myers Squibb Co.

3.6

0.0

Accenture Ltd. Class A

3.3

3.3

NIKE, Inc. Class B

3.3

0.0

NYMEX Holdings, Inc.

3.1

0.0

Kroger Co.

3.0

3.1

37.5

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Information Technology

26.0

26.9

Health Care

16.8

16.2

Industrials

14.2

14.2

Consumer Discretionary

11.9

13.5

Consumer Staples

10.9

9.9

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.4%

Stocks 99.8%

Short-Term
Investments and
Net Other Assets 0.6%

Short-Term
Investments and
Net Other Assets 0.2%

* Foreign investments

4.4%

**Foreign
investments

5.5%

Annual Report

Fidelity Large Cap Growth Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.4%

Shares

Value

CONSUMER DISCRETIONARY - 11.9%

Auto Components - 1.6%

The Goodyear Tire & Rubber Co. (a)

99,300

$ 2,499,381

Diversified Consumer Services - 1.2%

Apollo Group, Inc. Class A (non-vtg.) (a)

22,600

1,802,124

Hotels, Restaurants & Leisure - 2.6%

McDonald's Corp.

74,200

3,973,410

Media - 2.3%

Regal Entertainment Group Class A (d)

189,300

3,509,622

Specialty Retail - 0.9%

Gamestop Corp. Class A (a)

27,400

1,417,402

Textiles, Apparel & Luxury Goods - 3.3%

NIKE, Inc. Class B

79,900

4,934,624

TOTAL CONSUMER DISCRETIONARY

18,136,563

CONSUMER STAPLES - 10.9%

Beverages - 5.0%

Pepsi Bottling Group, Inc.

114,200

3,979,870

PepsiCo, Inc.

52,500

3,579,975

7,559,845

Food & Staples Retailing - 3.0%

Kroger Co.

181,400

4,616,630

Food Products - 2.4%

Kellogg Co.

77,200

3,697,880

Household Products - 0.5%

Energizer Holdings, Inc. (a)

8,300

777,046

TOTAL CONSUMER STAPLES

16,651,401

ENERGY - 7.9%

Energy Equipment & Services - 0.9%

Baker Hughes, Inc.

5,700

370,101

Nabors Industries Ltd. (a)

24,400

664,168

Schlumberger Ltd. (NY Shares)

5,000

377,300

1,411,569

Oil, Gas & Consumable Fuels - 7.0%

Exxon Mobil Corp.

78,000

6,739,200

Sunoco, Inc.

13,000

808,600

Valero Energy Corp.

53,000

3,137,070

10,684,870

TOTAL ENERGY

12,096,439

FINANCIALS - 7.5%

Capital Markets - 4.4%

Charles Schwab Corp.

198,200

4,419,860

Northern Trust Corp.

30,900

2,266,824

6,686,684

Shares

Value

Diversified Financial Services - 3.1%

NYMEX Holdings, Inc. (d)

41,700

$ 4,795,500

TOTAL FINANCIALS

11,482,184

HEALTH CARE - 16.8%

Biotechnology - 1.2%

Biogen Idec, Inc. (a)

29,300

1,785,835

Health Care Equipment & Supplies - 2.8%

Baxter International, Inc.

69,700

4,233,578

Health Care Providers & Services - 9.2%

Aetna, Inc.

86,000

4,580,360

Express Scripts, Inc. (a)

11,400

769,386

Medco Health Solutions, Inc. (a)

35,200

1,762,816

UnitedHealth Group, Inc.

47,700

2,425,068

WellPoint, Inc. (a)

57,800

4,519,960

14,057,590

Pharmaceuticals - 3.6%

Bristol-Myers Squibb Co.

234,400

5,435,736

TOTAL HEALTH CARE

25,512,739

INDUSTRIALS - 14.2%

Aerospace & Defense - 5.6%

Lockheed Martin Corp.

5,900

636,728

Precision Castparts Corp.

16,900

1,923,220

The Boeing Co.

71,500

5,947,370

8,507,318

Commercial Services & Supplies - 3.9%

Allied Waste Industries, Inc. (a)

460,200

4,532,970

HNI Corp. (d)

15,400

518,364

Manpower, Inc.

15,300

860,778

5,912,112

Construction & Engineering - 2.7%

Fluor Corp.

7,500

912,525

Shaw Group, Inc. (a)

13,800

779,700

URS Corp. (a)

54,700

2,401,330

4,093,555

Industrial Conglomerates - 0.8%

McDermott International, Inc. (a)

24,500

1,155,910

Machinery - 1.2%

AGCO Corp. (a)

8,700

523,914

Cummins, Inc.

16,500

796,620

Manitowoc Co., Inc.

15,400

587,048

1,907,582

TOTAL INDUSTRIALS

21,576,477

INFORMATION TECHNOLOGY - 26.0%

Communications Equipment - 3.8%

Cisco Systems, Inc. (a)

236,300

5,789,350

Common Stocks - continued

Shares

Value

INFORMATION TECHNOLOGY - continued

Computers & Peripherals - 0.5%

Lexmark International, Inc. Class A (a)

1,400

$ 50,694

Western Digital Corp. (a)

29,000

767,050

817,744

Electronic Equipment & Instruments - 0.5%

National Instruments Corp.

27,400

735,964

IT Services - 6.2%

Accenture Ltd. Class A

143,600

4,971,432

Mastercard, Inc. Class A (d)

7,000

1,449,000

The Western Union Co.

135,200

3,028,480

9,448,912

Semiconductors & Semiconductor Equipment - 2.2%

MEMC Electronic Materials, Inc. (a)

12,500

893,250

NVIDIA Corp. (a)

82,500

2,028,675

Texas Instruments, Inc.

12,000

371,160

3,293,085

Software - 12.8%

Autodesk, Inc. (a)

76,100

3,131,515

BMC Software, Inc. (a)

74,200

2,377,368

Microsoft Corp.

233,200

7,602,320

Oracle Corp. (a)

306,900

6,306,795

19,417,998

TOTAL INFORMATION TECHNOLOGY

39,503,053

MATERIALS - 3.0%

Chemicals - 2.0%

Air Products & Chemicals, Inc.

4,200

378,084

E.I. du Pont de Nemours & Co.

10,100

456,318

Monsanto Co.

12,800

1,439,232

Praxair, Inc.

4,700

380,277

The Mosaic Co. (a)

4,000

364,040

3,017,951

Metals & Mining - 1.0%

Freeport-McMoRan Copper & Gold, Inc. Class B

8,400

747,852

Newmont Mining Corp.

7,100

385,814

Southern Copper Corp.

4,400

412,896

1,546,562

TOTAL MATERIALS

4,564,513

Shares

Value

TELECOMMUNICATION SERVICES - 0.3%

Wireless Telecommunication Services - 0.3%

Crown Castle International Corp. (a)

10,700

$ 387,233

UTILITIES - 0.9%

Electric Utilities - 0.9%

Exelon Corp.

18,300

1,394,277

TOTAL COMMON STOCKS

(Cost $156,064,347)

151,304,879

Money Market Funds - 6.4%

Fidelity Cash Central Fund, 3.79% (b)

1,837,290

1,837,290

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

7,887,500

7,887,500

TOTAL MONEY MARKET FUNDS

(Cost $9,724,790)

9,724,790

TOTAL INVESTMENT PORTFOLIO - 105.8%

(Cost $165,789,137)

161,029,669

NET OTHER ASSETS - (5.8)%

(8,892,495)

NET ASSETS - 100%

$ 152,137,174

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Includes investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 60,857

Fidelity Securities Lending Cash Central Fund

56,422

Total

$ 117,279

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $7,837,041 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Growth Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $7,883,516) - See accompanying schedule:

Unaffiliated issuers (cost $156,064,347)

$ 151,304,879

Fidelity Central Funds (cost $9,724,790)

9,724,790

Total Investments (cost $165,789,137)

$ 161,029,669

Receivable for investments sold

18,559,475

Receivable for fund shares sold

222,238

Dividends receivable

82,079

Distributions receivable from Fidelity Central Funds

4,743

Prepaid expenses

514

Receivable from investment adviser for expense reductions

25,900

Total assets

179,924,618

Liabilities

Payable for investments purchased

$ 19,486,946

Payable for fund shares redeemed

257,786

Accrued management fee

52,051

Distribution fees payable

1,918

Other affiliated payables

48,536

Other payables and accrued expenses

52,707

Collateral on securities loaned, at value

7,887,500

Total liabilities

27,787,444

Net Assets

$ 152,137,174

Net Assets consist of:

Paid in capital

$ 162,808,253

Accumulated undistributed net realized gain (loss) on investments

(5,911,611)

Net unrealized appreciation (depreciation) on investments

(4,759,468)

Net Assets

$ 152,137,174

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($1,302,425 ÷ 132,172 shares)

$ 9.85

Maximum offering price per share (100/94.25 of $9.85)

$ 10.45

Class T:
Net Asset Value
and redemption price per share ($1,097,491 ÷ 111,448 shares)

$ 9.85

Maximum offering price per share (100/96.50 of $9.85)

$ 10.21

Class B:
Net Asset Value
and offering price per share ($543,149 ÷ 55,237 shares)A

$ 9.83

Class C:
Net Asset Value
and offering price per share ($944,595 ÷ 96,233 shares)A

$ 9.82

Large Cap Growth:
Net Asset Value
, offering price and redemption price per share ($147,863,609 ÷ 14,947,307 shares)

$ 9.89

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($385,905 ÷ 39,068 shares)

$ 9.88

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Growth Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 1,475,584

Interest

922

Income from Fidelity Central Funds (including $56,422 from security lending)

117,279

Total income

1,593,785

Expenses

Management fee
Basic fee

$ 979,536

Performance adjustment

37,062

Transfer agent fees

534,768

Distribution fees

10,735

Accounting and security lending fees

70,624

Custodian fees and expenses

18,147

Independent trustees' compensation

638

Registration fees

103,527

Audit

55,247

Legal

928

Miscellaneous

11,842

Total expenses before reductions

1,823,054

Expense reductions

(97,947)

1,725,107

Net investment income (loss)

(131,322)

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

10,237,592

Change in net unrealized appreciation (depreciation) on investment securities

(21,747,840)

Net gain (loss)

(11,510,248)

Net increase (decrease) in net assets resulting from operations

$ (11,641,570)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ (131,322)

$ 26,323

Net realized gain (loss)

10,237,592

4,571,573

Change in net unrealized appreciation (depreciation)

(21,747,840)

(297,654)

Net increase (decrease) in net assets resulting from operations

(11,641,570)

4,300,242

Distributions to shareholders from net investment income

-

(137,207)

Distributions to shareholders from net realized gain

(18,572,404)

(3,644,497)

Total distributions

(18,572,404)

(3,781,704)

Share transactions - net increase (decrease)

(1,164,336)

25,478,221

Redemption fees

-

5,257

Total increase (decrease) in net assets

(31,378,310)

26,002,016

Net Assets

Beginning of period

183,515,484

157,513,468

End of period

$ 152,137,174

$ 183,515,484

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.03)

Net realized and unrealized gain (loss)

(.72)

Total from investment operations

(.75)

Distributions from net realized gain

(1.25)

Net asset value, end of period

$ 9.85

Total Return B,C,D

(6.99)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.20% A

Expenses net of fee waivers, if any

1.20% A

Expenses net of all reductions

1.20% A

Net investment income (loss)

(.29)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,302

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.06)

Net realized and unrealized gain (loss)

(.69)

Total from investment operations

(.75)

Distributions from net realized gain

(1.25)

Net asset value, end of period

$ 9.85

Total Return B,C,D

(7.05)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.47% A

Expenses net of fee waivers, if any

1.47% A

Expenses net of all reductions

1.47% A

Net investment income (loss)

(.56)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,097

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.12)

Net realized and unrealized gain (loss)

(.70)

Total from investment operations

(.82)

Distributions from net realized gain

(1.20)

Net asset value, end of period

$ 9.83

Total Return B,C,D

(7.62)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.99% A

Expenses net of fee waivers, if any

1.99% A

Expenses net of all reductions

1.99% A

Net investment income (loss)

(1.07)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 543

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.11)

Net realized and unrealized gain (loss)

(.70)

Total from investment operations

(.81)

Distributions from net realized gain

(1.22)

Net asset value, end of period

$ 9.82

Total Return B,C,D

(7.54)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.96% A

Expenses net of fee waivers, if any

1.96% A

Expenses net of all reductions

1.96% A

Net investment income (loss)

(1.05)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 945

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Large Cap Growth

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 11.92

$ 11.82

$ 10.17

$ 9.21

$ 6.93

Income from Investment Operations

Net investment income (loss) B

(.01)

- H

.02 E

(.01) F

(.01)

Net realized and unrealized gain (loss)

(.77)

.37

1.87

.97

2.29

Total from investment operations

(.78)

.37

1.89

.96

2.28

Distributions from net investment income

-

(.01)

-

-

-

Distributions from net realized gain

(1.25)

(.26)

(.24)

-

-

Total distributions

(1.25)

(.27)

(.24)

-

-

Redemption fees added to paid in capital B

-

- H,I

- H

- H

- H

Net asset value, end of period

$ 9.89

$ 11.92

$ 11.82

$ 10.17

$ 9.21

Total Return A

(7.26)%

3.20%

18.66%

10.42%

32.90%

Ratios to Average Net Assets C,G

Expenses before reductions

1.03%

1.10%

1.12%

1.30%

1.53%

Expenses net of fee waivers, if any

.99%

1.00%

1.00%

1.20%

1.20%

Expenses net of all reductions

.98%

.99%

.94%

1.13%

1.18%

Net investment income (loss)

(.07)%

.02%

.15% E

(.07)% F

(.15)%

Supplemental Data

Net assets, end of period (000 omitted)

$ 147,864

$ 183,515

$ 157,513

$ 49,453

$ 23,079

Portfolio turnover rate D

428%

189%

268%

274%

81%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.07)%.

F Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.32)%.

G Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

H Amount represents less than $.01 per share.

I The redemption fee was eliminated during the year ended January 31, 2007.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net realized and unrealized gain (loss)

(.70)

Distributions from net realized gain

(1.27)

Net asset value, end of period

$ 9.88

Total Return B,C

(6.64)%

Ratios to Average Net Assets D,E

Expenses before reductions

.88% A

Expenses net of fee waivers, if any

.88% A

Expenses net of all reductions

.88% A

Net investment income (loss)

.03% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 386

Portfolio turnover rate F

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Advisor Mid Cap Growth Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the class' dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table does not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Class A (incl. 5.75% sales charge) B

-15.01%

11.83%

4.05%

Class T (incl. 3.50% sales charge) C

-13.32%

12.27%

4.38%

Class B (incl. contingent deferred sales charge) D

-14.75%

12.74%

4.92%

Class C (incl. contingent deferred sales charge) E

-11.37%

12.98%

4.92%

A From November 15, 2001.

B Class A shares bear a 0.25% 12b-1 fee. The initial offering of Class A shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Growth, the original class of the fund, which has no 12b-1 fee. Had Class A shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower.

C Class T shares bear a 0.50% 12b-1 fee. The initial offering of Class T shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Growth, the original class of the fund, which has no 12b-1 fee. Had Class T shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower.

D Class B shares bear a 1.00% 12b-1 fee. The initial offering of Class B shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Growth, the original class of the fund, which has no 12b-1 fee. Had Class B shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower. Class B shares' contingent deferred sales charge included in the past one year, past five years, and life of fund total return figures are 5%, 2%, and 0%, respectively.

E Class C shares bear a 1.00% 12b-1 fee. The initial offering of Class C shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Growth, the original class of the fund, which has no 12b-1 fee. Had Class C shares' 12b-1 fee been reflected, returns prior to February 13, 2007 would have been lower. Class C shares' contingent deferred sales charge included in the past one year, past five years, and life of fund total return figures are 1%, 0%, and 0%, respectively.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Fidelity Advisor Mid Cap Growth Fund - Class A on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell Midcap® Growth Index performed over the same period. The initial offering of Class A took place on February 13, 2007. See above for additional information regarding performance of Class A.



Annual Report

Fidelity Advisor Mid Cap Growth Fund

Management's Discussion of Fund Performance

Comments from Patrick Venanzi, Portfolio Manager of Fidelity Advisor Mid Cap Growth Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

The fund's Class A, Class T, Class B and Class C shares (excluding sales charges) lagged behind the -1.03% return for the Russell Midcap® Growth Index. (For specific class-level results, please refer to the performance section of this report.) Poor stock selection in consumer discretionary, industrials, materials and consumer staples hurt the most. While the fund struggled to keep pace with the index for most of the period, stock selection in energy and health care helped limit some of the downside. Within industrials, the fund's overweighting in transportation stocks, specifically in the airlines, detracted from performance. Our holdings in US Airways and AMR Corp., the parent company of American Airlines, suffered from rising energy costs. Overweighted holdings in the consumer durables and apparel segment - primarily the homebuilders - also took a bite out of fund performance. D.R. Horton, Ryland Group, KB Home and Lennar were particularly hard hit as the U.S. residential real estate market continued to slow down. I sold D.R. Horton, Ryland Group and Lennar. On the positive side, energy services company National Oilwell Varco was the fund's biggest contributor. Another contributor was the fund's out-of-index position in consumer electronics giant Apple, which continued to gain from strong sales of its iPod, iPhone and iMac product lines. I sold the fund's position in Apple to lock in profits. Heavy-lifting equipment and crane manufacturer Manitowoc advanced due to the continuing worldwide build-out of commercial and industrial infrastructure. I also sold this position by the end of the period. BE Aerospace, which makes products for aircraft cabin interiors, also contributed, as investors expressed confidence in the company's new contracts to supply airline carriers that are upgrading their fleets. Lastly, MEMC Electronic Materials, which produces silicon wafers, helped as well.

For the year, the fund's Institutional Class shares lagged behind the -1.03% return for the Russell Midcap® Growth Index. (For specific class-level results, please refer to the performance section of this report.) Poor stock selection in consumer discretionary, industrials, materials and consumer staples hurt the most. While the fund struggled to keep pace with the index for most of the period, stock selection in energy and health care helped limit some of the downside. Within industrials, the fund's overweighting in transportation stocks, specifically in the airlines, detracted from performance. Our holdings in US Airways and AMR Corp., the parent company of American Airlines, suffered from rising energy costs. Overweighted holdings in the consumer durables and apparel segment - primarily the homebuilders - also took a bite out of fund performance. D.R. Horton, Ryland Group, KB Home and Lennar were particularly hard hit as the U.S. residential real estate market continued to slow down. I sold D.R. Horton, Ryland Group and Lennar. On the positive side, energy services company National Oilwell Varco was the fund's biggest contributor. Another contributor was the fund's out-of-index position in consumer electronics giant Apple, which continued to gain from strong sales of its iPod, iPhone and iMac product lines. I sold the fund's position in Apple to lock in profits. Heavy-lifting equipment and crane manufacturer Manitowoc advanced due to the continuing worldwide build-out of commercial and industrial infrastructure. I also sold this position by the end of the period. BE Aerospace, which makes products for aircraft cabin interiors, also contributed, as investors expressed confidence in the company's new contracts to supply airline carriers that are upgrading their fleets. Lastly, MEMC Electronic Materials, which produces silicon wafers, helped as well.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Mid Cap Growth Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

PPL Corp.

1.4

0.7

Southwestern Energy Co.

1.4

0.3

Constellation Energy Group, Inc.

1.3

1.0

National Oilwell Varco, Inc.

1.3

3.9

Align Technology, Inc.

1.1

0.0

Centex Corp.

1.1

0.0

TranS1, Inc.

1.0

0.0

Polycom, Inc.

0.9

0.0

Intuitive Surgical, Inc.

0.9

0.0

Northwest Airlines Corp.

0.9

0.0

11.3

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Information Technology

19.8

19.0

Industrials

17.1

16.1

Health Care

16.6

12.9

Consumer Discretionary

14.1

17.6

Energy

11.6

10.9

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 100.2%

Stocks 97.8%

Short-Term
Investments and
Net Other Assets(dagger) (0.2)%

Short-Term
Investments and
Net Other Assets 2.2%

* Foreign
investments

11.7%

**Foreign investments

2.1%

(dagger) Short-Term Investments
and Net Other Assets
are not included in the pie chart.

Annual Report

Fidelity Mid Cap Growth Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 100.2%

Shares

Value

CONSUMER DISCRETIONARY - 14.1%

Automobiles - 0.3%

Renault SA

7,300

$ 833,337

Diversified Consumer Services - 1.1%

American Public Education, Inc.

7,200

283,752

Apollo Group, Inc. Class A (non-vtg.) (a)

8,200

653,868

DeVry, Inc.

19,000

1,048,610

Regis Corp.

56,400

1,428,612

3,414,842

Hotels, Restaurants & Leisure - 2.1%

Bally Technologies, Inc. (a)

20,200

962,328

Boyd Gaming Corp.

66,400

1,774,872

Buffalo Wild Wings, Inc. (a)

8,400

211,428

Burger King Holdings, Inc.

54,789

1,443,690

Gaylord Entertainment Co. (a)

29,700

866,943

Papa John's International, Inc. (a)

26,100

660,069

Peet's Coffee & Tea, Inc. (a)

7,000

153,580

Starwood Hotels & Resorts Worldwide, Inc.

5,400

244,350

6,317,260

Household Durables - 2.1%

Centex Corp. (d)

116,700

3,241,926

KB Home

100

2,750

Newell Rubbermaid, Inc.

15,100

364,212

Pulte Homes, Inc.

33,500

547,390

Whirlpool Corp.

25,700

2,187,327

6,343,605

Internet & Catalog Retail - 1.2%

Blue Nile, Inc. (a)

17,400

961,350

Expedia, Inc. (a)

10,600

244,012

Priceline.com, Inc. (a)

21,099

2,289,663

3,495,025

Leisure Equipment & Products - 0.7%

Hasbro, Inc.

12,400

322,028

Leapfrog Enterprises, Inc. Class A (a)

36,300

236,313

Mattel, Inc.

53,533

1,124,728

Nikon Corp.

14,000

381,830

2,064,899

Media - 2.7%

Cablevision Systems Corp. - NY Group Class A (a)

30,300

711,444

DISH Network Corp. Class A (a)

13,057

368,730

E.W. Scripps Co. Class A

35,800

1,457,776

Grupo Televisa SA de CV (CPO) sponsored ADR

94,000

2,095,260

Regal Entertainment Group Class A

87,100

1,614,834

SR Teleperformance SA

44,300

1,386,369

The Walt Disney Co.

11,700

350,181

Time Warner, Inc.

21,600

339,984

8,324,578

Shares

Value

Multiline Retail - 0.1%

Dollar Tree Stores, Inc. (a)

14,300

$ 400,543

Specialty Retail - 3.2%

Abercrombie & Fitch Co. Class A

5,100

406,419

Advance Auto Parts, Inc.

30,500

1,088,240

Asbury Automotive Group, Inc.

13,100

185,758

AutoZone, Inc. (a)

8,800

1,063,744

Gamestop Corp. Class A (a)

5,400

279,342

Group 1 Automotive, Inc.

8,700

230,028

Home Depot, Inc.

19,700

604,199

Lumber Liquidators, Inc.

3,400

30,260

OfficeMax, Inc.

49,800

1,233,546

Ross Stores, Inc.

36,300

1,058,145

Tween Brands, Inc. (a)(d)

44,700

1,431,741

Urban Outfitters, Inc. (a)

28,800

835,200

Williams-Sonoma, Inc.

49,500

1,330,560

Zumiez, Inc. (a)

3,500

67,305

9,844,487

Textiles, Apparel & Luxury Goods - 0.6%

Coach, Inc. (a)

13,000

416,650

Deckers Outdoor Corp. (a)

2,800

339,472

Lululemon Athletica, Inc.

15,700

532,073

Under Armour, Inc. Class A (sub. vtg.) (a)

16,000

644,000

1,932,195

TOTAL CONSUMER DISCRETIONARY

42,970,771

CONSUMER STAPLES - 3.4%

Beverages - 0.1%

Coca-Cola Femsa SA de CV sponsored ADR

7,200

338,040

Food & Staples Retailing - 1.3%

CVS Caremark Corp.

14,600

570,422

Performance Food Group Co. (a)

48,000

1,518,240

Safeway, Inc.

25,200

780,948

Sysco Corp.

42,100

1,223,005

4,092,615

Food Products - 1.5%

Campbell Soup Co.

17,300

546,853

Marine Harvest ASA (a)

2,600,000

1,400,493

McCormick & Co., Inc. (non-vtg.)

55,300

1,864,716

Seaboard Corp.

370

475,543

Wm. Wrigley Jr. Co.

3,000

172,290

4,459,895

Household Products - 0.4%

Energizer Holdings, Inc. (a)

13,300

1,245,146

Personal Products - 0.1%

Estee Lauder Companies, Inc. Class A

8,900

375,580

TOTAL CONSUMER STAPLES

10,511,276

Common Stocks - continued

Shares

Value

ENERGY - 11.6%

Energy Equipment & Services - 5.0%

Atwood Oceanics, Inc. (a)

21,000

$ 1,744,890

Cameron International Corp. (a)

31,700

1,276,242

Diamond Offshore Drilling, Inc.

16,700

1,885,931

Grey Wolf, Inc. (a)

297,600

1,773,696

Nabors Industries Ltd. (a)

35,000

952,700

National Oilwell Varco, Inc. (a)

65,120

3,922,178

North American Energy Partners, Inc. (a)

14,200

170,116

Ocean RIG ASA (a)(d)

114,500

787,337

Smith International, Inc.

23,900

1,295,619

Transocean, Inc. (a)

10,961

1,343,819

15,152,528

Oil, Gas & Consumable Fuels - 6.6%

Alpha Natural Resources, Inc. (a)

21,500

719,390

Arch Coal, Inc.

15,700

690,800

Boardwalk Pipeline Partners, LP

5,700

177,156

Canadian Natural Resources Ltd.

10,100

646,227

Chesapeake Energy Corp.

46,600

1,734,918

CONSOL Energy, Inc.

35,300

2,576,900

Copano Energy LLC

4,900

173,166

Denbury Resources, Inc. (a)

65,700

1,662,210

Energy Transfer Equity LP

5,200

171,652

Foundation Coal Holdings, Inc.

7,000

366,100

Frontier Oil Corp.

38,100

1,343,787

Holly Corp.

28,601

1,384,860

OPTI Canada, Inc. (a)

80,400

1,325,913

Petrobank Energy & Resources Ltd. (a)

7,000

351,761

Southwestern Energy Co. (a)

76,100

4,254,751

Tesoro Corp.

22,400

874,720

Valero Energy Corp.

100

5,919

Western Refining, Inc.

44,460

949,221

Williams Companies, Inc.

27,100

866,387

20,275,838

TOTAL ENERGY

35,428,366

FINANCIALS - 9.2%

Capital Markets - 2.6%

Franklin Resources, Inc.

23,600

2,459,828

KBW, Inc. (a)

6,300

187,173

Legg Mason, Inc.

14,500

1,044,000

MF Global Ltd.

73,200

2,199,660

SEI Investments Co.

67,400

1,866,980

T. Rowe Price Group, Inc.

3,300

166,947

7,924,588

Commercial Banks - 1.5%

Bank of Yokohama Ltd.

84,000

544,305

Chiba Bank Ltd.

52,000

384,877

Chuo Mitsui Trust Holdings, Inc.

55,000

377,598

Mitsubishi UFJ Financial Group, Inc.

70,000

691,600

Shinsei Bank Ltd.

117,000

534,769

Shares

Value

Shizuoka Bank Ltd.

40,000

$ 439,763

Sumitomo Mitsui Financial Group, Inc.

100

788,113

Tokyo Tomin Bank Ltd.

20,600

572,491

UMB Financial Corp.

7,600

320,188

4,653,704

Consumer Finance - 0.4%

Cash America International, Inc.

37,100

1,206,121

Diversified Financial Services - 1.5%

CME Group, Inc.

800

495,120

IntercontinentalExchange, Inc. (a)

9,400

1,315,624

MSCI, Inc. Class A

11,800

389,046

NYMEX Holdings, Inc.

18,400

2,116,000

Osaka Securities Exchange Co. Ltd.

64

346,092

4,661,882

Insurance - 0.9%

Assurant, Inc.

19,400

1,258,866

CNA Financial Corp.

11,480

390,205

CNinsure, Inc. ADR

44,000

472,560

LandAmerica Financial Group, Inc.

4,900

255,584

Philadelphia Consolidated Holdings Corp. (a)

9,600

343,680

Stewart Information Services Corp.

1,100

37,653

2,758,548

Real Estate Investment Trusts - 1.5%

Annaly Capital Management, Inc.

124,000

2,445,280

MFA Mortgage Investments, Inc.

189,300

1,930,860

4,376,140

Thrifts & Mortgage Finance - 0.8%

People's United Financial, Inc.

58,700

991,443

Washington Federal, Inc.

55,900

1,365,078

2,356,521

TOTAL FINANCIALS

27,937,504

HEALTH CARE - 16.6%

Biotechnology - 3.5%

Alexion Pharmaceuticals, Inc. (a)

18,500

1,208,420

Amylin Pharmaceuticals, Inc. (a)

74,607

2,212,098

Biogen Idec, Inc. (a)

26,400

1,609,080

Cephalon, Inc. (a)

11,300

741,619

Genentech, Inc. (a)

8,000

561,520

Gilead Sciences, Inc. (a)

41,362

1,889,830

Theravance, Inc. (a)(d)

127,038

2,506,460

10,729,027

Health Care Equipment & Supplies - 5.8%

Abiomed, Inc. (a)

25,000

377,500

Align Technology, Inc. (a)(d)

283,984

3,345,332

American Medical Systems Holdings, Inc. (a)

90,000

1,286,100

Conceptus, Inc. (a)

158,500

2,580,380

Hillenbrand Industries, Inc.

13,300

687,876

Hologic, Inc. (a)

14,000

901,040

Common Stocks - continued

Shares

Value

HEALTH CARE - continued

Health Care Equipment & Supplies - continued

Intuitive Surgical, Inc. (a)

11,000

$ 2,794,000

Inverness Medical Innovations, Inc. (a)

3,000

135,150

Mindray Medical International Ltd. sponsored ADR

10,000

341,000

NxStage Medical, Inc. (a)

14,100

171,174

Regeneration Technologies, Inc. (a)

215,000

1,720,000

TranS1, Inc. (d)

213,000

3,107,670

Varian Medical Systems, Inc. (a)

5,300

275,547

17,722,769

Health Care Providers & Services - 2.4%

athenahealth, Inc. (d)

49,400

1,551,160

Brookdale Senior Living, Inc. (d)

11,000

245,520

Coventry Health Care, Inc. (a)

5,500

311,190

Express Scripts, Inc. (a)

28,600

1,930,214

Health Net, Inc. (a)

10,300

478,847

Henry Schein, Inc. (a)

5,473

318,145

MWI Veterinary Supply, Inc. (a)

8,000

305,840

Pediatrix Medical Group, Inc. (a)

24,587

1,674,129

Psychiatric Solutions, Inc. (a)

5,900

178,003

VCA Antech, Inc. (a)

10,200

394,332

7,387,380

Health Care Technology - 1.0%

Cerner Corp. (a)

43,200

2,263,680

MedAssets, Inc.

7,900

159,185

TriZetto Group, Inc. (a)

38,900

759,328

3,182,193

Life Sciences Tools & Services - 2.0%

Affymetrix, Inc. (a)(d)

60,600

1,215,636

Applera Corp. - Applied Biosystems Group

30,000

945,900

Charles River Laboratories International, Inc. (a)

19,700

1,223,370

Covance, Inc. (a)

1,800

149,688

PAREXEL International Corp. (a)

10,000

544,100

Pharmaceutical Product Development, Inc.

30,200

1,309,472

Thermo Fisher Scientific, Inc. (a)

12,770

657,527

6,045,693

Pharmaceuticals - 1.9%

Allergan, Inc.

20,400

1,370,676

BioForm Medical, Inc. (d)

186,481

1,305,367

Perrigo Co.

15,700

484,188

Renovo Group PLC (a)

342,000

920,384

Teva Pharmaceutical Industries Ltd. sponsored ADR

22,300

1,026,692

XenoPort, Inc. (a)

8,600

527,696

5,635,003

TOTAL HEALTH CARE

50,702,065

Shares

Value

INDUSTRIALS - 17.1%

Aerospace & Defense - 1.0%

BE Aerospace, Inc. (a)

21,444

$ 827,953

Goodrich Corp.

19,900

1,244,745

Spirit AeroSystems Holdings, Inc.
Class A (a)

31,700

875,554

2,948,252

Air Freight & Logistics - 0.2%

United Parcel Service, Inc. Class B

6,400

468,224

Airlines - 2.3%

AMR Corp. (a)

103,000

1,435,820

Continental Airlines, Inc. Class B (a)

63,900

1,738,719

Northwest Airlines Corp. (a)

146,700

2,744,757

US Airways Group, Inc. (a)

89,200

1,231,852

7,151,148

Building Products - 0.3%

Ameron International Corp.

8,000

718,800

Universal Forest Products, Inc.

6,000

217,200

936,000

Commercial Services & Supplies - 5.1%

Advisory Board Co. (a)

2,600

165,672

American Reprographics Co. (a)

46,200

726,726

Copart, Inc. (a)

33,984

1,389,266

Corporate Executive Board Co.

9,400

540,970

Corrections Corp. of America (a)

94,182

2,499,590

Fuel Tech, Inc. (a)

35,600

677,824

GeoEye, Inc. (a)

15,000

524,400

IHS, Inc. Class A (a)

11,500

712,310

InnerWorkings, Inc. (a)

12,000

166,200

Manpower, Inc.

45,400

2,554,204

Republic Services, Inc.

56,600

1,698,000

The Geo Group, Inc. (a)

54,600

1,306,032

Waste Connections, Inc. (a)

45,600

1,329,696

Waste Management, Inc.

34,400

1,115,936

15,406,826

Construction & Engineering - 0.7%

Chicago Bridge & Iron Co. NV
(NY Shares)

4,900

218,001

Fluor Corp.

7,500

912,525

Grupo Acciona SA

2,000

509,808

Quanta Services, Inc. (a)

20,000

438,400

2,078,734

Electrical Equipment - 3.9%

Belden, Inc.

7,000

296,100

Energy Conversion Devices, Inc. (a)(d)

57,020

1,316,022

Evergreen Solar, Inc. (a)

154,800

1,887,012

First Solar, Inc. (a)

10,300

1,872,231

JA Solar Holdings Co. Ltd. ADR

16,000

813,280

Nexans SA

19,800

2,188,427

Q-Cells AG (a)

6,000

564,589

Sunpower Corp. Class A (a)(d)

35,400

2,445,786

Common Stocks - continued

Shares

Value

INDUSTRIALS - continued

Electrical Equipment - continued

Superior Essex, Inc. (a)

13,100

$ 315,055

Vestas Wind Systems AS (a)

2,800

271,999

11,970,501

Machinery - 1.3%

AGCO Corp. (a)

20,600

1,240,532

GEA Group AG (a)

32,000

988,991

Ingersoll-Rand Co. Ltd. Class A

18,000

711,360

Terex Corp. (a)

17,100

1,004,796

3,945,679

Marine - 0.4%

Ultrapetrol (Bahamas) Ltd. (a)

70,100

1,125,105

Road & Rail - 1.7%

J.B. Hunt Transport Services, Inc. (d)

56,100

1,744,710

Knight Transportation, Inc.

102,900

1,765,764

Landstar System, Inc.

36,500

1,826,095

5,336,569

Trading Companies & Distributors - 0.2%

Genesis Lease Ltd. ADR

31,600

635,160

TOTAL INDUSTRIALS

52,002,198

INFORMATION TECHNOLOGY - 19.8%

Communications Equipment - 2.0%

Ciena Corp. (a)

15,000

406,950

CommScope, Inc. (a)

11,700

518,895

Corning, Inc.

22,000

529,540

Juniper Networks, Inc. (a)

64,700

1,756,605

Polycom, Inc. (a)

111,000

2,802,750

6,014,740

Computers & Peripherals - 1.7%

Brocade Communications Systems, Inc. (a)

226,602

1,561,288

Diebold, Inc.

28,644

741,307

NCR Corp. (a)

61,600

1,323,168

Western Digital Corp. (a)

54,600

1,444,170

5,069,933

Electronic Equipment & Instruments - 1.6%

Agilent Technologies, Inc. (a)

8,700

295,017

Avnet, Inc. (a)

17,300

616,053

Bell Microproducts, Inc. (a)

29,818

156,843

Comverge, Inc.

6,100

113,704

Flextronics International Ltd. (a)

51,100

597,870

Ingram Micro, Inc. Class A (a)

30,700

545,846

Itron, Inc. (a)

24,000

1,977,600

Jabil Circuit, Inc.

47,300

626,725

4,929,658

Internet Software & Services - 2.8%

Akamai Technologies, Inc. (a)

16,800

507,360

DealerTrack Holdings, Inc. (a)

23,600

636,256

Shares

Value

Dice Holdings, Inc.

258,600

$ 1,649,868

DivX, Inc. (a)

50,000

712,500

Internet Brands, Inc. Class A

30,979

239,158

Omniture, Inc. (a)

85,700

2,118,504

Online Resources Corp. (a)

89,800

916,858

The Knot, Inc. (a)

49,700

727,608

ValueClick, Inc. (a)

44,100

962,703

8,470,815

IT Services - 1.1%

Cognizant Technology Solutions Corp. Class A (a)

72,700

2,028,330

CyberSource Corp. (a)

38,000

636,500

Mastercard, Inc. Class A

3,800

786,600

3,451,430

Semiconductors & Semiconductor Equipment - 7.4%

Altera Corp.

93,000

1,570,770

Applied Materials, Inc.

125,400

2,247,168

ARM Holdings PLC sponsored ADR

23,600

167,088

ASML Holding NV (NY Shares) (a)

20,000

531,800

Broadcom Corp. Class A (a)

66,500

1,468,320

Cypress Semiconductor Corp. (a)

62,200

1,321,750

FormFactor, Inc. (a)

21,063

510,146

Himax Technologies, Inc. sponsored ADR

285,100

1,405,543

Hittite Microwave Corp. (a)

12,400

493,768

Intersil Corp. Class A

31,400

723,142

Lam Research Corp. (a)

46,500

1,785,135

LDK Solar Co. Ltd. Sponsored ADR (d)

19,800

697,950

Marvell Technology Group Ltd. (a)

122,180

1,450,277

MEMC Electronic Materials, Inc. (a)

18,500

1,322,010

Microchip Technology, Inc.

35,500

1,132,805

Miraial Co. Ltd.

15,900

386,547

National Semiconductor Corp.

53,000

976,790

Skyworks Solutions, Inc. (a)(d)

124,700

1,003,835

Teradyne, Inc. (a)

5,400

59,238

Varian Semiconductor Equipment Associates, Inc. (a)

63,800

2,054,998

Xilinx, Inc.

61,200

1,338,444

22,647,524

Software - 3.2%

Activision, Inc. (a)

25,100

649,337

Citrix Systems, Inc. (a)

12,000

415,440

CompuGROUP Holding AG (a)

33,000

622,352

Concur Technologies, Inc. (a)

53,300

1,868,698

FactSet Research Systems, Inc.

6,000

335,580

Jack Henry & Associates, Inc.

53,300

1,310,114

Microsoft Corp.

20,800

678,080

Nintendo Co. Ltd.

400

197,600

Nuance Communications, Inc. (a)

87,000

1,382,430

Shanda Interactive Entertainment Ltd. sponsored ADR (a)

17,900

492,071

SPSS, Inc. (a)

9,000

297,450

Common Stocks - continued

Shares

Value

INFORMATION TECHNOLOGY - continued

Software - continued

Take-Two Interactive Software, Inc. (a)(d)

61,650

$ 1,013,526

THQ, Inc. (a)

33,800

608,738

9,871,416

TOTAL INFORMATION TECHNOLOGY

60,455,516

MATERIALS - 3.1%

Chemicals - 0.2%

FMC Corp.

11,300

600,708

Construction Materials - 0.5%

Martin Marietta Materials, Inc. (d)

13,800

1,693,536

Containers & Packaging - 0.4%

Smurfit-Stone Container Corp. (a)

39,600

375,804

Temple-Inland, Inc.

39,600

742,500

1,118,304

Metals & Mining - 1.9%

Allegheny Technologies, Inc.

19,542

1,375,757

Barrick Gold Corp.

6,900

356,224

Century Aluminum Co. (a)

14,000

727,860

Newmont Mining Corp.

10,000

543,400

Nucor Corp.

22,200

1,283,160

Randgold Resources Ltd. sponsored ADR

5,800

276,718

RTI International Metals, Inc. (a)

23,140

1,278,485

5,841,604

Paper & Forest Products - 0.1%

Glatfelter

12,400

179,304

TOTAL MATERIALS

9,433,456

TELECOMMUNICATION SERVICES - 1.3%

Wireless Telecommunication Services - 1.3%

American Tower Corp. Class A (a)

52,705

1,978,019

NII Holdings, Inc. (a)

48,691

2,077,158

4,055,177

UTILITIES - 4.0%

Electric Utilities - 2.1%

Allegheny Energy, Inc.

17,600

964,304

Enernoc, Inc.

14,300

504,075

Shares

Value

Entergy Corp.

4,500

$ 486,810

PPL Corp.

89,800

4,393,015

6,348,204

Independent Power Producers & Energy Traders - 1.9%

AES Corp. (a)

79,400

1,514,952

Clipper Windpower PLC (a)

2,400

28,693

Constellation Energy Group, Inc.

42,800

4,021,488

Ormat Technologies, Inc.

3,800

165,186

5,730,319

TOTAL UTILITIES

12,078,523

TOTAL COMMON STOCKS

(Cost $305,398,397)

305,574,852

Money Market Funds - 4.3%

Fidelity Cash Central Fund, 3.79% (b)

2,288,816

2,288,816

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

10,764,675

10,764,675

TOTAL MONEY MARKET FUNDS

(Cost $13,053,491)

13,053,491

TOTAL INVESTMENT PORTFOLIO - 104.5%

(Cost $318,451,888)

318,628,343

NET OTHER ASSETS - (4.5)%

(13,583,264)

NET ASSETS - 100%

$ 305,045,079

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 442,525

Fidelity Securities Lending Cash Central Fund

313,679

Total

$ 756,204

Other Information

Distribution of investments by country of issue, as a percentage of total net assets, is as follows:

United States of America

88.3%

Japan

1.8%

Bermuda

1.7%

France

1.5%

Cayman Islands

1.3%

Others (individually less than 1%)

5.4%

100.0%

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $18,706,577 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Growth Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $10,830,743) - See accompanying schedule:

Unaffiliated issuers (cost $305,398,397)

$ 305,574,852

Fidelity Central Funds (cost $13,053,491)

13,053,491

Total Investments (cost $318,451,888)

$ 318,628,343

Cash

944,763

Receivable for investments sold

60,992,695

Receivable for fund shares sold

386,427

Dividends receivable

50,720

Distributions receivable from Fidelity Central Funds

53,069

Prepaid expenses

1,182

Other receivables

2,063

Total assets

381,059,262

Liabilities

Payable for investments purchased

$ 23,187,980

Payable for fund shares redeemed

41,783,947

Accrued management fee

108,332

Distribution fees payable

1,571

Other affiliated payables

110,841

Other payables and accrued expenses

56,837

Collateral on securities loaned, at value

10,764,675

Total liabilities

76,014,183

Net Assets

$ 305,045,079

Net Assets consist of:

Paid in capital

$ 319,016,951

Accumulated undistributed net realized gain (loss) on investments and foreign currency transactions

(14,150,236)

Net unrealized appreciation (depreciation) on investments and assets and liabilities in foreign currencies

178,364

Net Assets

$ 305,045,079

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($1,936,249 ÷ 158,895 shares)

$ 12.19

Maximum offering price per share (100/94.25 of $12.19)

$ 12.93

Class T:
Net Asset Value
and redemption price per share ($591,188 ÷ 48,688 shares)

$ 12.14

Maximum offering price per share (100/96.50 of $12.14)

$ 12.58

Class B:
Net Asset Value
and offering price per share ($413,537 ÷ 34,216 shares)A

$ 12.09

Class C:
Net Asset Value
and offering price per share ($697,298 ÷ 57,672 shares)A

$ 12.09

Mid Cap Growth:
Net Asset Value
, offering price and redemption price per share ($301,224,741 ÷ 24,671,114 shares)

$ 12.21

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($182,066 ÷ 14,896 shares)

$ 12.22

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Growth Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 2,094,532

Interest

386

Income from Fidelity Central Funds (including $313,679 from security lending)

756,204

Total income

2,851,122

Expenses

Management fee
Basic fee

$ 2,332,314

Performance adjustment

(430,070)

Transfer agent fees

1,202,314

Distribution fees

10,363

Accounting and security lending fees

170,995

Custodian fees and expenses

18,214

Independent trustees' compensation

1,531

Registration fees

100,229

Audit

56,662

Legal

2,290

Miscellaneous

25,324

Total expenses before reductions

3,490,166

Expense reductions

(113,488)

3,376,678

Net investment income (loss)

(525,556)

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

13,581,162

Foreign currency transactions

16,150

Total net realized gain (loss)

13,597,312

Change in net unrealized appreciation (depreciation) on:

Investment securities

(50,635,596)

Assets and liabilities in foreign currencies

1,909

Total change in net unrealized appreciation (depreciation)

(50,633,687)

Net gain (loss)

(37,036,375)

Net increase (decrease) in net assets resulting from operations

$ (37,561,931)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ (525,556)

$ (1,458,396)

Net realized gain (loss)

13,597,312

(4,757,238)

Change in net unrealized appreciation (depreciation)

(50,633,687)

5,161,873

Net increase (decrease) in net assets resulting from operations

(37,561,931)

(1,053,761)

Distributions to shareholders from net realized gain

(21,909,497)

(5,089,264)

Share transactions - net increase (decrease)

(76,821,412)

97,426,141

Redemption fees

26,012

46,437

Total increase (decrease) in net assets

(136,266,828)

91,329,553

Net Assets

Beginning of period

441,311,907

349,982,354

End of period

$ 305,045,079

$ 441,311,907

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.05)

Net realized and unrealized gain (loss)

(1.30)

Total from investment operations

(1.35)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.19

Total Return B,C,D

(9.95)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.10% A

Expenses net of fee waivers, if any

1.10% A

Expenses net of all reductions

1.10% A

Net investment income (loss)

(.41)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,936

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.09)

Net realized and unrealized gain (loss)

(1.31)

Total from investment operations

(1.40)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.14

Total Return B,C,D

(10.30)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.36% A

Expenses net of fee waivers, if any

1.36% A

Expenses net of all reductions

1.36% A

Net investment income (loss)

(.68)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 591

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.16)

Net realized and unrealized gain (loss)

(1.29)

Total from investment operations

(1.45)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.09

Total Return B,C,D

(10.65)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.85% A

Expenses net of fee waivers, if any

1.85% A

Expenses net of all reductions

1.85% A

Net investment income (loss)

(1.16)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 414

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.15)

Net realized and unrealized gain (loss)

(1.30)

Total from investment operations

(1.45)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.09

Total Return B,C,D

(10.65)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.85% A

Expenses net of fee waivers, if any

1.85% A

Expenses net of all reductions

1.85% A

Net investment income (loss)

(1.16)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 697

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Mid Cap Growth

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 14.31

$ 14.38

$ 11.58

$ 10.63

$ 7.32

Income from Investment Operations

Net investment income (loss) B

(.02)

(.04)

.01 E

(.03) F

(.07)

Net realized and unrealized gain (loss)

(1.29)

.15

3.09

1.15

3.38

Total from investment operations

(1.31)

.11

3.10

1.12

3.31

Distributions from net realized gain

(.79)

(.18)

(.30)

(.17)

-

Redemption fees added to paid in capital B,H

-

-

-

-

-

Net asset value, end of period

$ 12.21

$ 14.31

$ 14.38

$ 11.58

$ 10.63

Total Return A

(9.68)%

.80%

27.15%

10.55%

45.22%

Ratios to Average Net Assets C,G

Expenses before reductions

.83%

1.02%

1.04%

1.02%

1.25%

Expenses net of fee waivers, if any

.81%

1.00%

1.00%

1.02%

1.20%

Expenses net of all reductions

.81%

.99%

.95%

.99%

1.16%

Net investment income (loss)

(.12)%

(.33)%

.07% E

(.31)% F

(.77)%

Supplemental Data

Net assets, end of period (000 omitted)

$ 301,225

$ 441,312

$ 349,982

$ 77,658

$ 60,660

Portfolio turnover rate D

245%

178%

173%

220%

94%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.03 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.18)%.

F Investment income per share reflects a special dividend which amounted to $.01 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.44)%.

G Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

H Amount represents less than $.01 per share.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net realized and unrealized gain (loss) D

(1.32)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.22

Total Return B,C

(9.74)%

Ratios to Average Net Assets E,H

Expenses before reductions

.72% A

Expenses net of fee waivers, if any

.72% A

Expenses net of all reductions

.72% A

Net investment income (loss)

(.03)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 182

Portfolio turnover rate F

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Notes to Financial Statements

For the period ended January 31, 2008

1. Organization.

Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund(the Funds) are funds of Fidelity Devonshire Trust(the trust) and are authorized to issue an unlimited number of shares. The trust is registered under the Investment Company Act of 1940, as amended(the 1940 Act), as an open-end management company organized as a Massachusetts business trust. The Funds offer Class A, Class T, Class B, Class C, Institutional Class and Large Cap Value, Mid Cap Value, Large Cap Growth and Mid Cap Growth shares, respectively, each of which has equal rights as to assets and voting privileges. Each class has exclusive voting rights with respect to matters that affect that class. The Funds commenced sale of Class A, Class T, Class B, Class C and Institutional Class shares and the existing class of each Fund was designated Large Cap Value, Mid Cap Value, Large Cap Growth and Mid Cap Growth, respectively, on February 13, 2007. Class B shares will automatically convert to Class A shares after a holding period of seven years from the initial date of purchase. Investment income, realized and unrealized capital gains and losses, the common expenses of the Fund, and certain fund-level expense reductions, if any, are allocated on a pro rata basis to each class based on the relative net assets of each class to the total net assets of the Fund. Each class differs with respect to transfer agent and distribution and service plan fees incurred. Certain expense reductions also differ by class. Certain Funds investments in emerging markets can be subject to social, economic, regulatory, and political uncertainties and can be extremely volatile.

2. Investments in Fidelity Central Funds.

The Funds may invest in Fidelity Central Funds, which are open-end investment companies available only to other investment companies and accounts managed by Fidelity Management & Research Company(FMR) and its affiliates. The Funds' Schedules of Investments list each of the Fidelity Central Funds as an investment of each Fund, but does not include the underlying holdings of each Fidelity Central Fund. As an Investing Fund, the Funds indirectly bear their proportionate share of the expenses of the underlying Fidelity Central Funds.

The Money Market Central Funds seek preservation of capital and current income and are managed by Fidelity Investments Money Management, Inc.(FIMM), an affiliate of FMR.

A complete list of holdings for each Fidelity Central Fund is available upon request or at the SEC's web site at www.sec.gov. In addition, the financial statements of the Fidelity Central Funds, which are not covered by the Funds' Report of Independent Registered Public Accounting Firm, are available on the SEC's web site or upon request.

3. Significant Accounting Policies.

The financial statements have been prepared in conformity with accounting principles generally accepted in the United States of America, which require management to make certain estimates and assumptions at the date of the financial statements. Actual results could differ from those estimates. The following summarizes the significant accounting policies of the Funds:

Security Valuation. Investments are valued and net asset value(NAV) per share is calculated(NAV calculation) as of the close of business of the New York Stock Exchange(NYSE), normally 4:00 p.m. Eastern time. Wherever possible, each Fund uses independent pricing services approved by the Board of Trustees to value their investments.

Equity securities, including restricted securities, for which market quotations are readily available, are valued at the last reported sale price or official closing price as reported by an independent pricing service on the primary market or exchange on which they are traded. In the event there were no sales during the day or closing prices are not available, securities are valued at the last quoted bid price. Investments in open-end mutual funds, including the Fidelity Central Funds, are valued at their closing net asset value each business day. Short-term securities with remaining maturities of sixty days or less for which quotations are not readily available are valued at amortized cost, which approximates value.

When current market prices or quotations are not readily available or do not accurately reflect fair value, valuations may be determined in accordance with procedures adopted by the Board of Trustees. For example, when developments occur between the close of a market and the close of the NYSE that may materially affect the value of some or all of the securities, or when trading in a security is halted, those securities may be fair valued. Factors used in the determination of fair value may include monitoring news to identify significant market or security specific events such as changes in the value of U.S. securities markets, reviewing developments in foreign markets and evaluating the performance of ADRs, futures contracts and exchange-traded funds. Because each Fund's utilization of fair value pricing depends on market activity, the frequency with which fair value pricing is used cannot be predicted and may be utilized to a significant extent. The value of securities used for NAV calculation under fair value pricing may differ from published prices for the same securities.

Foreign Currency. Certain Funds may use foreign currency contracts to facilitate transactions in foreign-denominated securities. Losses from these transactions may arise from changes in the value of the foreign currency or if the counterparties do not perform under the contracts' terms.

Annual Report

Notes to Financial Statements - continued

3. Significant Accounting Policies - continued

Foreign Currency - continued

Foreign-denominated assets, including investment securities, and liabilities are translated into U.S. dollars at the exchange rate at period end. Purchases and sales of investment securities, income and dividends received and expenses denominated in foreign currencies are translated into U.S. dollars at the exchange rate in effect on the transaction date.

The effects of exchange rate fluctuations on investments are included with the net realized and unrealized gain(loss) on investment securities. Other foreign currency transactions resulting in realized and unrealized gain(loss) are disclosed separately.

Investment Transactions and Income. For financial reporting purposes, the Funds' investment holdings and NAV include trades executed through the end of the last business day of the period. The NAV for processing shareholder transactions includes trades executed through the end of the prior business day. Gains and losses on securities sold are determined on the basis of identified cost and may include proceeds received from litigation. Dividend income is recorded on the ex-dividend date, except for certain dividends from foreign securities where the ex-dividend date may have passed, which are recorded as soon as the Funds are informed of the ex-dividend date. Non-cash dividends included in dividend income, if any, are recorded at the fair market value of the securities received. Distributions received on securities that represent a return of capital or capital gain are recorded as a reduction of cost of investments and/or as a realized gain. The Funds estimate the components of distributions received that may be considered return of capital distributions or capital gain distributions. Interest income and distributions from the Fidelity Central Funds are accrued as earned. Interest income includes coupon interest and amortization of premium and accretion of discount on debt securities.

Expenses. Most expenses of the trust can be directly attributed to a fund. Expenses which cannot be directly attributed are apportioned among each Fund in the trust. Expense estimates are accrued in the period to which they relate and adjustments are made when actual amounts are known.

Income Tax Information and Distributions to Shareholders. Each year, each Fund intends to qualify as a regulated investment company by distributing substantially all of its taxable income and realized gains under Subchapter M of the Internal Revenue Code and filing its U.S. federal tax return. As a result, no provision for income taxes is required. Each Fund adopted the provisions of FASB Interpretation No. 48, Accounting for Uncertainties in Income Taxes(FIN 48), on June 29, 2007. FIN 48 sets forth a minimum threshold for financial statement recognition of the benefit of a tax position taken or expected to be taken in a tax return. The implementation of FIN 48 did not result in any unrecognized tax benefits in the accompanying financial statements. Each of the Funds' federal tax returns for the prior three fiscal years remains subject to examination by the Internal Revenue Service. Foreign taxes are provided for based on each Fund's understanding of the tax rules and rates that exist in the foreign markets in which it invests.

Distributions are recorded on the ex-dividend date. Income dividends and capital gain distributions are declared separately for each class. Income and capital gain distributions are determined in accordance with income tax regulations, which may differ from generally accepted accounting principles. In addition, certain Funds will claim a portion of the payment made to redeeming shareholders as a distribution for income tax purposes.

Capital accounts within the financial statements are adjusted for permanent book-tax differences. These adjustments have no impact on net assets or the results of operations. Temporary book-tax differences will reverse in a subsequent period.

Book-tax differences are primarily due to foreign currency transactions, passive foreign investment companies (PFIC), partnerships, capital loss carryforwards and losses deferred due to wash sales and excise tax regulations.

The tax-basis components of distributable earnings and the federal tax cost as of period end were as follows for each Fund:

Cost for Federal
Income Tax
Purposes

Unrealized
Appreciation

Unrealized
Depreciation

Net Unrealized
Appreciation/
(Depreciation)

Fidelity Large Cap Value Fund

$ 1,571,171,429

$ 98,475,075

$ (92,166,547)

$ 6,308,528

Fidelity Mid Cap Value Fund

844,773,929

42,574,804

(57,370,158)

(14,795,354)

Fidelity Large Cap Growth Fund

166,610,940

4,786,892

(10,368,163)

(5,581,271)

Fidelity Mid Cap Growth Fund

319,883,846

21,520,033

(22,775,536)

(1,255,503)

Annual Report

3. Significant Accounting Policies - continued

Income Tax Information and Distributions to Shareholders - continued

The tax character of distributions paid was as follows:

January 31, 2008

Ordinary
Income

Long-term
Capital Gains

Total

Fidelity Large Cap Value Fund

$ 37,304,898

$ 67,715,379

$ 105,020,277

Fidelity Mid Cap Value Fund

10,958,373

29,219,096

40,177,469

Fidelity Large Cap Growth Fund

6,193,151

12,379,253

18,572,404

Fidelity Mid Cap Growth Fund

-

21,909,497

21,909,497

January 31, 2007

Ordinary Income

Long-term
Capital Gains

Total

Fidelity Large Cap Value Fund

$ 17,006,124

$ 7,416,999

$ 24,423,123

Fidelity Mid Cap Value Fund

7,722,049

9,225,041

16,947,090

Fidelity Large Cap Growth Fund

1,920,898

1,860,806

3,781,704

Fidelity Mid Cap Growth Fund

2,544,632

2,544,632

5,089,264

Short-Term Trading (Redemption) Fees. Shares held in the Fidelity Mid Cap Value Fund and Fidelity Mid Cap Growth Fund less than 30 days are subject to a redemption fee equal to 0.75% of the proceeds of the redeemed shares. All redemption fees, including any estimated redemption fees paid by FMR, are retained by the Funds and accounted for as an addition to paid in capital.

New Accounting Pronouncement. In September 2006, Statement of Financial Accounting Standards No. 157, Fair Value Measurements(SFAS 157), was issued and is effective for fiscal years beginning after November 15, 2007. SFAS 157 defines fair value, establishes a framework for measuring fair value and expands disclosures about fair value measurements. Management is currently evaluating the impact the adoption of SFAS 157 will have on the Funds' financial statement disclosures.

4. Operating Policies.

Repurchase Agreements. FMR has received an Exemptive Order from the Securities and Exchange Commission(the SEC) which permits certain Funds and other affiliated entities of FMR to transfer uninvested cash balances into joint trading accounts which are then invested in repurchase agreements. Certain Funds may also invest directly with institutions in repurchase agreements. Repurchase agreements are collateralized by government or non-government securities. Upon settlement date, collateral is held in segregated accounts with custodian banks and may be obtained in the event of a default of the counterparty. Each applicable Fund monitors, on a daily basis, the value of the collateral to ensure it is at least equal to the principal amount of the repurchase agreement(including accrued interest). In the event of a default by the counterparty, realization of the collateral proceeds could be delayed, during which time the value of the collateral may decline.

5. Purchases and Sales of Investments.

Purchases and sales of securities, other than short-term securities, are noted in the table below.

Purchases($)

Sales($)

Fidelity Large Cap Value Fund

3,485,833,978

3,246,788,832

Fidelity Mid Cap Value Fund

2,520,812,788

2,356,094,981

Fidelity Large Cap Growth Fund

745,952,270

764,990,241

Fidelity Mid Cap Growth Fund

995,464,036

1,090,139,044

6. Fees and Other Transactions with Affiliates.

Management Fee. FMR and its affiliates provide the Funds with investment management related services for which the Funds pay a monthly management fee. The management fee is the sum of an individual fund fee rate and a group fee rate. The individual fund fee rate is applied to each Fund's average net assets. The group fee rate is based upon the average net assets of all the mutual funds advised by FMR. The group fee rate decreases as assets under management increase and increases as assets under management decrease. In addition, the management fee for the funds is subject to a performance adjustment(up to a maximum ± .20% of each applicable Fund's average net assets over a 36 month performance period). The upward or downward adjustment to the management fee is based on each applicable Fund's relative investment performance of the asset-weighted return of all

Annual Report

Notes to Financial Statements - continued

6. Fees and Other Transactions with Affiliates - continued

Management Fee - continued

classes as compared to an appropriate benchmark index. For the period, each Fund's annual management fee rate expressed as a percentage of each Fund's average net assets, including the performance adjustment, if applicable was as follows:

Individual Rate

Group Rate

Total

Fidelity Large Cap Value Fund

.30%

.26%

.58%

Fidelity Mid Cap Value Fund

.30%

.26%

.52%

Fidelity Large Cap Growth Fund

.30%

.26%

.58%

Fidelity Mid Cap Growth Fund

.30%

.26%

.46%

Distribution and Service Plan. In accordance with Rule 12b-1 of the 1940 Act, the Fund has adopted separate Distribution and Service Plans for each class of shares. Certain classes pay Fidelity Distributors Corporation(FDC), an affiliate of FMR, separate Distribution and Service Fees, each of which is based on an annual percentage of each class' average net assets. In addition, FDC may pay financial intermediaries for selling shares of the Fund and providing shareholder support services. For the period, the Distribution and Service Fee rates and the total amounts paid to and retained by FDC were as follows:

Fidelity Large Cap Value Fund

Distribution
Fee

Service
Fee

Paid to
FDC

Retained
by FDC

Class A

0%

.25%

$ 14,716

$ 3,419

Class T

.25%

.25%

29,520

209

Class B

.75%

.25%

20,952

15,912

Class C

.75%

.25%

8,540

4,769

$ 73,728

$ 24,309

Fidelity Mid Cap Value Fund

Class A

0%

.25%

$ 9,176

$ 3,141

Class T

.25%

.25%

10,994

355

Class B

.75%

.25%

17,435

13,285

Class C

.75%

.25%

16,086

12,118

$ 53,691

$ 28,899

Fidelity Large Cap Growth Fund

Class A

0%

.25%

$ 1,502

$ 246

Class T

.25%

.25%

2,248

460

Class B

.75%

.25%

2,797

2,331

Class C

.75%

.25%

4,188

3,019

$ 10,735

$ 6,056

Fidelity Mid Cap Growth Fund

Class A

0%

.25%

$ 1,756

$ 245

Class T

.25%

.25%

1,796

463

Class B

.75%

.25%

2,959

2,452

Class C

.75%

.25%

3,852

3,025

$ 10,363

$ 6,185

Sales Load. FDC receives a front-end sales charge of up to 5.75% for selling Class A shares, and 3.50% for selling Class T shares, some of which is paid to financial intermediaries for selling shares of the Fund. FDC receives the proceeds of contingent deferred sales charges levied on Class A, Class T, Class B, and Class C redemptions. These charges depend on the holding period. The deferred sales charges range from 5% to 1% for Class B, 1% for Class C, 1.00% to .50% for certain purchases of Class A shares and .25% for certain purchases of Class T shares.

For the period, sales charge amounts retained by FDC were as follows:

Fidelity Large Cap Value Fund

Retained
by FDC

Class A

$ 18,265

Class T

4,391

Class B *

1,096

Class C *

11

$ 22,763

Annual Report

6. Fees and Other Transactions with Affiliates - continued

Sales Load - continued

For the period, sales charge amounts retained by FDC were as follows: - continued

Fidelity Mid Cap Value Fund

Retained
by FDC

Class A

$ 22,621

Class T

5,951

Class B*

2,448

Class C*

137

$ 31,157

Fidelity Large Cap Growth Fund

Class A

$ 4,515

Class T

1,123

Class B*

509

Class C*

62

$ 6,209

Fidelity Mid Cap Growth Fund

Class A

$ 6,603

Class T

1,049

Class B*

962

Class C*

165

$ 8,779

* When Class B and Class C shares are initially sold, FDC pays commissions from its own resources to financial intermediaries through which the sales are made.

Transfer Agent Fees. Fidelity Investments Institutional Operations Company, Inc.(FIIOC), an affiliate of FMR, is the transfer, dividend disbursing and shareholder servicing agent for each class of the Funds. FIIOC receives account fees and asset-based fees that vary according to the account size and type of account of the shareholders of the respective classes of each Fund. FIIOC pays for typesetting, printing and mailing of shareholder reports, except proxy statements. Prior to January 1, 2008, Fidelity Service Company, Inc. (FSC), also an affiliate of FMR, was the Fund's transfer agent. For the period, the total transfer agent fees paid by each class to FIIOC or FSC, were as follows:

Fidelity Large Cap Value Fund

Amount

% of
Average
Net Assets
*

Class A

$ 20,036

.34

Class T

20,697

.35

Class B

7,804

.37

Class C

2,694

.31

Large Cap Value

3,716,520

.23

Institutional Class

944

.23

$ 3,768,695

Fidelity Mid Cap Value Fund

Class A

$ 10,910

.30

Class T

6,730

.31

Class B

5,278

.30

Class C

5,017

.31

Mid Cap Value

2,206,272

.24

Institutional Class

1,803

.30

$ 2,236,010

Fidelity Large Cap Growth Fund

Class A

$ 1,787

.30

Class T

1,353

.31

Class B

833

.30

Class C

1,224

.29

Large Cap Growth

529,155

.31

Institutional Class

416

.20

$ 534,768

Annual Report

Notes to Financial Statements - continued

6. Fees and Other Transactions with Affiliates - continued

Transfer Agent Fees - continued

Fidelity Mid Cap Growth Fund

Amount

% of
Average
Net Assets
*

Class A

$ 2,094

.30

Class T

1,136

.32

Class B

899

.30

Class C

1,151

.30

Mid Cap Growth

1,196,809

.29

Institutional Class

225

.17

$ 1,202,314

* Annualized

Accounting and Security Lending Fees. Fidelity Service Company, Inc.(FSC), an affiliate of FMR, maintains each Fund's accounting records. The accounting fee is based on the level of average net assets for the month. Under a separate contract, FSC administers the security lending program. The security lending fee is based on the number and duration of lending transactions.

Brokerage Commissions. Certain Funds placed a portion of their portfolio transactions with brokerage firms which are affiliates of the investment adviser. The commissions paid to these affiliated firms were as follows:

Amount

Fidelity Large Cap Value Fund

$ 18,780

Fidelity Mid Cap Value Fund

12,752

Fidelity Large Cap Growth Fund

13,318

Fidelity Mid Cap Growth Fund

10,529

Interfund Lending Program. Pursuant to an Exemptive Order issued by the SEC, the Funds, along with other registered investment companies having management contracts with FMR, may participate in an interfund lending program. This program provides an alternative credit facility allowing the funds to borrow from, or lend money to, other participating affiliated funds. At period end, there were no interfund loans outstanding. Each applicable fund's activity in this program during the period for which loans were outstanding was as follows:

Borrower or
Lender

Average Daily Loan Balance

Weighted
Average Interest Rate

Interest
Expense

Fidelity Large Cap Value Fund

Borrower

$ 6,183,857

4.35%

$ 5,226

Fidelity Mid Cap Value Fund

Borrower

$ 5,637,000

5.27%

$ 4,946

7. Committed Line of Credit.

Certain Funds participate with other funds managed by FMR in a $4.2 billion credit facility(the "line of credit") to be utilized for temporary or emergency purposes to fund shareholder redemptions or for other short-term liquidity purposes. The participating funds have agreed to pay commitment fees on their pro rata portion of the line of credit, which is reflected in Miscellaneous Expense on the Statement of Operations, and is as follows:

Fidelity Large Cap Value Fund

$ 3,040

Fidelity Mid Cap Value Fund

1,696

Fidelity Large Cap Growth Fund

343

Fidelity Mid Cap Growth Fund

824

During the period, there were no borrowings on this line of credit.

8. Security Lending.

Certain Funds lend portfolio securities from time to time in order to earn additional income. On the settlement date of the loan, each applicable Fund receives collateral(in the form of U.S. Treasury obligations, letters of credit and/or cash) against the loaned securities and maintains collateral in an amount not less than 100% of the market value of the loaned securities during the period of the loan. The market value of the loaned securities is determined at the close of business of the Funds and any additional required collateral is delivered to the Funds on the next business day. If the borrower defaults on its obligation to return the securities loaned because of insolvency or other reasons, a fund could experience delays and costs in recovering the securities loaned or in gaining access to the collateral. Any cash collateral received is invested in the Fidelity Securities Lending Cash

Annual Report

8. Security Lending - continued

Central Fund. The value of loaned securities and cash collateral at period end are disclosed on each applicable Fund's Statement of Assets and Liabilities. Security lending income represents the income earned on investing cash collateral, less fees and expenses associated with the loan, plus any premium payments that may be received on the loan of certain types of securities. Security lending income is presented on each applicable Fund's Statement of Operations as a component of income from Fidelity Central Funds.

9. Expense Reductions.

FMR voluntarily agreed to reimburse funds to the extent annual operating expenses exceeded certain levels of average net assets as noted in the table below. Some expenses, for example interest expense, including commitment fees, are excluded from this reimbursement.

The following classes of each applicable Fund were in reimbursement during the period:

Expense
Limitations

Reimbursement
from adviser

Fidelity Large Cap Growth Fund

Large Cap Growth

1.00%

$ 58,552

In addition, FMR voluntarily agreed to reimburse a portion of certain fund's existing class' operating expenses. During the period, this reimbursement reduced certain fund's existing class' expenses by the following amounts:

Reimbursement
from adviser

Fidelity Large Cap Value Fund

Large Cap Value

$ 85,974

Fidelity Mid Cap Value Fund

Mid Cap Value

$ 85,078

Fidelity Large Cap Growth Fund

Large Cap Growth

$ 25,900

Fidelity Mid Cap Growth Fund

Mid Cap Growth

$ 84,552

In addition, through arrangements with each applicable Fund's custodian and each class' transfer agent, credits realized as a result of uninvested cash balances were used to reduce the Fund's expenses. All of the applicable expense reductions are noted in the table below.

Brokerage Service Arrangements

Transfer
Agent
expense
reduction

Fidelity Large Cap Value Fund

$ -

$ -

Large Cap Value

-

50,587

Institutional Class

-

50

Fidelity Mid Cap Value Fund

-

-

Class A

-

31

Class B

-

9

Mid Cap Value

-

19,810

Institutional Class

-

70

Fidelity Large Cap Growth Fund

-

-

Large Cap Growth

-

13,456

Fidelity Mid Cap Growth Fund

1,543

-

Mid Cap Growth

-

27,293

10. Other.

The Funds' organizational documents provide former and current trustees and officers with a limited indemnification against liabilities arising in connection with the performance of their duties to the Funds. In the normal course of business, the Funds may also enter into contracts that provide general indemnifications. The Funds' maximum exposure under these arrangements is unknown as this would be dependent on future claims that may be made against the Funds. The risk of material loss from such claims is considered remote.

Annual Report

Notes to Financial Statements - continued

10. Other - continued

In December 2006, the Independent Trustees, with the assistance of independent counsel, completed an investigation regarding gifts, gratuities and business entertainment provided by certain brokers to certain individuals who were employed on FMR's domestic equity trading desk during the period 2002 to 2004. The Independent Trustees and FMR agreed that, despite the absence of proof that the Fidelity mutual funds experienced diminished execution quality as a result of the improper receipt of gifts and business entertainment, the conduct at issue was serious and was worthy of redress. Accordingly, the Independent Trustees requested, and FMR agreed to make a payment of $42 million plus accrued interest, which equaled approximately $7.3 million, to certain Fidelity mutual funds. FMR also reimbursed the related legal expenses, which are recorded in the accompanying Statement of Operations as an expense reduction.

Subsequent to period end, the Trustees approved a method for allocating this payment among the funds and, in total, FMR paid each fund the following amounts:

Fidelity Large Cap Value Fund

$ 12,127

Fidelity Mid Cap Value Fund

$ 23,107

Fidelity Large Cap Growth Fund

$ 14,058

Fidelity Mid Cap Growth Fund

$ 16,394

In a related administrative order dated March 5, 2008, the U.S. Securities and Exchange Commission ("SEC") announced a settlement with FMR and FMR Co., Inc. (an affiliate of FMR) involving the SEC's regulatory rules for investment advisers and the improper receipt of gifts, gratuities and business entertainment. Without admitting or denying the SEC's findings, FMR agreed to pay an $8 million civil penalty to the United States Treasury.

11. Distributions to Shareholders.

Distributions to shareholders of each class were as follows:

Years ended January 31,

2008 A

2007

Fidelity Large Cap Value Fund

From net investment income

Class A

$ 75,483

$ -

Class T

44,031

-

Class B

824

-

Class C

4,115

-

Large Cap Value

14,152,017

10,401,399

Institutional Class

10,129

-

Total

$ 14,286,599

$ 10,401,399

From net realized gain

Class A

$ 372,920

$ -

Class T

317,190

-

Class B

97,442

-

Class C

51,607

-

Large Cap Value

89,854,195

14,021,724

Institutional Class

40,324

-

Total

$ 90,733,678

$ 14,021,724

Fidelity Mid Cap Value Fund

From net investment income

Class A

$ 18,691

$ -

Class T

5,836

-

Class B

259

-

Class C

66

-

Mid Cap Value

3,005,000

2,949,768

Institutional Class

7,163

-

Total

$ 3,037,015

$ 2,949,768

From net realized gain

Class A

$ 176,774

$ -

Class T

106,263

-

Class B

46,244

-

Class C

67,330

-

Mid Cap Value

36,700,729

13,997,322

Institutional Class

43,114

-

Total

$ 37,140,454

$ 13,997,322

Annual Report

11. Distributions to Shareholders - continued

Distributions to shareholders of each class were as follows: - continued

Years ended January 31,

2008 A

2007

Fidelity Large Cap Growth Fund

From net investment income

Large Cap Growth

$ -

$ 137,207

From net realized gain

Class A

$ 123,133

$ -

Class T

101,048

-

Class B

40,430

-

Class C

75,734

-

Large Cap Growth

18,196,705

3,644,497

Institutional Class

35,354

-

Total

$ 18,572,404

$ 3,644,497

Fidelity Mid Cap Growth Fund

From net realized gain

Class A

$ 100,774

$ -

Class T

34,957

-

Class B

25,358

-

Class C

40,118

-

Mid Cap Growth

21,697,261

5,089,264

Institutional Class

11,029

-

Total

$ 21,909,497

$ 5,089,264

A Distributions for Class A,T,B,C and Institutional are for the period February 13, 2007(commencement of sale of shares) to January 31, 2008.

12. Share Transactions.

Transactions for each class of shares were as follows:

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Fidelity Large Cap Value Fund

Class A

Shares sold

1,045,290

-

$ 16,232,942

$ -

Reinvestment of distributions

29,393

-

423,574

-

Shares redeemed

(352,749)

-

(5,365,826)

-

Net increase (decrease)

721,934

-

$ 11,290,690

$ -

Class T

Shares sold

935,752

-

$ 14,740,653

$ -

Reinvestment of distributions

24,935

-

359,076

-

Shares redeemed

(518,994)

-

(7,932,251)

-

Net increase (decrease)

441,693

-

$ 7,167,478

$ -

Class B

Shares sold

315,386

-

$ 4,951,665

$ -

Reinvestment of distributions

6,739

-

97,191

-

Shares redeemed

(184,721)

-

(2,835,906)

-

Net increase (decrease)

137,404

-

$ 2,212,950

$ -

Class C

Shares sold

138,185

-

$ 2,136,503

$ -

Reinvestment of distributions

3,506

-

50,496

-

Shares redeemed

(52,323)

-

(788,391)

-

Net increase (decrease)

89,368

-

$ 1,398,608

$ -

Annual Report

Notes to Financial Statements - continued

12. Share Transactions - continued

Transactions for each class of shares were as follows: - continued

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Fidelity Large Cap Value Fund - continued

Large Cap Value

Shares sold

52,779,386

72,550,724

$ 813,954,360

$ 1,029,577,278

Reinvestment of distributions

7,056,478

1,672,345

102,030,422

23,955,243

Shares redeemed

(40,898,915)

(25,616,795)

(619,081,485)

(368,491,524)

Net increase (decrease)

18,936,949

48,606,274

$ 296,903,297

$ 685,040,997

Institutional Class

Shares sold

92,735

-

$ 1,397,015

$ -

Reinvestment of distributions

3,477

-

50,127

-

Shares redeemed

(17,951)

-

(257,652)

-

Net increase (decrease)

78,261

-

$ 1,189,490

$ -

Fidelity Mid Cap Value Fund

Class A

Shares sold

615,995

-

$ 10,540,942

$ -

Reinvestment of distributions

11,024

-

178,558

-

Shares redeemed

(132,358)

-

(2,195,988)

-

Net increase (decrease)

494,661

-

$ 8,523,512

$ -

Class T

Shares sold

325,063

-

$ 5,643,056

$ -

Reinvestment of distributions

6,694

-

108,412

-

Shares redeemed

(84,728)

-

(1,422,111)

-

Net increase (decrease)

247,029

-

$ 4,329,357

$ -

Class B

Shares sold

204,574

-

$ 3,623,364

$ -

Reinvestment of distributions

2,791

-

45,364

-

Shares redeemed

(120,358)

-

(2,068,797)

-

Net increase (decrease)

87,007

-

$ 1,599,931

$ -

Class C

Shares sold

168,571

-

$ 2,987,730

$ -

Reinvestment of distributions

3,136

-

50,659

-

Shares redeemed

(61,027)

-

(991,710)

-

Net increase (decrease)

110,680

-

$ 2,046,679

$ -

Mid Cap Value

Shares sold

34,123,012

26,022,900

$ 603,742,926

$ 415,931,492

Reinvestment of distributions

2,338,406

1,003,867

38,409,538

16,337,499

Shares redeemed

(27,133,004)

(10,884,934)

(462,055,715)

(173,050,014)

Net increase (decrease)

9,328,414

16,141,833

$ 180,096,749

$ 259,218,977

Institutional Class

Shares sold

102,472

-

$ 1,790,525

$ -

Reinvestment of distributions

3,101

-

50,277

-

Shares redeemed

(9,162)

-

(148,581)

-

Net increase (decrease)

96,411

-

$ 1,692,221

$ -

Fidelity Large Cap Growth Fund

Class A

Shares sold

165,474

-

$ 1,900,337

$ -

Reinvestment of distributions

10,979

-

116,895

-

Shares redeemed

(44,281)

-

(486,246)

-

Net increase (decrease)

132,172

-

$ 1,530,986

$ -

Class T

Shares sold

167,227

-

$ 1,920,600

$ -

Reinvestment of distributions

9,490

-

101,048

-

Shares redeemed

(65,269)

-

(790,917)

-

Net increase (decrease)

111,448

-

$ 1,230,731

$ -

Annual Report

12. Share Transactions - continued

Transactions for each class of shares were as follows: - continued

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Class B

Shares sold

55,847

-

$ 636,287

$ -

Reinvestment of distributions

3,795

-

40,430

-

Shares redeemed

(4,405)

-

(49,003)

-

Net increase (decrease)

55,237

-

$ 627,714

$ -

Class C

Shares sold

106,310

-

$ 1,230,146

$ -

Reinvestment of distributions

6,439

-

68,402

-

Shares redeemed

(16,516)

-

(191,737)

-

Net increase (decrease)

96,233

-

$ 1,106,811

$ -

Large Cap Growth

Shares sold

5,794,783

10,973,541

$ 66,845,422

$ 124,315,764

Reinvestment of distributions

1,665,396

318,299

17,914,862

3,701,456

Shares redeemed

(7,903,937)

(9,222,108)

(90,877,571)

(102,538,999)

Net increase (decrease)

(443,758)

2,069,732

$ (6,117,287)

$ 25,478,221

Institutional Class

Shares sold

38,550

-

$ 453,557

$ -

Reinvestment of distributions

3,006

-

32,121

-

Shares redeemed

(2,488)

-

(28,969)

-

Net increase (decrease)

39,068

-

$ 456,709

$ -

Fidelity Mid Cap Growth Fund

Class A

Shares sold

165,234

-

$ 2,326,735

$ -

Reinvestment of distributions

7,218

-

97,305

-

Shares redeemed

(13,557)

-

(186,849)

-

Net increase (decrease)

158,895

-

$ 2,237,191

$ -

Class T

Shares sold

54,813

-

$ 796,065

$ -

Reinvestment of distributions

2,601

-

34,957

-

Shares redeemed

(8,726)

-

(121,838)

-

Net increase (decrease)

48,688

-

$ 709,184

$ -

Class B

Shares sold

35,917

-

$ 516,488

$ -

Reinvestment of distributions

1,882

-

25,182

-

Shares redeemed

(3,583)

-

(47,283)

-

Net increase (decrease)

34,216

-

$ 494,387

$ -

Class C

Shares sold

59,795

-

$ 849,458

$ -

Reinvestment of distributions

2,811

-

37,643

-

Shares redeemed

(4,934)

-

(66,440)

-

Net increase (decrease)

57,672

-

$ 820,661

$ -

Mid Cap Growth

Shares sold

7,994,956

24,363,804

$ 114,388,509

$ 336,430,868

Reinvestment of distributions

1,580,341

357,726

21,334,599

4,986,704

Shares redeemed

(15,751,578)

(18,209,467)

(217,016,915)

(243,991,431)

Net increase (decrease)

(6,176,281)

6,512,063

$ (81,293,807)

$ 97,426,141

Institutional Class

Shares sold

14,080

-

$ 199,943

$ -

Reinvestment of distributions

816

-

11,029

-

Net increase (decrease)

14,896

-

$ 210,972

$ -

A Share transactions for class A,T,B,C and Institutional are for the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

Annual Report

Report of Independent Registered

Public Accounting Firm

To the Trustees of Fidelity Devonshire Trust and the Shareholders of Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund:

In our opinion, the accompanying statements of assets and liabilities, including the schedules of investments, and the related statements of operations and of changes in net assets and the financial highlights present fairly, in all material respects, the financial position of Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund (funds of Fidelity Devonshire Trust) at January 31, 2008, the results of each of their operations for the year then ended, the changes in each of their net assets for each of the two years in the period then ended and the financial highlights for each of the periods indicated, in conformity with accounting principles generally accepted in the United States of America. These financial statements and financial highlights (hereafter referred to as "financial statements") are the responsibility of the Fidelity Devonshire Trust's management. Our responsibility is to express an opinion on these financial statements based on our audits. We conducted our audits of these financial statements in accordance with the standards of the Public Company Accounting Oversight Board (United States). Those standards require that we plan and perform the audit to obtain reasonable assurance about whether the financial statements are free of material misstatement. An audit includes examining, on a test basis, evidence supporting the amounts and disclosures in the financial statements, assessing the accounting principles used and significant estimates made by management, and evaluating the overall financial statement presentation. We believe that our audits, which included confirmation of securities at January 31, 2008 by correspondence with the custodian and brokers, provide a reasonable basis for our opinion.

/s/ PricewaterhouseCoopers LLP

PricewaterhouseCoopers LLP

Boston, Massachusetts
March 24, 2008

Annual Report

Trustees and Officers

The Trustees, Members of the Advisory Board, and executive officers of the trust and funds, as applicable, are listed below. The Board of Trustees governs each fund and is responsible for protecting the interests of shareholders. The Trustees are experienced executives who meet periodically throughout the year to oversee each fund's activities, review contractual arrangements with companies that provide services to each fund, and review each fund's performance. Except for James C. Curvey, each of the Trustees oversees 374 funds advised by FMR or an affiliate. Mr. Curvey oversees 369 funds advised by FMR or an affiliate.

The Trustees hold office without limit in time except that (a) any Trustee may resign; (b) any Trustee may be removed by written instrument, signed by at least two-thirds of the number of Trustees prior to such removal; (c) any Trustee who requests to be retired or who has become incapacitated by illness or injury may be retired by written instrument signed by a majority of the other Trustees; and (d) any Trustee may be removed at any special meeting of shareholders by a two-thirds vote of the outstanding voting securities of the trust. Each Trustee who is not an interested person (as defined in the 1940 Act) (Independent Trustee), shall retire not later than the last day of the calendar year in which his or her 72nd birthday occurs. The Independent Trustees may waive this mandatory retirement age policy with respect to individual Trustees. The executive officers and Advisory Board Members hold office without limit in time, except that any officer and Advisory Board Member may resign or may be removed by a vote of a majority of the Trustees at any regular meeting or any special meeting of the Trustees. Except as indicated, each individual has held the office shown or other offices in the same company for the past five years.

The funds' Statement of Additional Information (SAI) includes more information about the Trustees. To request a free copy, call Fidelity at 1-800-544-8544.

Interested Trustees*:

Correspondence intended for each Trustee who is an interested person may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Edward C. Johnson 3d (77)

Year of Election or Appointment: 1985

Mr. Johnson is Chairman of the Board of Trustees. Mr. Johnson serves as President (2006-present), Chief Executive Officer, Chairman, and a Director of FMR LLC; Chairman and a Director of FMR; Chairman and a Director of Fidelity Research & Analysis Company (FRAC); Chairman and a Director of Fidelity Investments Money Management, Inc.; and Chairman (2001-present) and a Director of FMR Co., Inc. In addition, Mr. Johnson serves as Chairman and Director of FIL Limited. Mr. Edward C. Johnson 3d and Mr. Arthur E. Johnson are not related.

James C. Curvey (72)

Year of Election or Appointment: 2007

Mr. Curvey also serves as Trustee (2007-present) or Member of the Advisory Board (2007-present) of other investment companies advised by FMR. Mr. Curvey is a Director of FMR and FMR Co., Inc. (2007-present). Mr. Curvey is also Vice Chairman (2006-present) and Director of FMR LLC. Mr. Curvey joined Fidelity in 1982 and served in numerous senior management positions, including President and Chief Operating Officer of FMR LLC (1997-2000) and President of Fidelity Strategic Investments (2000-2002). In addition, he serves as a member of the Board of Directors of Geerlings & Wade, Inc. (wine distribution).

* Trustees have been determined to be "Interested Trustees" by virtue of, among other things, their affiliation with the trust or various entities under common control with FMR. FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Independent Trustees:

Correspondence intended for each Independent Trustee (that is, the Trustees other than the Interested Trustees) may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235.

Name, Age; Principal Occupation

Dennis J. Dirks (59)

Year of Election or Appointment: 2005

Prior to his retirement in May 2003, Mr. Dirks was Chief Operating Officer and a member of the Board of The Depository Trust & Clearing Corporation (DTCC) (1999-2003). He also served as President, Chief Operating Officer, and Board member of The Depository Trust Company (DTC) (1999-2003) and President and Board member of the National Securities Clearing Corporation (NSCC) (1999-2003). In addition, Mr. Dirks served as Chief Executive Officer and Board member of the Government Securities Clearing Corporation (2001-2003) and Chief Executive Officer and Board member of the Mortgage-Backed Securities Clearing Corporation (2001-2003). Mr. Dirks also serves as a Trustee and a member of the Finance Committee of Manhattan College (2005-present) and a Trustee and a member of the Finance Committee of AHRC of Nassau County (2006-present).

Albert R. Gamper, Jr. (65)

Year of Election or Appointment: 2006

Prior to his retirement in December 2004, Mr. Gamper served as Chairman of the Board of CIT Group Inc. (commercial finance). During his tenure with CIT Group Inc. Mr. Gamper served in numerous senior management positions, including Chairman (1987-1989; 1999-2001; 2002-2004), Chief Executive Officer (1987-2004), and President (1989-2002). He currently serves as a member of the Board of Directors of Public Service Enterprise Group (utilities, 2001-present), Chairman of the Board of Governors, Rutgers University (2004-present), and Chairman of the Board of Saint Barnabas Health Care System.

George H. Heilmeier (71)

Year of Election or Appointment: 2004

Dr. Heilmeier is Chairman Emeritus of Telcordia Technologies (communication software and systems), where prior to his retirement, he served as company Chairman and Chief Executive Officer. He currently serves on the Boards of Directors of The Mitre Corporation (systems engineering and information technology support for the government), and HRL Laboratories (private research and development, 2004-present). He is Chairman of the General Motors Science & Technology Advisory Board and a Life Fellow of the Institute of Electrical and Electronics Engineers (IEEE). Dr. Heilmeier is a member of the Defense Science Board and the National Security Agency Advisory Board. He is also a member of the National Academy of Engineering, the American Academy of Arts and Sciences, and the Board of Overseers of the School of Engineering and Applied Science of the University of Pennsylvania. Previously, Dr. Heilmeier served as a Director of TRW Inc. (automotive, space, defense, and information technology, 1992-2002), Compaq (1994-2002), Automatic Data Processing, Inc. (ADP) (technology-based business outsourcing, 1995-2002), INET Technologies Inc. (telecommunications network surveillance, 2001-2004), and Teletech Holdings (customer management services). He is the recipient of the 2005 Kyoto Prize in Advanced Technology for his invention of the liquid crystal display, and a member of the Consumer Electronics Hall of Fame.

James H. Keyes (67)

Year of Election or Appointment: 2007

Prior to his retirement in 2003, Mr. Keyes was Chairman, President, and Chief Executive Officer of Johnson Controls, Inc. (automotive supplier, 1993-2003). He currently serves as a member of the boards of LSI Logic Corporation (semiconductor technologies), Navistar International Corporation (manufacture and sale of trucks, buses, and diesel engines, 2002-present), and Pitney Bowes, Inc. (integrated mail, messaging, and document management solutions).

Marie L. Knowles (61)

Year of Election or Appointment: 2001

Prior to Ms. Knowles' retirement in June 2000, she served as Executive Vice President and Chief Financial Officer of Atlantic Richfield Company (ARCO) (diversified energy, 1996-2000). From 1993 to 1996, she was a Senior Vice President of ARCO and President of ARCO Transportation Company. She served as a Director of ARCO from 1996 to 1998. She currently serves as a Director of Phelps Dodge Corporation (copper mining and manufacturing) and McKesson Corporation (healthcare service, 2002-present). Ms. Knowles is a Trustee of the Brookings Institution and the Catalina Island Conservancy and also serves as a member of the Advisory Board for the School of Engineering of the University of Southern California.

Ned C. Lautenbach (63)

Year of Election or Appointment: 2000

Mr. Lautenbach is Chairman of the Independent Trustees (2006-present). Mr. Lautenbach has been a partner of Clayton, Dubilier & Rice, Inc. (private equity investment firm) since September 1998. Previously, Mr. Lautenbach was with the International Business Machines Corporation (IBM) from 1968 until his retirement in 1998. Mr. Lautenbach serves as a Director of Sony Corporation (2006-present) and Eaton Corporation (diversified industrial) as well as the Philharmonic Center for the Arts in Naples, Florida. He also is a member of the Board of Trustees of Fairfield University (2005-present), as well as a member of the Council on Foreign Relations.

Cornelia M. Small (63)

Year of Election or Appointment: 2005

Ms. Small is a member (2000-present) and Chairperson (2002-present) of the Investment Committee, and a member (2002-present) of the Board of Trustees of Smith College. Previously, she served as Chief Investment Officer (1999-2000), Director of Global Equity Investments (1996-1999), and a member of the Board of Directors of Scudder, Stevens & Clark (1990-1997) and Scudder Kemper Investments (1997-1999). In addition, Ms. Small served as Co-Chair (2000-2003) of the Annual Fund for the Fletcher School of Law and Diplomacy.

William S. Stavropoulos (68)

Year of Election or Appointment: 2002

Mr. Stavropoulos is Chairman Emeritus of the Board of Directors of The Dow Chemical Company. Since joining The Dow Chemical Company in 1967, Mr. Stavropoulos served in numerous senior management positions, including President (1993-2000; 2002-2003), CEO (1995-2000; 2002-2004), and Chairman of the Executive Committee (2000-2004). Currently, he is a Director of NCR Corporation (data warehousing and technology solutions), Chemical Financial Corporation, Maersk Inc. (industrial conglomerate, 2002-present), Tyco International, Inc. (multinational manufacturing and services, 2007-present), and a member of the Advisory Board for Metalmark Capital (private equity investment firm, 2005-present). He is a special advisor to Clayton, Dubilier & Rice, Inc., a private equity investment firm. He also serves as a member of the Board of Trustees of the American Enterprise Institute for Public Policy Research. In addition, Mr. Stavropoulos is a member of The Business Council, J.P. Morgan International Council and the University of Notre Dame Advisory Council for the College of Science.

Kenneth L. Wolfe (68)

Year of Election or Appointment: 2005

Mr. Wolfe is Chairman and a Director of Hershey Foods Corporation (2007-present), where prior to his retirement in 2001, he was Chairman and Chief Executive Officer. Mr. Wolfe currently serves as a member of the board of Revlon Inc. (2004-present). Previously, Mr. Wolfe served as a member of the boards of Adelphia Communications Corporation (2003-2006) and Bausch & Lomb, Inc. (1993-2007).

Advisory Board Members and Executive Officers**:

Correspondence intended for Mr. Mauriello, Mr. Thomas, Mr. Wiley, Mr. Lacy, and Mr. Arthur Johnson may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235. Correspondence intended for each executive officer and Mr. Lynch may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Arthur E. Johnson (61)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Johnson serves as Senior Vice President of Corporate Strategic Development of Lockheed Martin Corporation (defense contractor). In addition, Mr. Johnson serves as a member of the Board of Directors of AGL Resources, Inc. (holding company, 2002-present), and IKON Office Solutions, Inc. (document management systems and services). Mr. Arthur E. Johnson and Mr. Edward C. Johnson 3d are not related.

Alan J. Lacy (54)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lacy serves as Senior Adviser (2007-present) of Oak Hill Capital Partners, L.P. (a private equity firm). Mr. Lacy also served as Vice Chairman and Chief Executive Officer of Sears Holdings Corporation and Sears, Roebuck and Co. (retail, 2005-2006; 2000-2005). In addition, Mr. Lacy serves as a member of the Board of Directors of The Western Union Company (global money transfer, 2006-present) and Bristol-Myers Squibb (global pharmaceuticals, 2007-present). Mr. Lacy is a Trustee of the National Parks Conservation Association and The Field Museum of Natural History.

Peter S. Lynch (64)

Year of Election or Appointment: 2003

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lynch is Vice Chairman and a Director of FMR, and Vice Chairman (2001-present) and a Director of FMR Co., Inc. Previously, Mr. Lynch served as a Trustee of the Fidelity funds (1990-2003). In addition, he serves as a Trustee of Boston College and as the Chairman of the Inner-City Scholarship Fund.

Joseph Mauriello (63)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Prior to his retirement in January 2006, Mr. Mauriello served in numerous senior management positions including Deputy Chairman and Chief Operating Officer (2004-2005), and Vice Chairman of Financial Services (2002-2004) of KPMG LLP US (professional services firm, 1965-2005). Mr. Mauriello currently serves as a member of the Board of Directors of XL Capital Ltd. (global insurance and re-insurance company, 2006-present) and of Arcadia Resources Inc. (health care services and products, 2007-present). He also served as a Director of the Hamilton Funds of the Bank of New York (2006-2007).

David M. Thomas (58)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Previously, Mr. Thomas served as Executive Chairman (2005-2006) and Chairman and Chief Executive Officer (2000-2005) of IMS Health, Inc. (pharmaceutical and healthcare information solutions). In addition, Mr. Thomas serves as a member of the Board of Directors of Fortune Brands, Inc. (consumer products holding company), and Interpublic Group of Companies, Inc. (marketing communication, 2004-present).

Michael E. Wiley (57)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Wiley also serves as a member of the Board of Trustees of the University of Tulsa (2000-2006; 2007-present). He serves as a Director of Tesoro Corporation (independent oil refiner and marketer, 2005-present), and a Director of Bill Barrett Corporation (exploration and production company, 2005-present). In addition, he also serves as a Director of Post Oak Bank (privately-held bank, 2004-present). Previously, Mr. Wiley served as a Sr. Energy Advisor of Katzenbach Partners, LLC (consulting firm, 2006-2007), as an Advisory Director of Riverstone Holdings (private investment firm), Chairman, President, and CEO of Baker Hughes, Inc. (oilfield services company, 2000-2004), and as Director of Spinnaker Exploration Company (exploration and production company, 2001-2005).

Kimberley H. Monasterio (44)

Year of Election or Appointment: 2007

President and Treasurer of the funds. Ms. Monasterio also serves as President and Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2004-present). Previously, Ms. Monasterio served as Deputy Treasurer of the Fidelity funds (2004-2006). Before joining Fidelity Investments, Ms. Monasterio served as Treasurer (2000-2004) and Chief Financial Officer (2002-2004) of the Franklin Templeton Funds and Senior Vice President of Franklin Templeton Services, LLC (2000-2004).

Walter C. Donovan (45)

Year of Election or Appointment: 2007

Vice President of the funds. Mr. Donovan also serves as Vice President of Fidelity's Equity Funds (2007-present). Mr. Donovan also serves as Executive Vice President of FMR (2005-present) and FMR Co., Inc. (2005-present). Previously, Mr. Donovan served as Vice President of Fidelity's High Income Funds (2005-2007), Fixed-Income Funds (2005-2006), certain Asset Allocation Funds (2005-2006), certain Balanced Funds (2005-2006), and as Vice President and Director of Fidelity's International Equity Trading group (1998-2005).

Bruce T. Herring (42)

Year of Election or Appointment: 2006

Vice President of the funds. Mr. Herring also serves as Vice President of certain Equity Funds (2006-present). Mr. Herring is Senior Vice President of FMR (2006-present) and Vice President of FMR Co., Inc. (2001-present). Previously, Mr. Herring served as a portfolio manager for Fidelity U.S. Equity Funds (2001-2005).

Eric D. Roiter (59)

Year of Election or Appointment: 2001

Secretary of the funds. He also serves as Secretary of other Fidelity funds; Vice President, General Counsel, and Secretary of FMR Co., Inc. (2001-present) and FMR; Assistant Secretary of Fidelity Management & Research (U.K.) Inc. (2001-present), Fidelity Research & Analysis Company (2001-present), and Fidelity Investments Money Management, Inc. (2001-present). Mr. Roiter is an Adjunct Member, Faculty of Law, at Boston College Law School (2003-present). Previously, Mr. Roiter served as Vice President and Secretary of Fidelity Distributors Corporation (FDC) (1998-2005).

John B. McGinty, Jr. (45)

Year of Election or Appointment: 2008

Assistant Secretary of the funds. Mr. McGinty also serves as Assistant Secretary of other Fidelity funds (2008-present) and is an employee of FMR LLC (2004-present). Mr. McGinty also serves as Senior Vice President, Secretary, and Chief Legal Officer of FDC (2007-present). Before joining Fidelity Investments, Mr. McGinty practiced law at Ropes & Gray, LLP.

R. Stephen Ganis (41)

Year of Election or Appointment: 2006

Anti-Money Laundering (AML) officer of the funds. Mr. Ganis also serves as AML officer of other Fidelity funds (2006-present) and FMR LLC (2003-present). Before joining Fidelity Investments, Mr. Ganis practiced law at Goodwin Procter, LLP (2000-2002).

Joseph B. Hollis (59)

Year of Election or Appointment: 2006

Chief Financial Officer of the funds. Mr. Hollis also serves as Chief Financial Officer of other Fidelity funds. Mr. Hollis is President of Fidelity Pricing and Cash Management Services (FPCMS) (2005-present). Mr. Hollis also serves as President and Director of Fidelity Service Company, Inc. (2006-present). Previously, Mr. Hollis served as Senior Vice President of Cash Management Services (1999-2002) and Investment Management Operations (2002-2005).

Kenneth A. Rathgeber (60)

Year of Election or Appointment: 2004

Chief Compliance Officer of the funds. Mr. Rathgeber also serves as Chief Compliance Officer of other Fidelity funds (2004-present) and Executive Vice President of Risk Oversight for Fidelity Investments (2002-present). He is Chief Compliance Officer of FMR (2005-present), FMR Co., Inc. (2005-present), Fidelity Management & Research (U.K.) Inc. (2005-present), Fidelity Research & Analysis Company (2005-present), Fidelity Investments Money Management, Inc. (2005-present), and Strategic Advisers, Inc. (2005-present). Previously, Mr. Rathgeber served as Executive Vice President and Chief Operating Officer for Fidelity Investments Institutional Services Company, Inc. (1998-2002).

Bryan A. Mehrmann (46)

Year of Election or Appointment: 2005

Deputy Treasurer of the funds. Mr. Mehrmann also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR. Previously, Mr. Mehrmann served as Vice President of Fidelity Investments Institutional Services Group (FIIS)/Fidelity Investments Institutional Operations Corporation, Inc. (FIIOC) Client Services (1998-2004).

Kenneth B. Robins (38)

Year of Election or Appointment: 2005

Deputy Treasurer of the funds. Mr. Robins also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2004-present). Before joining Fidelity Investments, Mr. Robins worked at KPMG LLP, where he was a partner in KPMG's department of professional practice (2002-2004) and a Senior Manager (1999-2000). In addition, Mr. Robins served as Assistant Chief Accountant, United States Securities and Exchange Commission (2000-2002).

Robert G. Byrnes (41)

Year of Election or Appointment: 2005

Assistant Treasurer of the funds. Mr. Byrnes also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Byrnes served as Vice President of FPCMS (2003-2005). Before joining Fidelity Investments, Mr. Byrnes worked at Deutsche Asset Management where he served as Vice President of the Investment Operations Group (2000-2003).

Peter L. Lydecker (54)

Year of Election or Appointment: 2004

Assistant Treasurer of the funds. Mr. Lydecker also serves as Assistant Treasurer of other Fidelity funds (2004) and is an employee of FMR.

Paul M. Murphy (60)

Year of Election or Appointment: 2007

Assistant Treasurer of the funds. Mr. Murphy also serves as Assistant Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2007-present). Previously, Mr. Murphy served as Chief Financial Officer of the Fidelity Funds (2005-2006), Vice President and Associate General Counsel of FMR (2007), and Senior Vice President of Fidelity Pricing and Cash Management Services Group (FPCMS) (1994-2007).

Gary W. Ryan (49)

Year of Election or Appointment: 2005

Assistant Treasurer of the funds. Mr. Ryan also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Ryan served as Vice President of Fund Reporting in FPCMS (1999-2005).

** FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Annual Report

Distributions

The funds hereby designate as capital gain dividend the amounts noted below for the taxable year ended January 31, 2008, or, if subsequently determined to be different, the net capital gain of such year.

Fidelity Large Cap Value Fund

$53,592,103

Fidelity Mid Cap Value Fund

$30,917,128

Fidelity Large Cap Growth Fund

$9,534,777

Fidelity Mid Cap Growth Fund

$25,157,607

A percentage of the dividends distributed during the fiscal year for the following funds qualifies for the dividends-received deduction for corporate shareholders:

March,
2007

December,
2007

Fidelity Large Cap Value Fund

Class A

45%

100%

Class T

45%

100%

Class B

45%

100%

Class C

45%

100%

Fidelity Mid Cap Value Fund

Class A

26%

100%

Class T

26%

100%

Class B

26%

0%

Class C

26%

0%

Fidelity Large Cap Growth Fund

Class A

91%

18%

Class T

91%

18%

Class B

91%

21%

Class C

91%

20%

A percentage of the dividends distributed during the fiscal year for the following funds may be taken into account as a dividend for purposes of the maximum rate under section 1(h)(11) of the Internal Revenue Code.

March,
2007

December,
2007

Fidelity Large Cap Value Fund

Class A

45%

100%

Class T

45%

100%

Class B

45%

100%

Class C

45%

100%

Fidelity Mid Cap Value Fund

Class A

26%

100%

Class T

26%

100%

Class B

26%

0%

Class C

26%

0%

Fidelity Large Cap Growth Fund

Class A

92%

19%

Class T

92%

20%

Class B

92%

22%

Class C

92%

21%

The fund will notify shareholders in January 2009 of amounts for use in preparing 2008 income tax returns.

Annual Report

Annual Report

Annual Report

Annual Report

Investment Adviser

Fidelity Management & Research Company
Boston, MA

Investment Sub-Adviser

FMR Co., Inc.

Fidelity Management & Research
(U.K.) Inc.

Fidelity Research & Analysis Company

Fidelity Investments Japan Limited

Fidelity International Investment Advisors

Fidelity International Investment Advisors
(U.K.) Limited

General Distributor

Fidelity Distributors Corporation
Boston, MA

Transfer and Service Agents

Fidelity Investments Institutional Operations
Company, Inc.
Boston, MA

Fidelity Service Company, Inc.
Boston, MA

Custodian

Citibank, N.A.
New York, NY

The Fidelity Telephone Connection

Mutual Fund 24-Hour Service

Exchanges/Redemptions
and Account Assistance 1-800-544-6666

Product Information 1-800-544-6666

Retirement Accounts 1-800-544-4774
(8 a.m. - 9 p.m.)

TDD Service 1-800-544-0118
for the deaf and hearing impaired
(9 a.m. - 9 p.m. Eastern time)

Fidelity Automated Service
Telephone (FAST®) (automated graphic)    1-800-544-5555

(automated graphic)    Automated line for quickest service

(Fidelity Investment logo)(registered trademark)
Corporate Headquarters
82 Devonshire St., Boston, MA 02109
www.fidelity.com

ALMC-UANN-0308
1.863127.100

Fidelity Advisor Large Cap Value

Institutional Class

Fidelity Advisor Mid Cap Value

Institutional Class

Fidelity Advisor Large Cap Growth

Institutional Class

Fidelity Advisor Mid Cap Growth

Institutional Class

Funds

Annual Report

January 31, 2008

(2_fidelity_logos) (Registered_Trademark)

Each Institutional Class
listed above is a class of
Fidelity® Large Cap Value,
Fidelity Mid Cap Value,
Fidelity Large Cap Growth, and
Fidelity Mid Cap Growth
Funds

Contents

Chairman's Message

<Click Here>

Ned Johnson's message to shareholders.

Shareholder Expense Example

<Click Here>

An example of shareholder expenses.

Fidelity Advisor Large Cap Value Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Advisor Mid Cap Value Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Advisor Large Cap Growth Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Fidelity Advisor Mid Cap Growth Fund

<Click Here>

Performance

<Click Here>

Management's Discussion

<Click Here>

Investment Changes

<Click Here>

Investments

<Click Here>

Financial Statements

Notes

<Click Here>

Notes to the Financial Statements

Report of Independent Registered Public Accounting Firm

<Click Here>

Trustees and Officers

<Click Here>

Distributions

<Click Here>

To view a fund's proxy voting guidelines and proxy voting record for the 12-month period ended June 30, visit http://www.fidelity.com (search for "proxy voting guidelines") or visit the Securities and Exchange Commission's (SEC) web site at http://www.sec.gov. You may also call 1-800-544-8544 to request a free copy of the proxy voting guidelines.

Standard & Poor's, S&P and S&P 500 are registered service marks of The McGraw-Hill Companies, Inc. and have been licensed for use by Fidelity Distributors Corporation.

Other third party marks appearing herein are the property of their respective owners.

All other marks appearing herein are registered or unregistered trademarks or service marks of FMR LLC or an affiliated company.

This report and the financial statements contained herein are submitted for the general information of the shareholders of the funds. This report is not authorized for distribution to prospective investors in the funds unless preceded or accompanied by an effective prospectus.

A fund files its complete schedule of portfolio holdings with the SEC for the first and third quarters of each fiscal year on Form N-Q. Forms N-Q are available on the SEC's web site at http://www.sec.gov. A fund's Forms N-Q may be reviewed and copied at the SEC's Public Reference Room in Washington, DC. Information regarding the operation of the SEC's Public Reference Room may be obtained by calling 1-800-SEC-0330. For a complete list of a fund's portfolio holdings, view the most recent holdings listing, semiannual report, or annual report on Fidelity's web site at http://www.fidelity.com or http://www.advisor.fidelity.com, as applicable.

NOT FDIC INSURED · MAY LOSE VALUE · NO BANK GUARANTEE

Neither the funds nor Fidelity Distributors Corporation is a bank.

Annual Report

Chairman's Message

(Photograph of Edward C. Johnson 3d.)

Dear Shareholder:

Stocks got off to a poor start in 2008, while investment-grade bonds and money markets showed positive returns, once again underscoring the importance of a diversified portfolio. Financial markets are always unpredictable, but there are a number of time-tested principles that can put the historical odds in your favor.

One of the basic tenets is to invest for the long term. Over time, riding out the markets' inevitable ups and downs has proven much more effective than selling into panic or chasing the hottest trend. Even missing only a few of the markets' best days can significantly diminish investor returns. Patience also affords the benefits of compounding - of earning interest on additional income or reinvested dividends and capital gains. There are tax advantages and cost benefits to consider as well. The more you sell, the more taxes you pay, and the more you trade, the higher the costs. While staying the course doesn't eliminate risk, it can considerably lessen the effect of short-term declines.

You can further manage your investing risk through diversification. And today, more than ever, geographic diversification should be taken into account. Studies

indicate that asset allocation is the single most important determinant of a portfolio's long-term success. The right mix of stocks, bonds and cash - aligned to your particular risk tolerance and investment objective - is very important. Age-appropriate rebalancing is also an essential aspect of asset allocation. For younger investors, an emphasis on equities - which historically have been the best-performing asset class over time - is encouraged. As investors near their specific goal, such as retirement or sending a child to college, consideration may be given to replacing volatile assets (e.g. common stocks) with more-stable fixed investments (bonds or savings plans).

A third investment principle - investing regularly - can help lower the average cost of your purchases. Investing a certain amount of money each month or quarter helps ensure you won't pay for all your shares at market highs. This strategy - known as dollar cost averaging - also reduces unconstructive "emotion" from investing, helping shareholders avoid selling weak performers just prior to an upswing, or chasing a hot performer just before a correction.

We invite you to contact us via the Internet, through our Investor Centers or over the phone. It is our privilege to provide you the information you need to make the investments that are right for you.

Sincerely,

/s/Edward C. Johnson 3d

Edward C. Johnson 3d

Annual Report

Shareholder Expense Example

As a shareholder of a Fund, you incur two types of costs: (1) transaction costs, including sales charges (loads) on purchase payments or redemption proceeds, redemption fees, and (2) ongoing costs, including management fees, distribution and/or service (12b-1) fees and other Fund expenses. This Example is intended to help you understand your ongoing costs (in dollars) of investing in the Funds and to compare these costs with the ongoing costs of investing in other mutual funds.

The Example is based on an investment of $1,000 invested at the beginning of the period and held for the entire period (August 1, 2007 to January 31, 2008).

Actual Expenses

The first line of the accompanying table for each class of each fund provides information about actual account values and actual expenses. You may use the information in this line, together with the amount you invested, to estimate the expenses that you paid over the period. Simply divide your account value by $1,000.00 (for example, an $8,600 account value divided by $1,000.00 = 8.6), then multiply the result by the number in the first line for a class of the fund under the heading entitled "Expenses Paid During Period" to estimate the expenses you paid on your account during this period. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, each Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Hypothetical Example for Comparison Purposes

The second line of the accompanying table for each class of each fund provides information about hypothetical account values and hypothetical expenses based on a Class' actual expense ratio and an assumed rate of return of 5% per year before expenses, which is not the Class' actual return. The hypothetical account values and expenses may not be used to estimate the actual ending account balance or expenses you paid for the period. You may use this information to compare the ongoing costs of investing in the Fund and other funds. To do so, compare this 5% hypothetical example with the 5% hypothetical examples that appear in the shareholder reports of the other funds. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, each Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Please note that the expenses shown in the table are meant to highlight your ongoing costs only and do not reflect any transaction costs. Therefore, the second line of the table is useful in comparing ongoing costs only, and will not help you determine the relative total costs of owning different funds. In addition, if these transactional costs were included, your costs would have been higher.

Beginning
Account Value
August 1, 2007

Ending
Account Value
January 31, 2008

Expenses Paid
During Period
*
August 1, 2007
to January 31, 2008

Fidelity Large Cap Value Fund

Class A

Actual

$ 1,000.00

$ 946.40

$ 6.13

HypotheticalA

$ 1,000.00

$ 1,018.90

$ 6.36

Class T

Actual

$ 1,000.00

$ 944.70

$ 7.35

HypotheticalA

$ 1,000.00

$ 1,017.64

$ 7.63

Class B

Actual

$ 1,000.00

$ 943.10

$ 9.80

HypotheticalA

$ 1,000.00

$ 1,015.12

$ 10.16

Class C

Actual

$ 1,000.00

$ 943.20

$ 9.65

HypotheticalA

$ 1,000.00

$ 1,015.27

$ 10.01

Large Cap Value

Actual

$ 1,000.00

$ 948.00

$ 4.27

HypotheticalA

$ 1,000.00

$ 1,020.82

$ 4.43

Institutional Class

Actual

$ 1,000.00

$ 948.00

$ 4.22

HypotheticalA

$ 1,000.00

$ 1,020.87

$ 4.38

Fidelity Mid Cap Value Fund

Class A

Actual

$ 1,000.00

$ 912.50

$ 5.54

HypotheticalA

$ 1,000.00

$ 1,019.41

$ 5.85

Beginning
Account Value
August 1, 2007

Ending
Account Value
January 31, 2008

Expenses Paid
During Period
*
August 1, 2007
to January 31, 2008

Class T

Actual

$ 1,000.00

$ 911.80

$ 6.79

HypotheticalA

$ 1,000.00

$ 1,018.10

$ 7.17

Class B

Actual

$ 1,000.00

$ 909.00

$ 9.09

HypotheticalA

$ 1,000.00

$ 1,015.68

$ 9.60

Class C

Actual

$ 1,000.00

$ 909.00

$ 9.24

HypotheticalA

$ 1,000.00

$ 1,015.53

$ 9.75

Mid Cap Value

Actual

$ 1,000.00

$ 913.40

$ 4.05

HypotheticalA

$ 1,000.00

$ 1,020.97

$ 4.28

Institutional Class

Actual

$ 1,000.00

$ 913.70

$ 4.34

HypotheticalA

$ 1,000.00

$ 1,020.67

$ 4.58

Fidelity Large Cap Growth Fund

Class A

Actual

$ 1,000.00

$ 955.40

$ 5.91

HypotheticalA

$ 1,000.00

$ 1,019.16

$ 6.11

Class T

Actual

$ 1,000.00

$ 955.60

$ 7.30

HypotheticalA

$ 1,000.00

$ 1,017.74

$ 7.53

Class B

Actual

$ 1,000.00

$ 952.20

$ 9.74

HypotheticalA

$ 1,000.00

$ 1,015.22

$ 10.06

Class C

Actual

$ 1,000.00

$ 953.00

$ 9.65

HypotheticalA

$ 1,000.00

$ 1,015.32

$ 9.96

Large Cap Growth

Actual

$ 1,000.00

$ 956.60

$ 4.78

HypotheticalA

$ 1,000.00

$ 1,020.32

$ 4.94

Institutional Class

Actual

$ 1,000.00

$ 958.20

$ 4.10

HypotheticalA

$ 1,000.00

$ 1,021.02

$ 4.23

Fidelity Mid Cap Growth Fund

Class A

Actual

$ 1,000.00

$ 909.40

$ 5.29

HypotheticalA

$ 1,000.00

$ 1,019.66

$ 5.60

Class T

Actual

$ 1,000.00

$ 905.80

$ 6.58

HypotheticalA

$ 1,000.00

$ 1,018.30

$ 6.97

Class B

Actual

$ 1,000.00

$ 904.90

$ 8.79

HypotheticalA

$ 1,000.00

$ 1,015.98

$ 9.30

Class C

Actual

$ 1,000.00

$ 904.20

$ 8.88

HypotheticalA

$ 1,000.00

$ 1,015.88

$ 9.40

Mid Cap Growth

Actual

$ 1,000.00

$ 909.50

$ 3.75

HypotheticalA

$ 1,000.00

$ 1,021.27

$ 3.97

Institutional Class

Actual

$ 1,000.00

$ 909.60

$ 3.32

HypotheticalA

$ 1,000.00

$ 1,021.73

$ 3.52

A 5% return per year before expenses

* Expenses are equal to each Class' annualized expense ratio (shown in the table below); multiplied by the average account value over the period, multiplied by 184/ 365 (to reflect the one-half year period).

Annual Report

Annualized
Expense Ratio

Fidelity Large Cap Value Fund

Class A

1.25%

Class T

1.50%

Class B

2.00%

Class C

1.97%

Large Cap Value

.87%

Institutional Class

.86%

Fidelity Mid Cap Value Fund

Class A

1.15%

Class T

1.41%

Class B

1.89%

Class C

1.92%

Mid Cap Value

.84%

Institutional Class

.90%

Fidelity Large Cap Growth Fund

Class A

1.20%

Class T

1.48%

Class B

1.98%

Class C

1.96%

Large Cap Growth

.97%

Institutional Class

.83%

Fidelity Mid Cap Growth Fund

Class A

1.10%

Class T

1.37%

Class B

1.83%

Class C

1.85%

Mid Cap Growth

.78%

Institutional Class

.69%

Annual Report

Fidelity Advisor Large Cap Value Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the class' dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table does not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Institutional Class B

-4.45%

14.09%

7.81%

A From November 15, 2001.

B The initial offering of Institutional Class shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Value, the original class of the fund.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Fidelity Advisor Large Cap Value Fund - Institutional Class on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell 1000® Value Index performed over the same period. The initial offering of Institutional Class took place on February 13, 2007. See above for additional information regarding the performance of Institutional Class.



Annual Report

Fidelity Advisor Large Cap Value Fund

Management's Discussion of Fund Performance

Comments from Bruce Dirks, Portfolio Manager of Fidelity Advisor Large Cap Value Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the year, Advisor Large Cap Value's Class A, Class T, Class B and Class C shares (excluding sales charges) all finished about even with or slightly ahead of the Russell 1000® Value Index, which was off 5.38%. (For specific class-level results, please refer to the performance section of this report.) During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among banks and real estate investment trusts (REITs). Favorable industry weightings within the weak financials and consumer discretionary sectors also added value, as did good stock picking in health care and pockets of information technology. Underweighting diversified financials giant Citigroup and domestic banking firm Wachovia added to relative performance, as both stocks fared poorly within an environment of deteriorating business fundamentals for financials. Among other strong contributors were Cummins, a diesel engine manufacturer, and CF Industries, an out-of-index company that produces fertilizer products. On the flip side, relative results were held back by unfavorable stock picks in consumer staples - especially among food retailers - energy and the transportation industry. Among the biggest detractors were Tyson Foods, a large processor of poultry and meat products, and YRC Worldwide, a transportation company. Underweighting such strong-performing energy stocks as Occidental Petroleum also hurt. Tyson, YRC and Occidental were no longer in the portfolio at period end.

For the year, Advisor Large Cap Value's Institutional Class shares finished ahead of the Russell 1000® Value Index, which was off 5.38%. (For specific class-level results, please refer to the performance section of this report.) During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among banks and real estate investment trusts (REITs). Favorable industry weightings within the weak financials and consumer discretionary sectors also added value, as did good stock picking in health care and pockets of information technology. Underweighting diversified financials giant Citigroup and domestic banking firm Wachovia added to relative performance, as both stocks fared poorly within an environment of deteriorating business fundamentals for financials. Among other strong contributors were Cummins, a diesel engine manufacturer, and CF Industries, an out-of-index company that produces fertilizer products. On the flip side, relative results were held back by unfavorable stock picks in consumer staples - especially among food retailers - energy and the transportation industry. Among the biggest detractors were Tyson Foods, a large processor of poultry and meat products, and YRC Worldwide, a transportation company. Underweighting such strong-performing energy stocks as Occidental Petroleum also hurt. Tyson, YRC and Occidental were no longer in the portfolio at period end.

The views expressed above reflect those of the portfolio manager only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Large Cap Value Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Exxon Mobil Corp.

4.3

4.5

AT&T, Inc.

3.8

3.8

General Electric Co.

3.4

3.0

ConocoPhillips

3.1

0.0

JPMorgan Chase & Co.

3.0

3.0

American International Group, Inc.

2.7

2.3

Chevron Corp.

2.7

2.5

Bank of America Corp.

2.4

2.4

Altria Group, Inc.

2.3

1.8

Verizon Communications, Inc.

2.2

2.3

29.9

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Financials

30.7

31.7

Energy

15.6

14.0

Industrials

10.6

10.7

Consumer Staples

8.5

7.3

Health Care

7.6

7.6

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.7%

Stocks 99.7%

Short-Term
Investments and
Net Other Assets 0.3%

Short-Term
Investments and
Net Other Assets 0.3%

* Foreign investments

5.3%

**Foreign investments

2.0%

Annual Report

Fidelity Large Cap Value Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.7%

Shares

Value

CONSUMER DISCRETIONARY - 7.3%

Automobiles - 0.5%

General Motors Corp. (d)

283,401

$ 8,023,082

Hotels, Restaurants & Leisure - 2.0%

McDonald's Corp.

557,500

29,854,125

Household Durables - 0.3%

Whirlpool Corp.

56,000

4,766,160

Internet & Catalog Retail - 0.3%

Expedia, Inc. (a)

198,600

4,571,772

Leisure Equipment & Products - 0.4%

Eastman Kodak Co.

261,400

5,209,702

Media - 1.2%

Liberty Media Corp. - Capital
Series A (a)

64,300

6,919,966

Viacom, Inc. Class B (non-vtg.) (a)

296,500

11,492,340

18,412,306

Multiline Retail - 0.7%

Macy's, Inc.

387,800

10,718,792

Specialty Retail - 0.8%

Home Depot, Inc.

414,300

12,706,581

Textiles, Apparel & Luxury Goods - 1.1%

NIKE, Inc. Class B (d)

255,700

15,792,032

TOTAL CONSUMER DISCRETIONARY

110,054,552

CONSUMER STAPLES - 8.5%

Beverages - 1.1%

Molson Coors Brewing Co. Class B

392,400

17,528,508

Food & Staples Retailing - 2.9%

BJ's Wholesale Club, Inc. (a)

147,100

4,771,924

Kroger Co.

621,500

15,817,175

Safeway, Inc.

267,700

8,296,023

SUPERVALU, Inc.

484,600

14,567,076

43,452,198

Food Products - 0.3%

Del Monte Foods Co.

459,500

4,121,715

Household Products - 1.9%

Procter & Gamble Co.

433,300

28,576,135

Tobacco - 2.3%

Altria Group, Inc.

450,000

34,119,000

TOTAL CONSUMER STAPLES

127,797,556

ENERGY - 15.6%

Energy Equipment & Services - 1.9%

National Oilwell Varco, Inc. (a)

232,500

14,003,475

Transocean, Inc. (a)

112,200

13,755,720

27,759,195

Oil, Gas & Consumable Fuels - 13.7%

Chevron Corp.

482,300

40,754,350

ConocoPhillips

574,900

46,175,968

Shares

Value

Exxon Mobil Corp.

744,700

$ 64,342,081

Hess Corp.

265,100

24,079,033

Marathon Oil Corp.

308,900

14,471,965

Valero Energy Corp.

274,801

16,265,471

206,088,868

TOTAL ENERGY

233,848,063

FINANCIALS - 30.7%

Capital Markets - 6.5%

GLG Partners, Inc. (a)(d)

1,247,500

14,708,025

Goldman Sachs Group, Inc.

143,200

28,750,264

Janus Capital Group, Inc. (d)

575,300

15,538,853

Lehman Brothers Holdings, Inc. (d)

459,300

29,473,281

State Street Corp.

120,500

9,895,460

98,365,883

Commercial Banks - 3.8%

Associated Banc-Corp.

181,900

5,125,942

PNC Financial Services Group, Inc.

113,200

7,428,184

Regions Financial Corp. (d)

222,500

5,615,900

Wachovia Corp.

507,800

19,768,654

Wells Fargo & Co.

359,100

12,212,991

Zions Bancorp

132,200

7,236,628

57,388,299

Consumer Finance - 0.9%

Capital One Financial Corp. (d)

252,700

13,850,487

Diversified Financial Services - 6.7%

Bank of America Corp.

813,700

36,087,595

Citigroup, Inc.

546,460

15,421,101

JPMorgan Chase & Co.

947,400

45,048,870

MSCI, Inc. Class A

115,197

3,798,045

100,355,611

Insurance - 10.5%

ACE Ltd.

281,000

16,393,540

American International Group, Inc.

742,300

40,945,268

Assurant, Inc.

142,600

9,253,314

Axis Capital Holdings Ltd.

226,400

9,065,056

Endurance Specialty Holdings Ltd.

252,800

10,243,456

Loews Corp.

357,400

16,687,006

PartnerRe Ltd.

157,400

12,478,672

Prudential Financial, Inc.

255,600

21,564,972

RenaissanceRe Holdings Ltd.

215,900

12,304,141

W.R. Berkley Corp.

284,000

8,593,840

157,529,265

Real Estate Investment Trusts - 2.0%

Annaly Capital Management, Inc.

716,400

14,127,408

Boston Properties, Inc.

77,000

7,077,840

ProLogis Trust

91,600

5,436,460

Public Storage

46,700

3,654,275

30,295,983

Common Stocks - continued

Shares

Value

FINANCIALS - continued

Thrifts & Mortgage Finance - 0.3%

Astoria Financial Corp.

167,400

$ 4,549,932

TOTAL FINANCIALS

462,335,460

HEALTH CARE - 7.6%

Biotechnology - 1.0%

Biogen Idec, Inc. (a)

239,600

14,603,620

Health Care Equipment & Supplies - 0.5%

Baxter International, Inc.

118,200

7,179,468

Health Care Providers & Services - 1.8%

Humana, Inc. (a)

169,900

13,642,970

Medco Health Solutions, Inc. (a)

275,400

13,792,032

27,435,002

Life Sciences Tools & Services - 1.4%

Charles River Laboratories International, Inc. (a)

160,000

9,936,000

Invitrogen Corp. (a)

127,800

10,948,626

20,884,626

Pharmaceuticals - 2.9%

Johnson & Johnson

349,000

22,077,740

Pfizer, Inc.

960,700

22,470,773

44,548,513

TOTAL HEALTH CARE

114,651,229

INDUSTRIALS - 10.6%

Aerospace & Defense - 3.2%

Honeywell International, Inc.

94,300

5,570,301

Northrop Grumman Corp.

165,400

13,126,144

Raytheon Co.

170,600

11,112,884

The Boeing Co.

212,800

17,700,704

47,510,033

Airlines - 0.5%

Northwest Airlines Corp. (a)

193,200

3,614,772

UAL Corp.

121,400

4,607,130

8,221,902

Commercial Services & Supplies - 0.6%

Allied Waste Industries, Inc. (a)

920,000

9,062,000

Construction & Engineering - 1.4%

KBR, Inc. (a)

488,600

15,434,874

URS Corp. (a)

133,200

5,847,480

21,282,354

Industrial Conglomerates - 3.4%

General Electric Co.

1,424,000

50,423,840

Machinery - 1.2%

AGCO Corp. (a)

127,000

7,647,940

Cummins, Inc.

220,900

10,665,052

18,312,992

Shares

Value

Road & Rail - 0.3%

Hertz Global Holdings, Inc. (a)

266,933

$ 3,982,640

TOTAL INDUSTRIALS

158,795,761

INFORMATION TECHNOLOGY - 3.1%

Computers & Peripherals - 1.6%

Hewlett-Packard Co.

147,600

6,457,500

International Business Machines Corp.

124,200

13,331,628

Western Digital Corp. (a)

176,300

4,663,135

24,452,263

Electronic Equipment & Instruments - 0.8%

Tyco Electronics Ltd.

337,100

11,397,351

IT Services - 0.5%

Accenture Ltd. Class A

215,500

7,460,610

Software - 0.2%

Oracle Corp. (a)

172,000

3,534,600

TOTAL INFORMATION TECHNOLOGY

46,844,824

MATERIALS - 4.1%

Chemicals - 2.2%

Celanese Corp. Class A

164,400

6,112,392

CF Industries Holdings, Inc.

76,500

8,180,145

Chemtura Corp.

1,255,300

8,410,510

The Mosaic Co. (a)

108,600

9,883,686

32,586,733

Containers & Packaging - 0.6%

Ball Corp.

91,600

4,203,524

Owens-Illinois, Inc. (a)

94,300

4,752,720

8,956,244

Metals & Mining - 1.3%

Freeport-McMoRan Copper & Gold, Inc. Class B

104,800

9,330,344

United States Steel Corp.

99,300

10,139,523

19,469,867

TOTAL MATERIALS

61,012,844

TELECOMMUNICATION SERVICES - 6.0%

Diversified Telecommunication Services - 6.0%

AT&T, Inc.

1,480,362

56,979,133

Verizon Communications, Inc.

842,100

32,707,164

89,686,297

UTILITIES - 6.2%

Electric Utilities - 1.7%

Entergy Corp.

112,000

12,116,160

PPL Corp.

179,000

8,756,680

Reliant Energy, Inc. (a)

255,100

5,425,977

26,298,817

Gas Utilities - 1.0%

Energen Corp.

244,700

15,391,630

Common Stocks - continued

Shares

Value

UTILITIES - continued

Independent Power Producers & Energy Traders - 2.4%

Constellation Energy Group, Inc.

92,200

$ 8,663,112

Mirant Corp. (a)

324,700

11,961,948

NRG Energy, Inc. (a)

382,800

14,772,252

35,397,312

Multi-Utilities - 1.1%

Public Service Enterprise Group, Inc.

170,100

16,329,600

TOTAL UTILITIES

93,417,359

TOTAL COMMON STOCKS

(Cost $1,476,386,384)

1,498,443,945

Money Market Funds - 5.2%

Fidelity Cash Central Fund, 3.79% (b)

9,464,505

9,464,505

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

69,571,507

69,571,507

TOTAL MONEY MARKET FUNDS

(Cost $79,036,012)

79,036,012

TOTAL INVESTMENT PORTFOLIO - 104.9%

(Cost $1,555,422,396)

1,577,479,957

NET OTHER ASSETS - (4.9)%

(74,027,771)

NET ASSETS - 100%

$ 1,503,452,186

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 625,964

Fidelity Securities Lending Cash Central Fund

89,576

Total

$ 715,540

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $42,711,239 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Value Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $69,644,477) - See accompanying schedule:

Unaffiliated issuers (cost $1,476,386,384)

$ 1,498,443,945

Fidelity Central Funds (cost $79,036,012)

79,036,012

Total Investments (cost $1,555,422,396)

$ 1,577,479,957

Receivable for investments sold

23,766,119

Receivable for fund shares sold

2,565,058

Dividends receivable

1,696,869

Distributions receivable from Fidelity Central Funds

33,811

Prepaid expenses

4,921

Total assets

1,605,546,735

Liabilities

Payable for investments purchased

$ 23,494,925

Payable for fund shares redeemed

7,847,405

Accrued management fee

720,873

Distribution fees payable

6,898

Other affiliated payables

363,497

Other payables and accrued expenses

89,444

Collateral on securities loaned, at value

69,571,507

Total liabilities

102,094,549

Net Assets

$ 1,503,452,186

Net Assets consist of:

Paid in capital

$ 1,523,198,617

Undistributed net investment income

4,864,259

Accumulated undistributed net realized gain (loss) on investments

(46,668,251)

Net unrealized appreciation (depreciation) on investments

22,057,561

Net Assets

$ 1,503,452,186

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering
Price

Class A:
Net Asset Value
and redemption price per share ($9,774,232 ÷ 721,934 shares)

$ 13.54

Maximum offering price per share (100/94.25 of $13.54)

$ 14.37

Class T:
Net Asset Value
and redemption price per share ($5,975,654 ÷ 441,693 shares)

$ 13.53

Maximum offering price per share (100/96.50 of $13.53)

$ 14.02

Class B:
Net Asset Value
and offering price per share ($1,859,809 ÷ 137,404 shares)A

$ 13.54

Class C:
Net Asset Value
and offering price per share ($1,208,181 ÷ 89,368 shares)A

$ 13.52

Large Cap Value:
Net Asset Value
, offering price and redemption price per share ($1,483,574,281 ÷ 109,324,284 shares)

$ 13.57

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($1,060,029 ÷ 78,261 shares)

$ 13.54

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Value Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 32,039,820

Interest

4,832

Income from Fidelity Central Funds (including $89,576 from security lending)

715,540

Total income

32,760,192

Expenses

Management fee
Basic fee

$ 9,024,987

Performance adjustment

288,595

Transfer agent fees

3,768,695

Distribution fees

73,728

Accounting and security lending fees

507,630

Custodian fees and expenses

37,007

Independent trustees' compensation

5,753

Registration fees

122,860

Audit

60,620

Legal

7,461

Interest

5,226

Miscellaneous

53,563

Total expenses before reductions

13,956,125

Expense reductions

(136,896)

13,819,229

Net investment income (loss)

18,940,963

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

18,026,031

Change in net unrealized appreciation (depreciation) on investment securities

(121,407,791)

Net gain (loss)

(103,381,760)

Net increase (decrease) in net assets resulting from operations

$ (84,440,797)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ 18,940,963

$ 9,973,926

Net realized gain (loss)

18,026,031

34,353,479

Change in net unrealized appreciation (depreciation)

(121,407,791)

98,674,962

Net increase (decrease) in net assets resulting from operations

(84,440,797)

143,002,367

Distributions to shareholders from net investment income

(14,286,599)

(10,401,399)

Distributions to shareholders from net realized gain

(90,733,678)

(14,021,724)

Total distributions

(105,020,277)

(24,423,123)

Share transactions - net increase (decrease)

320,162,513

685,040,997

Redemption fees

-

21,396

Total increase (decrease) in net assets

130,701,439

803,641,637

Net Assets

Beginning of period

1,372,750,747

569,109,110

End of period (including undistributed net investment income of $4,864,259 and undistributed net investment income
of $274,346, respectively)

$ 1,503,452,186

$ 1,372,750,747

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.12

Net realized and unrealized gain (loss)

(1.00)

Total from investment operations

(.88)

Distributions from net investment income

(.12)

Distributions from net realized gain

(.87)

Total distributions

(.99)

Net asset value, end of period

$ 13.54

Total Return B,C

(6.04)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.22% A

Expenses net of fee waivers, if any

1.22% A

Expenses net of all reductions

1.22% A

Net investment income (loss)

.81% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 9,774

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the sales charges.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.08

Net realized and unrealized gain (loss)

(1.01)

Total from investment operations

(.93)

Distributions from net investment income

(.08)

Distributions from net realized gain

(.87)

Total distributions

(.95)

Net asset value, end of period

$ 13.53

Total Return B,C

(6.34)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.47% A

Expenses net of fee waivers, if any

1.47% A

Expenses net of all reductions

1.47% A

Net investment income (loss)

.56% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 5,976

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the sales charges.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.01

Net realized and unrealized gain (loss)

(1.00)

Total from investment operations

(.99)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.87)

Total distributions

(.88)

Net asset value, end of period

$ 13.54

Total Return B,C

(6.74)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.99% A

Expenses net of fee waivers, if any

1.99% A

Expenses net of all reductions

1.99% A

Net investment income (loss)

.04% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,860

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the contingent deferred sales charge.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.01

Net realized and unrealized gain (loss)

(.98)

Total from investment operations

(.97)

Distributions from net investment income

(.05)

Distributions from net realized gain

(.87)

Total distributions

(.92)

Net asset value, end of period

$ 13.52

Total Return B,C

(6.61)%

Ratios to Average Net Assets E,H

Expenses before reductions

1.94% A

Expenses net of fee waivers, if any

1.94% A

Expenses net of all reductions

1.94% A

Net investment income (loss)

.09% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,208

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns do not include the effect of the contingent deferred sales charge.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Large Cap Value

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 15.19

$ 13.62

$ 12.04

$ 10.64

$ 8.17

Income from Investment Operations

Net investment income (loss) B

.18

.16

.17

.09 E

.09

Net realized and unrealized gain (loss)

(.80)

1.80

1.87

1.47

2.47

Total from investment operations

(.62)

1.96

2.04

1.56

2.56

Distributions from net investment income

(.13)

(.13)

(.11)

(.05)

(.09)

Distributions from net realized gain

(.87)

(.26)

(.35)

(.11)

-

Total distributions

(1.00)

(.39)

(.46)

(.16)

(.09)

Redemption fees added to paid in capital B

-

- G,H

- G

- G

- G

Net asset value, end of period

$ 13.57

$ 15.19

$ 13.62

$ 12.04

$ 10.64

Total Return A

(4.39)%

14.63%

17.09%

14.68%

31.44%

Ratios to Average Net Assets C,F

Expenses before reductions

.86%

.89%

.89%

1.07%

1.45%

Expenses net of fee waivers, if any

.85%

.89%

.89%

1.07%

1.20%

Expenses net of all reductions

.85%

.89%

.84%

1.05%

1.18%

Net investment income (loss)

1.18%

1.10%

1.32%

.79% E

.99%

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,483,574

$ 1,372,751

$ 569,109

$ 177,004

$ 25,168

Portfolio turnover rate D

204%

164%

175%

170%

72%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been .59%.

F Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

G Amount represents less than $.01 per share.

H The redemption fee was eliminated during the year January 31, 2007.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 15.41

Income from Investment Operations

Net investment income (loss) D

.17

Net realized and unrealized gain (loss)

(1.02)

Total from investment operations

(.85)

Distributions from net investment income

(.15)

Distributions from net realized gain

(.87)

Total distributions

(1.02)

Net asset value, end of period

$ 13.54

Total Return B,C

(5.82)%

Ratios to Average Net Assets E,H

Expenses before reductions

.85% A

Expenses net of fee waivers, if any

.85% A

Expenses net of all reductions

.84% A

Net investment income (loss)

1.19% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,060

Portfolio turnover rate F

204%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Advisor Mid Cap Value Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the class' dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table does not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Institutional Class B

-7.70%

15.63%

10.37%

A From November 15, 2001.

B The initial offering of Institutional Class shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Value, the original class of the fund.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Fidelity Advisor Mid Cap Value Fund - Institutional Class on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell Midcap® Value Index performed over the same period. The initial offering of Institutional Class took place on February 13, 2007. See above for additional information regarding the performance of Institutional Class.



Annual Report

Fidelity Advisor Mid Cap Value Fund

Management's Discussion of Fund Performance

Comments from Bruce Dirks, Portfolio Manager of Fidelity Advisor Mid Cap Value Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the year, Advisor Mid Cap Value's Class A, Class T, Class B and Class C shares (excluding sales charges) all finished slightly ahead of the benchmark Russell Midcap® Value Index, which declined 8.73%. (For specific class-level results, please refer to the performance section of this report.) During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among bank and asset management companies. Favorable industry weightings within the weak financials and consumer discretionary sectors also added value. Good stock picking in materials, energy and the equipment/services group of health care also helped. Among the fund's strongest contributors for the period were CF Industries, an out-of-index company that produces fertilizer products; Cummins, a diesel engine manufacturer; Annaly Capital Management, a real estate investment trust that owned AAA-rated mortgages; and National Oilwell Varco, an energy services contractor. Conversely, relative results were held back by some unfavorable stock selection decisions in the utilities, consumer staples and consumer discretionary sectors. The fund's poorest performers included transportation firm YRC Worldwide; original equipment manufacturer TRW Automotive; and photographic imaging company Eastman Kodak. YRC and TRW were no longer held at period end.

For the year, Advisor Mid Cap Value Fund's Institutional Class shares finished slightly ahead of the benchmark Russell Midcap® Value Index, which declined 8.73%. (For specific class-level results, please refer to the performance section of this report.) During a volatile period for equities, especially late in the period, the fund outperformed its benchmark in part due to solid stock selection in pockets of the financials sector - mainly among bank and asset management companies. Favorable industry weightings within the weak financials and consumer discretionary sectors also added value. Good stock picking in materials, energy and the equipment/services group of health care also helped. Among the fund's strongest contributors for the period were CF Industries, an out-of-index company that produces fertilizer products; Cummins, a diesel engine manufacturer; Annaly Capital Management, a real estate investment trust that owned AAA-rated mortgages; and National Oilwell Varco, an energy services contractor. Conversely, relative results were held back by some unfavorable stock selection decisions in the utilities, consumer staples and consumer discretionary sectors. The fund's poorest performers included transportation firm YRC Worldwide; original equipment manufacturer TRW Automotive; and photographic imaging company Eastman Kodak. YRC and TRW were no longer held at period end.

The views expressed above reflect those of the portfolio manager only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

10.Fidelity Mid Cap Value Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Hess Corp.

2.5

2.3

Edison International

2.1

1.5

Everest Re Group Ltd.

1.8

0.0

Annaly Capital Management, Inc.

1.7

0.8

Boston Properties, Inc.

1.7

0.0

Questar Corp.

1.7

0.0

Axis Capital Holdings Ltd.

1.6

0.8

Assurant, Inc.

1.6

1.2

PartnerRe Ltd.

1.6

1.2

Energen Corp.

1.6

1.8

17.9

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Financials

30.0

29.5

Utilities

14.0

12.5

Consumer Discretionary

13.1

13.1

Industrials

10.4

10.7

Energy

8.1

7.0

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.7%

Stocks 99.4%

Short-Term
Investments and
Net Other Assets 0.3%

Short-Term
Investments and
Net Other Assets 0.6%

* Foreign investments

13.1%

**Foreign investments

4.2%

Annual Report

Fidelity Mid Cap Value Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.7%

Shares

Value

CONSUMER DISCRETIONARY - 13.1%

Diversified Consumer Services - 0.5%

Apollo Group, Inc. Class A (non-vtg.) (a)

50,800

$ 4,050,792

Hotels, Restaurants & Leisure - 1.1%

Burger King Holdings, Inc.

303,600

7,999,860

Household Durables - 2.3%

Leggett & Platt, Inc. (d)

296,500

5,639,430

Whirlpool Corp. (d)

133,500

11,362,185

17,001,615

Internet & Catalog Retail - 1.2%

Expedia, Inc. (a)

242,600

5,584,652

Liberty Media Corp. - Interactive
Series A (a)

239,500

3,810,445

9,395,097

Leisure Equipment & Products - 1.4%

Eastman Kodak Co. (d)

534,300

10,648,599

Media - 2.1%

E.W. Scripps Co. Class A

181,100

7,374,392

Liberty Media Corp. - Capital
Series A (a)

74,900

8,060,738

15,435,130

Specialty Retail - 2.6%

Abercrombie & Fitch Co. Class A

96,200

7,666,178

AutoZone, Inc. (a)

48,200

5,826,416

RadioShack Corp.

136,800

2,373,480

TJX Companies, Inc.

125,600

3,963,936

19,830,010

Textiles, Apparel & Luxury Goods - 1.9%

Crocs, Inc. (a)(d)

219,600

7,639,884

Jones Apparel Group, Inc. (d)

399,000

6,703,200

14,343,084

TOTAL CONSUMER DISCRETIONARY

98,704,187

CONSUMER STAPLES - 7.1%

Beverages - 1.9%

Coca-Cola Enterprises, Inc.

311,600

7,188,612

Molson Coors Brewing Co. Class B

168,900

7,544,763

14,733,375

Food & Staples Retailing - 2.8%

BJ's Wholesale Club, Inc. (a)(d)

170,600

5,534,264

Safeway, Inc.

259,000

8,026,410

SUPERVALU, Inc.

239,600

7,202,376

20,763,050

Food Products - 2.2%

Corn Products International, Inc.

136,600

4,617,080

Dean Foods Co.

158,400

4,435,200

Del Monte Foods Co.

316,300

2,837,211

Tyson Foods, Inc. Class A

308,000

4,389,000

16,278,491

Shares

Value

Personal Products - 0.2%

Estee Lauder Companies, Inc. Class A

44,100

$ 1,861,020

TOTAL CONSUMER STAPLES

53,635,936

ENERGY - 8.1%

Energy Equipment & Services - 4.8%

Helmerich & Payne, Inc.

206,500

8,098,930

Nabors Industries Ltd. (a)

152,100

4,140,162

National Oilwell Varco, Inc. (a)

121,000

7,287,830

Patterson-UTI Energy, Inc. (d)

450,800

8,826,664

Superior Energy Services, Inc. (a)

74,400

2,982,696

Tidewater, Inc.

81,800

4,332,128

35,668,410

Oil, Gas & Consumable Fuels - 3.3%

Frontline Ltd. (NY Shares)

137,300

5,844,861

Hess Corp. (d)

211,800

19,237,794

25,082,655

TOTAL ENERGY

60,751,065

FINANCIALS - 30.0%

Capital Markets - 3.0%

GLG Partners, Inc. (a)(d)

756,200

8,915,598

Janus Capital Group, Inc. (d)

352,500

9,521,025

MF Global Ltd.

151,200

4,543,560

22,980,183

Commercial Banks - 3.5%

Associated Banc-Corp.

277,700

7,825,586

BOK Financial Corp.

71,195

3,878,704

East West Bancorp, Inc. (d)

188,900

4,544,934

Synovus Financial Corp.

219,900

2,904,879

Zions Bancorp

125,800

6,886,292

26,040,395

Diversified Financial Services - 0.6%

MSCI, Inc. Class A

136,800

4,510,296

Insurance - 13.9%

ACE Ltd.

158,000

9,217,720

Allied World Assurance Co. Holdings Ltd.

147,800

7,039,714

Assurant, Inc.

187,200

12,147,408

Axis Capital Holdings Ltd.

308,900

12,368,356

CNA Financial Corp.

189,800

6,451,302

Endurance Specialty Holdings Ltd.

286,800

11,621,136

Everest Re Group Ltd.

133,100

13,534,939

PartnerRe Ltd.

152,100

12,058,488

RenaissanceRe Holdings Ltd.

197,800

11,272,622

W.R. Berkley Corp.

303,200

9,174,832

104,886,517

Real Estate Investment Trusts - 7.0%

Annaly Capital Management, Inc.

658,200

12,979,704

Boston Properties, Inc.

138,600

12,740,112

Common Stocks - continued

Shares

Value

FINANCIALS - continued

Real Estate Investment Trusts - continued

Colonial Properties Trust (SBI)

97,800

$ 2,409,792

Kimco Realty Corp.

198,300

7,101,123

ProLogis Trust

155,700

9,240,795

Public Storage

73,700

5,767,025

Taubman Centers, Inc.

47,400

2,377,110

52,615,661

Thrifts & Mortgage Finance - 2.0%

Astoria Financial Corp.

262,500

7,134,750

Washington Federal, Inc.

323,700

7,904,754

15,039,504

TOTAL FINANCIALS

226,072,556

HEALTH CARE - 2.3%

Health Care Equipment & Supplies - 0.3%

Kinetic Concepts, Inc. (a)(d)

41,100

2,045,958

Health Care Providers & Services - 0.5%

Humana, Inc. (a)

42,500

3,412,750

Life Sciences Tools & Services - 1.2%

Charles River Laboratories International, Inc. (a)

74,200

4,607,820

Invitrogen Corp. (a)

54,600

4,677,582

9,285,402

Pharmaceuticals - 0.3%

Watson Pharmaceuticals, Inc. (a)

85,000

2,219,350

TOTAL HEALTH CARE

16,963,460

INDUSTRIALS - 10.4%

Aerospace & Defense - 1.7%

DRS Technologies, Inc.

88,400

4,744,428

L-3 Communications Holdings, Inc.

72,500

8,035,175

12,779,603

Airlines - 0.8%

Northwest Airlines Corp. (a)

152,600

2,855,146

UAL Corp.

75,800

2,876,610

5,731,756

Building Products - 0.6%

Lennox International, Inc.

126,800

4,711,888

Commercial Services & Supplies - 2.2%

Allied Waste Industries, Inc. (a)

700,300

6,897,955

R.R. Donnelley & Sons Co.

267,200

9,322,608

16,220,563

Construction & Engineering - 1.7%

KBR, Inc. (a)

309,000

9,761,310

URS Corp. (a)

69,800

3,064,220

12,825,530

Shares

Value

Industrial Conglomerates - 0.6%

McDermott International, Inc. (a)

94,500

$ 4,458,510

Machinery - 2.5%

AGCO Corp. (a)

106,000

6,383,320

Cummins, Inc.

117,300

5,663,244

Ingersoll-Rand Co. Ltd. Class A

168,400

6,655,168

18,701,732

Road & Rail - 0.3%

Hertz Global Holdings, Inc. (a)

160,200

2,390,184

TOTAL INDUSTRIALS

77,819,766

INFORMATION TECHNOLOGY - 6.4%

Communications Equipment - 0.7%

Juniper Networks, Inc. (a)

195,900

5,318,685

Computers & Peripherals - 2.2%

NCR Corp. (a)

422,600

9,077,448

Western Digital Corp. (a)

278,300

7,361,035

16,438,483

Electronic Equipment & Instruments - 1.4%

Avnet, Inc. (a)

152,500

5,430,525

Ingram Micro, Inc. Class A (a)

173,400

3,083,052

Jabil Circuit, Inc.

174,200

2,308,150

10,821,727

Semiconductors & Semiconductor Equipment - 1.6%

Atmel Corp. (a)

1,316,800

4,161,088

MEMC Electronic Materials, Inc. (a)

25,000

1,786,500

Novellus Systems, Inc. (a)

111,700

2,653,992

Varian Semiconductor Equipment Associates, Inc. (a)

109,100

3,514,111

12,115,691

Software - 0.5%

Compuware Corp. (a)

405,900

3,450,150

TOTAL INFORMATION TECHNOLOGY

48,144,736

MATERIALS - 6.8%

Chemicals - 3.1%

Celanese Corp. Class A

146,800

5,458,024

CF Industries Holdings, Inc.

57,100

6,105,703

Chemtura Corp. (d)

867,400

5,811,580

The Mosaic Co. (a)

71,200

6,479,912

23,855,219

Containers & Packaging - 1.9%

Ball Corp.

85,100

3,905,239

Owens-Illinois, Inc. (a)

91,500

4,611,600

Smurfit-Stone Container Corp. (a)(d)

603,500

5,727,215

14,244,054

Common Stocks - continued

Shares

Value

MATERIALS - continued

Metals & Mining - 1.8%

Cleveland-Cliffs, Inc. (d)

42,700

$ 4,348,568

United States Steel Corp.

88,000

8,985,680

13,334,248

TOTAL MATERIALS

51,433,521

TELECOMMUNICATION SERVICES - 1.5%

Diversified Telecommunication Services - 1.5%

Embarq Corp.

116,500

5,277,450

Qwest Communications International, Inc.

1,043,800

6,137,544

11,414,994

UTILITIES - 14.0%

Electric Utilities - 4.6%

Allegheny Energy, Inc.

123,300

6,755,607

Edison International

305,600

15,940,096

Reliant Energy, Inc. (a)

558,600

11,881,422

34,577,125

Gas Utilities - 3.3%

Energen Corp.

191,700

12,057,930

Questar Corp.

247,300

12,590,043

24,647,973

Independent Power Producers & Energy Traders - 4.2%

Constellation Energy Group, Inc.

103,700

9,743,652

Mirant Corp. (a)

315,100

11,608,284

NRG Energy, Inc. (a)

268,200

10,349,838

31,701,774

Multi-Utilities - 1.9%

CMS Energy Corp.

583,000

9,135,610

OGE Energy Corp.

162,500

5,318,625

14,454,235

TOTAL UTILITIES

105,381,107

TOTAL COMMON STOCKS

(Cost $760,656,463)

750,321,328

Money Market Funds - 10.6%

Shares

Value

Fidelity Cash Central Fund, 3.79% (b)

5,496,422

$ 5,496,422

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

74,160,825

74,160,825

TOTAL MONEY MARKET FUNDS

(Cost $79,657,247)

79,657,247

TOTAL INVESTMENT PORTFOLIO - 110.3%

(Cost $840,313,710)

829,978,575

NET OTHER ASSETS - (10.3)%

(77,171,615)

NET ASSETS - 100%

$ 752,806,960

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 394,997

Fidelity Securities Lending Cash Central Fund

169,409

Total

$ 564,406

Other Information

Distribution of investments by country of issue, as a percentage of total net assets, is as follows:

United States of America

86.9%

Bermuda

11.3%

Cayman Islands

1.2%

Others (individually less than 1%)

0.6%

100.0%

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $31,051,104 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Value Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $73,863,006) - See accompanying schedule:

Unaffiliated issuers (cost $760,656,463)

$ 750,321,328

Fidelity Central Funds (cost $79,657,247)

79,657,247

Total Investments (cost $840,313,710)

$ 829,978,575

Receivable for investments sold

20,806,082

Receivable for fund shares sold

2,652,227

Dividends receivable

223,102

Distributions receivable from Fidelity Central Funds

36,747

Prepaid expenses

2,661

Other receivables

46

Total assets

853,699,440

Liabilities

Payable for investments purchased

$ 20,180,467

Payable for fund shares redeemed

5,956,855

Accrued management fee

312,903

Distribution fees payable

5,417

Other affiliated payables

203,019

Other payables and accrued expenses

72,994

Collateral on securities loaned, at value

74,160,825

Total liabilities

100,892,480

Net Assets

$ 752,806,960

Net Assets consist of:

Paid in capital

$ 788,210,259

Undistributed net investment income

1,465,714

Accumulated undistributed net realized gain (loss) on investments

(26,533,878)

Net unrealized appreciation (depreciation) on investments

(10,335,135)

Net Assets

$ 752,806,960

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($7,444,673 ÷ 494,661 shares)

$ 15.05

Maximum offering price per share (100/94.25 of $15.05)

$ 15.97

Class T:
Net Asset Value
and redemption price per share ($3,714,323 ÷ 247,029 shares)

$ 15.04

Maximum offering price per share (100/96.50 of $15.04)

$ 15.59

Class B:
Net Asset Value
and offering price per share ($1,304,436 ÷ 87,007 shares)A

$ 14.99

Class C:
Net Asset Value
and offering price per share ($1,657,889 ÷ 110,680 shares)A

$ 14.98

Mid Cap Value:
Net Asset Value
, offering price and redemption price per share ($737,234,029 ÷ 48,847,921 shares)

$ 15.09

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($1,451,610 ÷ 96,411 shares)

$ 15.06

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Value Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 11,189,048

Interest

8,457

Income from Fidelity Central Funds (including $169,409 from security lending)

564,406

Total income

11,761,911

Expenses

Management fee
Basic fee

$ 5,084,079

Performance adjustment

(317,451)

Transfer agent fees

2,236,010

Distribution fees

53,691

Accounting and security lending fees

315,091

Custodian fees and expenses

28,074

Independent trustees' compensation

3,227

Registration fees

145,678

Audit

58,698

Legal

4,195

Interest

4,946

Miscellaneous

29,800

Total expenses before reductions

7,646,038

Expense reductions

(105,134)

7,540,904

Net investment income (loss)

4,221,007

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

(3,216,421)

Change in net unrealized appreciation (depreciation) on investment securities

(85,150,820)

Net gain (loss)

(88,367,241)

Net increase (decrease) in net assets resulting from operations

$ (84,146,234)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ 4,221,007

$ 2,688,685

Net realized gain (loss)

(3,216,421)

24,851,516

Change in net unrealized appreciation (depreciation)

(85,150,820)

43,131,851

Net increase (decrease) in net assets resulting from operations

(84,146,234)

70,672,052

Distributions to shareholders from net investment income

(3,037,015)

(2,949,768)

Distributions to shareholders from net realized gain

(37,140,454)

(13,997,322)

Total distributions

(40,177,469)

(16,947,090)

Share transactions - net increase (decrease)

198,288,449

259,218,977

Redemption fees

47,852

33,264

Total increase (decrease) in net assets

74,012,598

312,977,203

Net Assets

Beginning of period

678,794,362

365,817,159

End of period (including undistributed net investment income of $1,465,714 and undistributed net investment income
of $375,291, respectively)

$ 752,806,960

$ 678,794,362

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

.03

Net realized and unrealized gain (loss)

(1.78)

Total from investment operations

(1.75)

Distributions from net investment income

(.06)

Distributions from net realized gain

(.77)

Total distributions

(.83)

Net asset value, end of period

$ 15.05

Total Return B,C,D

(10.28)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.14% A

Expenses net of fee waivers, if any

1.14% A

Expenses net of all reductions

1.13% A

Net investment income (loss)

.16% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 7,445

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.02)

Net realized and unrealized gain (loss)

(1.76)

Total from investment operations

(1.78)

Distributions from net investment income

(.04)

Distributions from net realized gain

(.77)

Total distributions

(.81)

Net asset value, end of period

$ 15.04

Total Return B,C,D

(10.46)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.39% A

Expenses net of fee waivers, if any

1.39% A

Expenses net of all reductions

1.39% A

Net investment income (loss)

(.10)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 3,714

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.10)

Net realized and unrealized gain (loss)

(1.76)

Total from investment operations

(1.86)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.77)

Total distributions

(.78)

Net asset value, end of period

$ 14.99

Total Return B,C,D

(10.88)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.89% A

Expenses net of fee waivers, if any

1.89% A

Expenses net of all reductions

1.89% A

Net investment income (loss)

(.59)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,304

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) E

(.10)

Net realized and unrealized gain (loss)

(1.77)

Total from investment operations

(1.87)

Distributions from net investment income

(.01)

Distributions from net realized gain

(.77)

Total distributions

(.78)

Net asset value, end of period

$ 14.98

Total Return B,C,D

(10.94)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.90% A

Expenses net of fee waivers, if any

1.90% A

Expenses net of all reductions

1.90% A

Net investment income (loss)

(.60)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,658

Portfolio turnover rate G

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Mid Cap Value

Years ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 17.18

$ 15.65

$ 14.14

$ 12.32

$ 8.85

Income from Investment Operations

Net investment income (loss) B

.08

.09

.16 E

.08

.06

Net realized and unrealized gain (loss)

(1.34)

1.98

2.59

2.10

3.45

Total from investment operations

(1.26)

2.07

2.75

2.18

3.51

Distributions from net investment income

(.06)

(.09)

(.10)

(.04)

(.04)

Distributions from net realized gain

(.77)

(.45)

(1.15)

(.32)

-

Total distributions

(.83)

(.54)

(1.24) H

(.36)

(.04)

Redemption fees added to paid in capital B,G

-

-

-

-

-

Net asset value, end of period

$ 15.09

$ 17.18

$ 15.65

$ 14.14

$ 12.32

Total Return A

(7.67)%

13.48%

19.97%

17.75%

39.69%

Ratios to Average Net Assets C,F

Expenses before reductions

.83%

.84%

.86%

.91%

1.07%

Expenses net of fee waivers, if any

.82%

.84%

.86%

.91%

1.07%

Expenses net of all reductions

.82%

.84%

.81%

.90%

1.05%

Net investment income (loss)

.47%

.56%

1.08% E

.59%

.55%

Supplemental Data

Net assets, end of period (000 omitted)

$ 737,234

$ 678,794

$ 365,817

$ 153,231

$ 95,797

Portfolio turnover rate D

264%

187%

207%

196%

97%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.04 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been .81%.

F Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

G Amount represents less than $.01 per share.

H Total distributions of $1.24 per share is comprised of distributions from net investment income of $0.95 and distributions from net realized gain of $1.145 per share.

Financial Highlights - Institutional Class

Years ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 17.63

Income from Investment Operations

Net investment income (loss) D

.07

Net realized and unrealized gain (loss)

(1.78)

Total from investment operations

(1.71)

Distributions from net investment income

(.09)

Distributions from net realized gain

(.77)

Total distributions

(.86)

Net asset value, end of period

$ 15.06

Total Return B,C

(10.06)%

Ratios to Average Net Assets E,H

Expenses before reductions

.89% A

Expenses net of fee waivers, if any

.89% A

Expenses net of all reductions

.88% A

Net investment income (loss)

.41% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,452

Portfolio turnover rate F

264%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Advisor Large Cap Growth Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the class' dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table does not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Institutional Class B

-7.18%

10.77%

2.36%

A From November 15, 2001.

B The initial offering of Institutional Class shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Large Cap Growth, the original class of the fund.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Fidelity Advisor Large Cap Growth Fund - Institutional Class on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell 1000® Growth Index performed over the same period. The initial offering of Institutional Class took place on February 13, 2007. See above for additional information regarding the performance of Institutional Class.



Annual Report

Fidelity Advisor Large Cap Growth Fund

Management's Discussion of Fund Performance

Comments from Edward Best, Portfolio Manager of Fidelity Advisor Large Cap Growth Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the 12 months ending January 31, 2008, the fund's Class A, Class T, Class B and Class C shares (excluding sales charges) trailed the 0.51% gain of the Russell 1000® Growth Index. (For specific class-level results, please refer to the performance section of this report.) Poor security selection in consumer discretionary, information technology and industrials accounted for the majority of the fund's underperformance compared with the index. Within consumer discretionary, the fund was hurt most by unproductive picks within the retailing and homebuilding segments. Consumer electronics retailer RadioShack and discount retailer Big Lots fell on investors' concerns about the companies' ability to weather recessionary concerns and a weak U.S. consumer. Pharmaceuticals company Sepracor suffered due to a change in Medicare reimbursement for one of its top-selling patented drugs. Airlines US Airways and AMR Corp. - parent company of American Airlines - underperformed due to increased pressure from rising energy prices. The fund's positions in homebuilders KB Home, Ryland Group, D.R. Horton and Lennar, all of which I sold when I took over the fund in June, also held back performance. While our overall stock picking within industrials detracted, stock selection within the sector's capital goods segment actually helped the fund. Specifically, Manitowoc, a major supplier of lifting equipment to the global construction industry, aided performance. Also within capital goods, defense contractor Lockheed Martin benefited from an increase in U.S. defense spending. Stock selection among health care names was positive as well. Pharmaceutical company Biogen Idec gained on news of the re-launch of its multiple sclerosis drug, Tysabri. While stock selection in technology hurt overall, there were some contributors. Apple advanced as the stock did well on the success of its iPod, iPhone and iMac products. Software developer Microsoft also aided performance. By the end of the period, I sold US Airways, AMR Corp., RadioShack, Big Lots, Sepracor and Apple to take advantage of other opportunities in the market.

For the 12 months ending January 31, 2008, the fund's Institutional Class shares trailed the 0.51% gain of the Russell 1000® Growth Index. (For specific class-level results, please refer to the performance section of this report.) Poor security selection in consumer discretionary, information technology and industrials accounted for the majority of the fund's underperformance compared with the index. Within consumer discretionary, the fund was hurt most by unproductive picks within the retailing and homebuilding segments. Consumer electronics retailer RadioShack and discount retailer Big Lots fell on investors' concerns about the companies' ability to weather recessionary concerns and a weak U.S. consumer. Pharmaceuticals company Sepracor suffered due to a change in Medicare reimbursement for one of its top-selling patented drugs. Airlines US Airways and AMR Corp. - parent company of American Airlines - underperformed due to increased pressure from rising energy prices. The fund's positions in homebuilders KB Home, Ryland Group, D.R. Horton and Lennar, all of which I sold when I took over the fund in June, also held back performance. While our overall stock picking within industrials detracted, stock selection within the sector's capital goods segment actually helped the fund. Specifically, Manitowoc, a major supplier of lifting equipment to the global construction industry, aided performance. Also within capital goods, defense contractor Lockheed Martin benefited from an increase in U.S. defense spending. Stock selection among health care names also was positive. Pharmaceutical company Biogen Idec gained on news of the re-launch of its multiple sclerosis drug, Tysabri. While stock selection in technology hurt overall, there were some contributors. Apple advanced as the stock did well on the success of its iPod, iPhone and iMac products. Software developer Microsoft also aided performance. By the end of the period, I sold US Airways, AMR Corp., RadioShack, Big Lots, Sepracor, and Apple to take advantage of other opportunities in the market.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Large Cap Growth Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Microsoft Corp.

5.0

5.0

Exxon Mobil Corp.

4.4

1.6

Oracle Corp.

4.1

0.0

The Boeing Co.

3.9

0.0

Cisco Systems, Inc.

3.8

2.4

Bristol-Myers Squibb Co.

3.6

0.0

Accenture Ltd. Class A

3.3

3.3

NIKE, Inc. Class B

3.3

0.0

NYMEX Holdings, Inc.

3.1

0.0

Kroger Co.

3.0

3.1

37.5

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Information Technology

26.0

26.9

Health Care

16.8

16.2

Industrials

14.2

14.2

Consumer Discretionary

11.9

13.5

Consumer Staples

10.9

9.9

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 99.4%

Stocks 99.8%

Short-Term
Investments and
Net Other Assets 0.6%

Short-Term
Investments and
Net Other Assets 0.2%

* Foreign investments

4.4%

**Foreign
investments

5.5%

Annual Report

Fidelity Large Cap Growth Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 99.4%

Shares

Value

CONSUMER DISCRETIONARY - 11.9%

Auto Components - 1.6%

The Goodyear Tire & Rubber Co. (a)

99,300

$ 2,499,381

Diversified Consumer Services - 1.2%

Apollo Group, Inc. Class A (non-vtg.) (a)

22,600

1,802,124

Hotels, Restaurants & Leisure - 2.6%

McDonald's Corp.

74,200

3,973,410

Media - 2.3%

Regal Entertainment Group Class A (d)

189,300

3,509,622

Specialty Retail - 0.9%

Gamestop Corp. Class A (a)

27,400

1,417,402

Textiles, Apparel & Luxury Goods - 3.3%

NIKE, Inc. Class B

79,900

4,934,624

TOTAL CONSUMER DISCRETIONARY

18,136,563

CONSUMER STAPLES - 10.9%

Beverages - 5.0%

Pepsi Bottling Group, Inc.

114,200

3,979,870

PepsiCo, Inc.

52,500

3,579,975

7,559,845

Food & Staples Retailing - 3.0%

Kroger Co.

181,400

4,616,630

Food Products - 2.4%

Kellogg Co.

77,200

3,697,880

Household Products - 0.5%

Energizer Holdings, Inc. (a)

8,300

777,046

TOTAL CONSUMER STAPLES

16,651,401

ENERGY - 7.9%

Energy Equipment & Services - 0.9%

Baker Hughes, Inc.

5,700

370,101

Nabors Industries Ltd. (a)

24,400

664,168

Schlumberger Ltd. (NY Shares)

5,000

377,300

1,411,569

Oil, Gas & Consumable Fuels - 7.0%

Exxon Mobil Corp.

78,000

6,739,200

Sunoco, Inc.

13,000

808,600

Valero Energy Corp.

53,000

3,137,070

10,684,870

TOTAL ENERGY

12,096,439

FINANCIALS - 7.5%

Capital Markets - 4.4%

Charles Schwab Corp.

198,200

4,419,860

Northern Trust Corp.

30,900

2,266,824

6,686,684

Shares

Value

Diversified Financial Services - 3.1%

NYMEX Holdings, Inc. (d)

41,700

$ 4,795,500

TOTAL FINANCIALS

11,482,184

HEALTH CARE - 16.8%

Biotechnology - 1.2%

Biogen Idec, Inc. (a)

29,300

1,785,835

Health Care Equipment & Supplies - 2.8%

Baxter International, Inc.

69,700

4,233,578

Health Care Providers & Services - 9.2%

Aetna, Inc.

86,000

4,580,360

Express Scripts, Inc. (a)

11,400

769,386

Medco Health Solutions, Inc. (a)

35,200

1,762,816

UnitedHealth Group, Inc.

47,700

2,425,068

WellPoint, Inc. (a)

57,800

4,519,960

14,057,590

Pharmaceuticals - 3.6%

Bristol-Myers Squibb Co.

234,400

5,435,736

TOTAL HEALTH CARE

25,512,739

INDUSTRIALS - 14.2%

Aerospace & Defense - 5.6%

Lockheed Martin Corp.

5,900

636,728

Precision Castparts Corp.

16,900

1,923,220

The Boeing Co.

71,500

5,947,370

8,507,318

Commercial Services & Supplies - 3.9%

Allied Waste Industries, Inc. (a)

460,200

4,532,970

HNI Corp. (d)

15,400

518,364

Manpower, Inc.

15,300

860,778

5,912,112

Construction & Engineering - 2.7%

Fluor Corp.

7,500

912,525

Shaw Group, Inc. (a)

13,800

779,700

URS Corp. (a)

54,700

2,401,330

4,093,555

Industrial Conglomerates - 0.8%

McDermott International, Inc. (a)

24,500

1,155,910

Machinery - 1.2%

AGCO Corp. (a)

8,700

523,914

Cummins, Inc.

16,500

796,620

Manitowoc Co., Inc.

15,400

587,048

1,907,582

TOTAL INDUSTRIALS

21,576,477

INFORMATION TECHNOLOGY - 26.0%

Communications Equipment - 3.8%

Cisco Systems, Inc. (a)

236,300

5,789,350

Common Stocks - continued

Shares

Value

INFORMATION TECHNOLOGY - continued

Computers & Peripherals - 0.5%

Lexmark International, Inc. Class A (a)

1,400

$ 50,694

Western Digital Corp. (a)

29,000

767,050

817,744

Electronic Equipment & Instruments - 0.5%

National Instruments Corp.

27,400

735,964

IT Services - 6.2%

Accenture Ltd. Class A

143,600

4,971,432

Mastercard, Inc. Class A (d)

7,000

1,449,000

The Western Union Co.

135,200

3,028,480

9,448,912

Semiconductors & Semiconductor Equipment - 2.2%

MEMC Electronic Materials, Inc. (a)

12,500

893,250

NVIDIA Corp. (a)

82,500

2,028,675

Texas Instruments, Inc.

12,000

371,160

3,293,085

Software - 12.8%

Autodesk, Inc. (a)

76,100

3,131,515

BMC Software, Inc. (a)

74,200

2,377,368

Microsoft Corp.

233,200

7,602,320

Oracle Corp. (a)

306,900

6,306,795

19,417,998

TOTAL INFORMATION TECHNOLOGY

39,503,053

MATERIALS - 3.0%

Chemicals - 2.0%

Air Products & Chemicals, Inc.

4,200

378,084

E.I. du Pont de Nemours & Co.

10,100

456,318

Monsanto Co.

12,800

1,439,232

Praxair, Inc.

4,700

380,277

The Mosaic Co. (a)

4,000

364,040

3,017,951

Metals & Mining - 1.0%

Freeport-McMoRan Copper & Gold, Inc. Class B

8,400

747,852

Newmont Mining Corp.

7,100

385,814

Southern Copper Corp.

4,400

412,896

1,546,562

TOTAL MATERIALS

4,564,513

Shares

Value

TELECOMMUNICATION SERVICES - 0.3%

Wireless Telecommunication Services - 0.3%

Crown Castle International Corp. (a)

10,700

$ 387,233

UTILITIES - 0.9%

Electric Utilities - 0.9%

Exelon Corp.

18,300

1,394,277

TOTAL COMMON STOCKS

(Cost $156,064,347)

151,304,879

Money Market Funds - 6.4%

Fidelity Cash Central Fund, 3.79% (b)

1,837,290

1,837,290

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

7,887,500

7,887,500

TOTAL MONEY MARKET FUNDS

(Cost $9,724,790)

9,724,790

TOTAL INVESTMENT PORTFOLIO - 105.8%

(Cost $165,789,137)

161,029,669

NET OTHER ASSETS - (5.8)%

(8,892,495)

NET ASSETS - 100%

$ 152,137,174

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Includes investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 60,857

Fidelity Securities Lending Cash Central Fund

56,422

Total

$ 117,279

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $7,837,041 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Growth Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $7,883,516) - See accompanying schedule:

Unaffiliated issuers (cost $156,064,347)

$ 151,304,879

Fidelity Central Funds (cost $9,724,790)

9,724,790

Total Investments (cost $165,789,137)

$ 161,029,669

Receivable for investments sold

18,559,475

Receivable for fund shares sold

222,238

Dividends receivable

82,079

Distributions receivable from Fidelity Central Funds

4,743

Prepaid expenses

514

Receivable from investment adviser for expense reductions

25,900

Total assets

179,924,618

Liabilities

Payable for investments purchased

$ 19,486,946

Payable for fund shares redeemed

257,786

Accrued management fee

52,051

Distribution fees payable

1,918

Other affiliated payables

48,536

Other payables and accrued expenses

52,707

Collateral on securities loaned, at value

7,887,500

Total liabilities

27,787,444

Net Assets

$ 152,137,174

Net Assets consist of:

Paid in capital

$ 162,808,253

Accumulated undistributed net realized gain (loss) on investments

(5,911,611)

Net unrealized appreciation (depreciation) on investments

(4,759,468)

Net Assets

$ 152,137,174

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($1,302,425 ÷ 132,172 shares)

$ 9.85

Maximum offering price per share (100/94.25 of $9.85)

$ 10.45

Class T:
Net Asset Value
and redemption price per share ($1,097,491 ÷ 111,448 shares)

$ 9.85

Maximum offering price per share (100/96.50 of $9.85)

$ 10.21

Class B:
Net Asset Value
and offering price per share ($543,149 ÷ 55,237 shares)A

$ 9.83

Class C:
Net Asset Value
and offering price per share ($944,595 ÷ 96,233 shares)A

$ 9.82

Large Cap Growth:
Net Asset Value
, offering price and redemption price per share ($147,863,609 ÷ 14,947,307 shares)

$ 9.89

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($385,905 ÷ 39,068 shares)

$ 9.88

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Large Cap Growth Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 1,475,584

Interest

922

Income from Fidelity Central Funds (including $56,422 from security lending)

117,279

Total income

1,593,785

Expenses

Management fee
Basic fee

$ 979,536

Performance adjustment

37,062

Transfer agent fees

534,768

Distribution fees

10,735

Accounting and security lending fees

70,624

Custodian fees and expenses

18,147

Independent trustees' compensation

638

Registration fees

103,527

Audit

55,247

Legal

928

Miscellaneous

11,842

Total expenses before reductions

1,823,054

Expense reductions

(97,947)

1,725,107

Net investment income (loss)

(131,322)

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

10,237,592

Change in net unrealized appreciation (depreciation) on investment securities

(21,747,840)

Net gain (loss)

(11,510,248)

Net increase (decrease) in net assets resulting from operations

$ (11,641,570)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ (131,322)

$ 26,323

Net realized gain (loss)

10,237,592

4,571,573

Change in net unrealized appreciation (depreciation)

(21,747,840)

(297,654)

Net increase (decrease) in net assets resulting from operations

(11,641,570)

4,300,242

Distributions to shareholders from net investment income

-

(137,207)

Distributions to shareholders from net realized gain

(18,572,404)

(3,644,497)

Total distributions

(18,572,404)

(3,781,704)

Share transactions - net increase (decrease)

(1,164,336)

25,478,221

Redemption fees

-

5,257

Total increase (decrease) in net assets

(31,378,310)

26,002,016

Net Assets

Beginning of period

183,515,484

157,513,468

End of period

$ 152,137,174

$ 183,515,484

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.03)

Net realized and unrealized gain (loss)

(.72)

Total from investment operations

(.75)

Distributions from net realized gain

(1.25)

Net asset value, end of period

$ 9.85

Total Return B,C,D

(6.99)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.20% A

Expenses net of fee waivers, if any

1.20% A

Expenses net of all reductions

1.20% A

Net investment income (loss)

(.29)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,302

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.06)

Net realized and unrealized gain (loss)

(.69)

Total from investment operations

(.75)

Distributions from net realized gain

(1.25)

Net asset value, end of period

$ 9.85

Total Return B,C,D

(7.05)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.47% A

Expenses net of fee waivers, if any

1.47% A

Expenses net of all reductions

1.47% A

Net investment income (loss)

(.56)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,097

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.12)

Net realized and unrealized gain (loss)

(.70)

Total from investment operations

(.82)

Distributions from net realized gain

(1.20)

Net asset value, end of period

$ 9.83

Total Return B,C,D

(7.62)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.99% A

Expenses net of fee waivers, if any

1.99% A

Expenses net of all reductions

1.99% A

Net investment income (loss)

(1.07)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 543

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net investment income (loss) E

(.11)

Net realized and unrealized gain (loss)

(.70)

Total from investment operations

(.81)

Distributions from net realized gain

(1.22)

Net asset value, end of period

$ 9.82

Total Return B,C,D

(7.54)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.96% A

Expenses net of fee waivers, if any

1.96% A

Expenses net of all reductions

1.96% A

Net investment income (loss)

(1.05)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 945

Portfolio turnover rate G

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Large Cap Growth

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 11.92

$ 11.82

$ 10.17

$ 9.21

$ 6.93

Income from Investment Operations

Net investment income (loss) B

(.01)

- H

.02 E

(.01) F

(.01)

Net realized and unrealized gain (loss)

(.77)

.37

1.87

.97

2.29

Total from investment operations

(.78)

.37

1.89

.96

2.28

Distributions from net investment income

-

(.01)

-

-

-

Distributions from net realized gain

(1.25)

(.26)

(.24)

-

-

Total distributions

(1.25)

(.27)

(.24)

-

-

Redemption fees added to paid in capital B

-

- H,I

- H

- H

- H

Net asset value, end of period

$ 9.89

$ 11.92

$ 11.82

$ 10.17

$ 9.21

Total Return A

(7.26)%

3.20%

18.66%

10.42%

32.90%

Ratios to Average Net Assets C,G

Expenses before reductions

1.03%

1.10%

1.12%

1.30%

1.53%

Expenses net of fee waivers, if any

.99%

1.00%

1.00%

1.20%

1.20%

Expenses net of all reductions

.98%

.99%

.94%

1.13%

1.18%

Net investment income (loss)

(.07)%

.02%

.15% E

(.07)% F

(.15)%

Supplemental Data

Net assets, end of period (000 omitted)

$ 147,864

$ 183,515

$ 157,513

$ 49,453

$ 23,079

Portfolio turnover rate D

428%

189%

268%

274%

81%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.07)%.

F Investment income per share reflects a special dividend which amounted to $.02 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.32)%.

G Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

H Amount represents less than $.01 per share.

I The redemption fee was eliminated during the year ended January 31, 2007.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 11.85

Income from Investment Operations

Net realized and unrealized gain (loss)

(.70)

Distributions from net realized gain

(1.27)

Net asset value, end of period

$ 9.88

Total Return B,C

(6.64)%

Ratios to Average Net Assets D,E

Expenses before reductions

.88% A

Expenses net of fee waivers, if any

.88% A

Expenses net of all reductions

.88% A

Net investment income (loss)

.03% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 386

Portfolio turnover rate F

428%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Advisor Mid Cap Growth Fund

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the class' dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table does not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Life of
fund
A

Institutional Class B

-9.62%

13.21%

5.09%

A From November 15, 2001.

B The initial offering of Institutional Class shares took place on February 13, 2007. Returns prior to February 13, 2007 are those of Mid Cap Growth, the original class of the fund.

$10,000 Over Life of Fund

Let's say hypothetically that $10,000 was invested in Fidelity Advisor Mid Cap Growth Fund - Institutional Class on November 15, 2001, when the fund started. The chart shows how the value of your investment would have changed, and also shows how the Russell Midcap® Growth Index performed over the same period. The initial offering of Institutional Class took place on February 13, 2007. See above for additional information regarding the performance of Institutional Class.



Annual Report

Fidelity Advisor Mid Cap Growth Fund

Management's Discussion of Fund Performance

Comments from Patrick Venanzi, Portfolio Manager of Fidelity Advisor Mid Cap Growth Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

The fund's Class A, Class T, Class B and Class C shares (excluding sales charges) lagged behind the -1.03% return for the Russell Midcap® Growth Index. (For specific class-level results, please refer to the performance section of this report.) Poor stock selection in consumer discretionary, industrials, materials and consumer staples hurt the most. While the fund struggled to keep pace with the index for most of the period, stock selection in energy and health care helped limit some of the downside. Within industrials, the fund's overweighting in transportation stocks, specifically in the airlines, detracted from performance. Our holdings in US Airways and AMR Corp., the parent company of American Airlines, suffered from rising energy costs. Overweighted holdings in the consumer durables and apparel segment - primarily the homebuilders - also took a bite out of fund performance. D.R. Horton, Ryland Group, KB Home and Lennar were particularly hard hit as the U.S. residential real estate market continued to slow down. I sold D.R. Horton, Ryland Group and Lennar. On the positive side, energy services company National Oilwell Varco was the fund's biggest contributor. Another contributor was the fund's out-of-index position in consumer electronics giant Apple, which continued to gain from strong sales of its iPod, iPhone and iMac product lines. I sold the fund's position in Apple to lock in profits. Heavy-lifting equipment and crane manufacturer Manitowoc advanced due to the continuing worldwide build-out of commercial and industrial infrastructure. I also sold this position by the end of the period. BE Aerospace, which makes products for aircraft cabin interiors, also contributed, as investors expressed confidence in the company's new contracts to supply airline carriers that are upgrading their fleets. Lastly, MEMC Electronic Materials, which produces silicon wafers, helped as well.

For the year, the fund's Institutional Class shares lagged behind the -1.03% return for the Russell Midcap® Growth Index. (For specific class-level results, please refer to the performance section of this report.) Poor stock selection in consumer discretionary, industrials, materials and consumer staples hurt the most. While the fund struggled to keep pace with the index for most of the period, stock selection in energy and health care helped limit some of the downside. Within industrials, the fund's overweighting in transportation stocks, specifically in the airlines, detracted from performance. Our holdings in US Airways and AMR Corp., the parent company of American Airlines, suffered from rising energy costs. Overweighted holdings in the consumer durables and apparel segment - primarily the homebuilders - also took a bite out of fund performance. D.R. Horton, Ryland Group, KB Home and Lennar were particularly hard hit as the U.S. residential real estate market continued to slow down. I sold D.R. Horton, Ryland Group and Lennar. On the positive side, energy services company National Oilwell Varco was the fund's biggest contributor. Another contributor was the fund's out-of-index position in consumer electronics giant Apple, which continued to gain from strong sales of its iPod, iPhone and iMac product lines. I sold the fund's position in Apple to lock in profits. Heavy-lifting equipment and crane manufacturer Manitowoc advanced due to the continuing worldwide build-out of commercial and industrial infrastructure. I also sold this position by the end of the period. BE Aerospace, which makes products for aircraft cabin interiors, also contributed, as investors expressed confidence in the company's new contracts to supply airline carriers that are upgrading their fleets. Lastly, MEMC Electronic Materials, which produces silicon wafers, helped as well.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Fidelity Mid Cap Growth Fund

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

PPL Corp.

1.4

0.7

Southwestern Energy Co.

1.4

0.3

Constellation Energy Group, Inc.

1.3

1.0

National Oilwell Varco, Inc.

1.3

3.9

Align Technology, Inc.

1.1

0.0

Centex Corp.

1.1

0.0

TranS1, Inc.

1.0

0.0

Polycom, Inc.

0.9

0.0

Intuitive Surgical, Inc.

0.9

0.0

Northwest Airlines Corp.

0.9

0.0

11.3

Top Five Market Sectors as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Information Technology

19.8

19.0

Industrials

17.1

16.1

Health Care

16.6

12.9

Consumer Discretionary

14.1

17.6

Energy

11.6

10.9

Asset Allocation (% of fund's net assets)

As of January 31, 2008*

As of July 31, 2007**

Stocks 100.2%

Stocks 97.8%

Short-Term
Investments and
Net Other Assets(dagger) (0.2)%

Short-Term
Investments and
Net Other Assets 2.2%

* Foreign
investments

11.7%

**Foreign investments

2.1%

(dagger) Short-Term Investments
and Net Other Assets
are not included in the pie chart.

Annual Report

Fidelity Mid Cap Growth Fund

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 100.2%

Shares

Value

CONSUMER DISCRETIONARY - 14.1%

Automobiles - 0.3%

Renault SA

7,300

$ 833,337

Diversified Consumer Services - 1.1%

American Public Education, Inc.

7,200

283,752

Apollo Group, Inc. Class A (non-vtg.) (a)

8,200

653,868

DeVry, Inc.

19,000

1,048,610

Regis Corp.

56,400

1,428,612

3,414,842

Hotels, Restaurants & Leisure - 2.1%

Bally Technologies, Inc. (a)

20,200

962,328

Boyd Gaming Corp.

66,400

1,774,872

Buffalo Wild Wings, Inc. (a)

8,400

211,428

Burger King Holdings, Inc.

54,789

1,443,690

Gaylord Entertainment Co. (a)

29,700

866,943

Papa John's International, Inc. (a)

26,100

660,069

Peet's Coffee & Tea, Inc. (a)

7,000

153,580

Starwood Hotels & Resorts Worldwide, Inc.

5,400

244,350

6,317,260

Household Durables - 2.1%

Centex Corp. (d)

116,700

3,241,926

KB Home

100

2,750

Newell Rubbermaid, Inc.

15,100

364,212

Pulte Homes, Inc.

33,500

547,390

Whirlpool Corp.

25,700

2,187,327

6,343,605

Internet & Catalog Retail - 1.2%

Blue Nile, Inc. (a)

17,400

961,350

Expedia, Inc. (a)

10,600

244,012

Priceline.com, Inc. (a)

21,099

2,289,663

3,495,025

Leisure Equipment & Products - 0.7%

Hasbro, Inc.

12,400

322,028

Leapfrog Enterprises, Inc. Class A (a)

36,300

236,313

Mattel, Inc.

53,533

1,124,728

Nikon Corp.

14,000

381,830

2,064,899

Media - 2.7%

Cablevision Systems Corp. - NY Group Class A (a)

30,300

711,444

DISH Network Corp. Class A (a)

13,057

368,730

E.W. Scripps Co. Class A

35,800

1,457,776

Grupo Televisa SA de CV (CPO) sponsored ADR

94,000

2,095,260

Regal Entertainment Group Class A

87,100

1,614,834

SR Teleperformance SA

44,300

1,386,369

The Walt Disney Co.

11,700

350,181

Time Warner, Inc.

21,600

339,984

8,324,578

Shares

Value

Multiline Retail - 0.1%

Dollar Tree Stores, Inc. (a)

14,300

$ 400,543

Specialty Retail - 3.2%

Abercrombie & Fitch Co. Class A

5,100

406,419

Advance Auto Parts, Inc.

30,500

1,088,240

Asbury Automotive Group, Inc.

13,100

185,758

AutoZone, Inc. (a)

8,800

1,063,744

Gamestop Corp. Class A (a)

5,400

279,342

Group 1 Automotive, Inc.

8,700

230,028

Home Depot, Inc.

19,700

604,199

Lumber Liquidators, Inc.

3,400

30,260

OfficeMax, Inc.

49,800

1,233,546

Ross Stores, Inc.

36,300

1,058,145

Tween Brands, Inc. (a)(d)

44,700

1,431,741

Urban Outfitters, Inc. (a)

28,800

835,200

Williams-Sonoma, Inc.

49,500

1,330,560

Zumiez, Inc. (a)

3,500

67,305

9,844,487

Textiles, Apparel & Luxury Goods - 0.6%

Coach, Inc. (a)

13,000

416,650

Deckers Outdoor Corp. (a)

2,800

339,472

Lululemon Athletica, Inc.

15,700

532,073

Under Armour, Inc. Class A (sub. vtg.) (a)

16,000

644,000

1,932,195

TOTAL CONSUMER DISCRETIONARY

42,970,771

CONSUMER STAPLES - 3.4%

Beverages - 0.1%

Coca-Cola Femsa SA de CV sponsored ADR

7,200

338,040

Food & Staples Retailing - 1.3%

CVS Caremark Corp.

14,600

570,422

Performance Food Group Co. (a)

48,000

1,518,240

Safeway, Inc.

25,200

780,948

Sysco Corp.

42,100

1,223,005

4,092,615

Food Products - 1.5%

Campbell Soup Co.

17,300

546,853

Marine Harvest ASA (a)

2,600,000

1,400,493

McCormick & Co., Inc. (non-vtg.)

55,300

1,864,716

Seaboard Corp.

370

475,543

Wm. Wrigley Jr. Co.

3,000

172,290

4,459,895

Household Products - 0.4%

Energizer Holdings, Inc. (a)

13,300

1,245,146

Personal Products - 0.1%

Estee Lauder Companies, Inc. Class A

8,900

375,580

TOTAL CONSUMER STAPLES

10,511,276

Common Stocks - continued

Shares

Value

ENERGY - 11.6%

Energy Equipment & Services - 5.0%

Atwood Oceanics, Inc. (a)

21,000

$ 1,744,890

Cameron International Corp. (a)

31,700

1,276,242

Diamond Offshore Drilling, Inc.

16,700

1,885,931

Grey Wolf, Inc. (a)

297,600

1,773,696

Nabors Industries Ltd. (a)

35,000

952,700

National Oilwell Varco, Inc. (a)

65,120

3,922,178

North American Energy Partners, Inc. (a)

14,200

170,116

Ocean RIG ASA (a)(d)

114,500

787,337

Smith International, Inc.

23,900

1,295,619

Transocean, Inc. (a)

10,961

1,343,819

15,152,528

Oil, Gas & Consumable Fuels - 6.6%

Alpha Natural Resources, Inc. (a)

21,500

719,390

Arch Coal, Inc.

15,700

690,800

Boardwalk Pipeline Partners, LP

5,700

177,156

Canadian Natural Resources Ltd.

10,100

646,227

Chesapeake Energy Corp.

46,600

1,734,918

CONSOL Energy, Inc.

35,300

2,576,900

Copano Energy LLC

4,900

173,166

Denbury Resources, Inc. (a)

65,700

1,662,210

Energy Transfer Equity LP

5,200

171,652

Foundation Coal Holdings, Inc.

7,000

366,100

Frontier Oil Corp.

38,100

1,343,787

Holly Corp.

28,601

1,384,860

OPTI Canada, Inc. (a)

80,400

1,325,913

Petrobank Energy & Resources Ltd. (a)

7,000

351,761

Southwestern Energy Co. (a)

76,100

4,254,751

Tesoro Corp.

22,400

874,720

Valero Energy Corp.

100

5,919

Western Refining, Inc.

44,460

949,221

Williams Companies, Inc.

27,100

866,387

20,275,838

TOTAL ENERGY

35,428,366

FINANCIALS - 9.2%

Capital Markets - 2.6%

Franklin Resources, Inc.

23,600

2,459,828

KBW, Inc. (a)

6,300

187,173

Legg Mason, Inc.

14,500

1,044,000

MF Global Ltd.

73,200

2,199,660

SEI Investments Co.

67,400

1,866,980

T. Rowe Price Group, Inc.

3,300

166,947

7,924,588

Commercial Banks - 1.5%

Bank of Yokohama Ltd.

84,000

544,305

Chiba Bank Ltd.

52,000

384,877

Chuo Mitsui Trust Holdings, Inc.

55,000

377,598

Mitsubishi UFJ Financial Group, Inc.

70,000

691,600

Shinsei Bank Ltd.

117,000

534,769

Shares

Value

Shizuoka Bank Ltd.

40,000

$ 439,763

Sumitomo Mitsui Financial Group, Inc.

100

788,113

Tokyo Tomin Bank Ltd.

20,600

572,491

UMB Financial Corp.

7,600

320,188

4,653,704

Consumer Finance - 0.4%

Cash America International, Inc.

37,100

1,206,121

Diversified Financial Services - 1.5%

CME Group, Inc.

800

495,120

IntercontinentalExchange, Inc. (a)

9,400

1,315,624

MSCI, Inc. Class A

11,800

389,046

NYMEX Holdings, Inc.

18,400

2,116,000

Osaka Securities Exchange Co. Ltd.

64

346,092

4,661,882

Insurance - 0.9%

Assurant, Inc.

19,400

1,258,866

CNA Financial Corp.

11,480

390,205

CNinsure, Inc. ADR

44,000

472,560

LandAmerica Financial Group, Inc.

4,900

255,584

Philadelphia Consolidated Holdings Corp. (a)

9,600

343,680

Stewart Information Services Corp.

1,100

37,653

2,758,548

Real Estate Investment Trusts - 1.5%

Annaly Capital Management, Inc.

124,000

2,445,280

MFA Mortgage Investments, Inc.

189,300

1,930,860

4,376,140

Thrifts & Mortgage Finance - 0.8%

People's United Financial, Inc.

58,700

991,443

Washington Federal, Inc.

55,900

1,365,078

2,356,521

TOTAL FINANCIALS

27,937,504

HEALTH CARE - 16.6%

Biotechnology - 3.5%

Alexion Pharmaceuticals, Inc. (a)

18,500

1,208,420

Amylin Pharmaceuticals, Inc. (a)

74,607

2,212,098

Biogen Idec, Inc. (a)

26,400

1,609,080

Cephalon, Inc. (a)

11,300

741,619

Genentech, Inc. (a)

8,000

561,520

Gilead Sciences, Inc. (a)

41,362

1,889,830

Theravance, Inc. (a)(d)

127,038

2,506,460

10,729,027

Health Care Equipment & Supplies - 5.8%

Abiomed, Inc. (a)

25,000

377,500

Align Technology, Inc. (a)(d)

283,984

3,345,332

American Medical Systems Holdings, Inc. (a)

90,000

1,286,100

Conceptus, Inc. (a)

158,500

2,580,380

Hillenbrand Industries, Inc.

13,300

687,876

Hologic, Inc. (a)

14,000

901,040

Common Stocks - continued

Shares

Value

HEALTH CARE - continued

Health Care Equipment & Supplies - continued

Intuitive Surgical, Inc. (a)

11,000

$ 2,794,000

Inverness Medical Innovations, Inc. (a)

3,000

135,150

Mindray Medical International Ltd. sponsored ADR

10,000

341,000

NxStage Medical, Inc. (a)

14,100

171,174

Regeneration Technologies, Inc. (a)

215,000

1,720,000

TranS1, Inc. (d)

213,000

3,107,670

Varian Medical Systems, Inc. (a)

5,300

275,547

17,722,769

Health Care Providers & Services - 2.4%

athenahealth, Inc. (d)

49,400

1,551,160

Brookdale Senior Living, Inc. (d)

11,000

245,520

Coventry Health Care, Inc. (a)

5,500

311,190

Express Scripts, Inc. (a)

28,600

1,930,214

Health Net, Inc. (a)

10,300

478,847

Henry Schein, Inc. (a)

5,473

318,145

MWI Veterinary Supply, Inc. (a)

8,000

305,840

Pediatrix Medical Group, Inc. (a)

24,587

1,674,129

Psychiatric Solutions, Inc. (a)

5,900

178,003

VCA Antech, Inc. (a)

10,200

394,332

7,387,380

Health Care Technology - 1.0%

Cerner Corp. (a)

43,200

2,263,680

MedAssets, Inc.

7,900

159,185

TriZetto Group, Inc. (a)

38,900

759,328

3,182,193

Life Sciences Tools & Services - 2.0%

Affymetrix, Inc. (a)(d)

60,600

1,215,636

Applera Corp. - Applied Biosystems Group

30,000

945,900

Charles River Laboratories International, Inc. (a)

19,700

1,223,370

Covance, Inc. (a)

1,800

149,688

PAREXEL International Corp. (a)

10,000

544,100

Pharmaceutical Product Development, Inc.

30,200

1,309,472

Thermo Fisher Scientific, Inc. (a)

12,770

657,527

6,045,693

Pharmaceuticals - 1.9%

Allergan, Inc.

20,400

1,370,676

BioForm Medical, Inc. (d)

186,481

1,305,367

Perrigo Co.

15,700

484,188

Renovo Group PLC (a)

342,000

920,384

Teva Pharmaceutical Industries Ltd. sponsored ADR

22,300

1,026,692

XenoPort, Inc. (a)

8,600

527,696

5,635,003

TOTAL HEALTH CARE

50,702,065

Shares

Value

INDUSTRIALS - 17.1%

Aerospace & Defense - 1.0%

BE Aerospace, Inc. (a)

21,444

$ 827,953

Goodrich Corp.

19,900

1,244,745

Spirit AeroSystems Holdings, Inc.
Class A (a)

31,700

875,554

2,948,252

Air Freight & Logistics - 0.2%

United Parcel Service, Inc. Class B

6,400

468,224

Airlines - 2.3%

AMR Corp. (a)

103,000

1,435,820

Continental Airlines, Inc. Class B (a)

63,900

1,738,719

Northwest Airlines Corp. (a)

146,700

2,744,757

US Airways Group, Inc. (a)

89,200

1,231,852

7,151,148

Building Products - 0.3%

Ameron International Corp.

8,000

718,800

Universal Forest Products, Inc.

6,000

217,200

936,000

Commercial Services & Supplies - 5.1%

Advisory Board Co. (a)

2,600

165,672

American Reprographics Co. (a)

46,200

726,726

Copart, Inc. (a)

33,984

1,389,266

Corporate Executive Board Co.

9,400

540,970

Corrections Corp. of America (a)

94,182

2,499,590

Fuel Tech, Inc. (a)

35,600

677,824

GeoEye, Inc. (a)

15,000

524,400

IHS, Inc. Class A (a)

11,500

712,310

InnerWorkings, Inc. (a)

12,000

166,200

Manpower, Inc.

45,400

2,554,204

Republic Services, Inc.

56,600

1,698,000

The Geo Group, Inc. (a)

54,600

1,306,032

Waste Connections, Inc. (a)

45,600

1,329,696

Waste Management, Inc.

34,400

1,115,936

15,406,826

Construction & Engineering - 0.7%

Chicago Bridge & Iron Co. NV
(NY Shares)

4,900

218,001

Fluor Corp.

7,500

912,525

Grupo Acciona SA

2,000

509,808

Quanta Services, Inc. (a)

20,000

438,400

2,078,734

Electrical Equipment - 3.9%

Belden, Inc.

7,000

296,100

Energy Conversion Devices, Inc. (a)(d)

57,020

1,316,022

Evergreen Solar, Inc. (a)

154,800

1,887,012

First Solar, Inc. (a)

10,300

1,872,231

JA Solar Holdings Co. Ltd. ADR

16,000

813,280

Nexans SA

19,800

2,188,427

Q-Cells AG (a)

6,000

564,589

Sunpower Corp. Class A (a)(d)

35,400

2,445,786

Common Stocks - continued

Shares

Value

INDUSTRIALS - continued

Electrical Equipment - continued

Superior Essex, Inc. (a)

13,100

$ 315,055

Vestas Wind Systems AS (a)

2,800

271,999

11,970,501

Machinery - 1.3%

AGCO Corp. (a)

20,600

1,240,532

GEA Group AG (a)

32,000

988,991

Ingersoll-Rand Co. Ltd. Class A

18,000

711,360

Terex Corp. (a)

17,100

1,004,796

3,945,679

Marine - 0.4%

Ultrapetrol (Bahamas) Ltd. (a)

70,100

1,125,105

Road & Rail - 1.7%

J.B. Hunt Transport Services, Inc. (d)

56,100

1,744,710

Knight Transportation, Inc.

102,900

1,765,764

Landstar System, Inc.

36,500

1,826,095

5,336,569

Trading Companies & Distributors - 0.2%

Genesis Lease Ltd. ADR

31,600

635,160

TOTAL INDUSTRIALS

52,002,198

INFORMATION TECHNOLOGY - 19.8%

Communications Equipment - 2.0%

Ciena Corp. (a)

15,000

406,950

CommScope, Inc. (a)

11,700

518,895

Corning, Inc.

22,000

529,540

Juniper Networks, Inc. (a)

64,700

1,756,605

Polycom, Inc. (a)

111,000

2,802,750

6,014,740

Computers & Peripherals - 1.7%

Brocade Communications Systems, Inc. (a)

226,602

1,561,288

Diebold, Inc.

28,644

741,307

NCR Corp. (a)

61,600

1,323,168

Western Digital Corp. (a)

54,600

1,444,170

5,069,933

Electronic Equipment & Instruments - 1.6%

Agilent Technologies, Inc. (a)

8,700

295,017

Avnet, Inc. (a)

17,300

616,053

Bell Microproducts, Inc. (a)

29,818

156,843

Comverge, Inc.

6,100

113,704

Flextronics International Ltd. (a)

51,100

597,870

Ingram Micro, Inc. Class A (a)

30,700

545,846

Itron, Inc. (a)

24,000

1,977,600

Jabil Circuit, Inc.

47,300

626,725

4,929,658

Internet Software & Services - 2.8%

Akamai Technologies, Inc. (a)

16,800

507,360

DealerTrack Holdings, Inc. (a)

23,600

636,256

Shares

Value

Dice Holdings, Inc.

258,600

$ 1,649,868

DivX, Inc. (a)

50,000

712,500

Internet Brands, Inc. Class A

30,979

239,158

Omniture, Inc. (a)

85,700

2,118,504

Online Resources Corp. (a)

89,800

916,858

The Knot, Inc. (a)

49,700

727,608

ValueClick, Inc. (a)

44,100

962,703

8,470,815

IT Services - 1.1%

Cognizant Technology Solutions Corp. Class A (a)

72,700

2,028,330

CyberSource Corp. (a)

38,000

636,500

Mastercard, Inc. Class A

3,800

786,600

3,451,430

Semiconductors & Semiconductor Equipment - 7.4%

Altera Corp.

93,000

1,570,770

Applied Materials, Inc.

125,400

2,247,168

ARM Holdings PLC sponsored ADR

23,600

167,088

ASML Holding NV (NY Shares) (a)

20,000

531,800

Broadcom Corp. Class A (a)

66,500

1,468,320

Cypress Semiconductor Corp. (a)

62,200

1,321,750

FormFactor, Inc. (a)

21,063

510,146

Himax Technologies, Inc. sponsored ADR

285,100

1,405,543

Hittite Microwave Corp. (a)

12,400

493,768

Intersil Corp. Class A

31,400

723,142

Lam Research Corp. (a)

46,500

1,785,135

LDK Solar Co. Ltd. Sponsored ADR (d)

19,800

697,950

Marvell Technology Group Ltd. (a)

122,180

1,450,277

MEMC Electronic Materials, Inc. (a)

18,500

1,322,010

Microchip Technology, Inc.

35,500

1,132,805

Miraial Co. Ltd.

15,900

386,547

National Semiconductor Corp.

53,000

976,790

Skyworks Solutions, Inc. (a)(d)

124,700

1,003,835

Teradyne, Inc. (a)

5,400

59,238

Varian Semiconductor Equipment Associates, Inc. (a)

63,800

2,054,998

Xilinx, Inc.

61,200

1,338,444

22,647,524

Software - 3.2%

Activision, Inc. (a)

25,100

649,337

Citrix Systems, Inc. (a)

12,000

415,440

CompuGROUP Holding AG (a)

33,000

622,352

Concur Technologies, Inc. (a)

53,300

1,868,698

FactSet Research Systems, Inc.

6,000

335,580

Jack Henry & Associates, Inc.

53,300

1,310,114

Microsoft Corp.

20,800

678,080

Nintendo Co. Ltd.

400

197,600

Nuance Communications, Inc. (a)

87,000

1,382,430

Shanda Interactive Entertainment Ltd. sponsored ADR (a)

17,900

492,071

SPSS, Inc. (a)

9,000

297,450

Common Stocks - continued

Shares

Value

INFORMATION TECHNOLOGY - continued

Software - continued

Take-Two Interactive Software, Inc. (a)(d)

61,650

$ 1,013,526

THQ, Inc. (a)

33,800

608,738

9,871,416

TOTAL INFORMATION TECHNOLOGY

60,455,516

MATERIALS - 3.1%

Chemicals - 0.2%

FMC Corp.

11,300

600,708

Construction Materials - 0.5%

Martin Marietta Materials, Inc. (d)

13,800

1,693,536

Containers & Packaging - 0.4%

Smurfit-Stone Container Corp. (a)

39,600

375,804

Temple-Inland, Inc.

39,600

742,500

1,118,304

Metals & Mining - 1.9%

Allegheny Technologies, Inc.

19,542

1,375,757

Barrick Gold Corp.

6,900

356,224

Century Aluminum Co. (a)

14,000

727,860

Newmont Mining Corp.

10,000

543,400

Nucor Corp.

22,200

1,283,160

Randgold Resources Ltd. sponsored ADR

5,800

276,718

RTI International Metals, Inc. (a)

23,140

1,278,485

5,841,604

Paper & Forest Products - 0.1%

Glatfelter

12,400

179,304

TOTAL MATERIALS

9,433,456

TELECOMMUNICATION SERVICES - 1.3%

Wireless Telecommunication Services - 1.3%

American Tower Corp. Class A (a)

52,705

1,978,019

NII Holdings, Inc. (a)

48,691

2,077,158

4,055,177

UTILITIES - 4.0%

Electric Utilities - 2.1%

Allegheny Energy, Inc.

17,600

964,304

Enernoc, Inc.

14,300

504,075

Shares

Value

Entergy Corp.

4,500

$ 486,810

PPL Corp.

89,800

4,393,015

6,348,204

Independent Power Producers & Energy Traders - 1.9%

AES Corp. (a)

79,400

1,514,952

Clipper Windpower PLC (a)

2,400

28,693

Constellation Energy Group, Inc.

42,800

4,021,488

Ormat Technologies, Inc.

3,800

165,186

5,730,319

TOTAL UTILITIES

12,078,523

TOTAL COMMON STOCKS

(Cost $305,398,397)

305,574,852

Money Market Funds - 4.3%

Fidelity Cash Central Fund, 3.79% (b)

2,288,816

2,288,816

Fidelity Securities Lending Cash Central Fund, 3.84% (b)(c)

10,764,675

10,764,675

TOTAL MONEY MARKET FUNDS

(Cost $13,053,491)

13,053,491

TOTAL INVESTMENT PORTFOLIO - 104.5%

(Cost $318,451,888)

318,628,343

NET OTHER ASSETS - (4.5)%

(13,583,264)

NET ASSETS - 100%

$ 305,045,079

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

(c) Investment made with cash collateral received from securities on loan.

(d) Security or a portion of the security is on loan at period end.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned

Fidelity Cash Central Fund

$ 442,525

Fidelity Securities Lending Cash Central Fund

313,679

Total

$ 756,204

Other Information

Distribution of investments by country of issue, as a percentage of total net assets, is as follows:

United States of America

88.3%

Japan

1.8%

Bermuda

1.7%

France

1.5%

Cayman Islands

1.3%

Others (individually less than 1%)

5.4%

100.0%

Income Tax Information

The fund intends to elect to defer to its fiscal year ending January 31, 2009 approximately $18,706,577 of losses recognized during the period November 1, 2007 to January 31, 2008.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Growth Fund

Financial Statements

Statement of Assets and Liabilities

January 31, 2008

Assets

Investment in securities, at value (including securities loaned of $10,830,743) - See accompanying schedule:

Unaffiliated issuers (cost $305,398,397)

$ 305,574,852

Fidelity Central Funds (cost $13,053,491)

13,053,491

Total Investments (cost $318,451,888)

$ 318,628,343

Cash

944,763

Receivable for investments sold

60,992,695

Receivable for fund shares sold

386,427

Dividends receivable

50,720

Distributions receivable from Fidelity Central Funds

53,069

Prepaid expenses

1,182

Other receivables

2,063

Total assets

381,059,262

Liabilities

Payable for investments purchased

$ 23,187,980

Payable for fund shares redeemed

41,783,947

Accrued management fee

108,332

Distribution fees payable

1,571

Other affiliated payables

110,841

Other payables and accrued expenses

56,837

Collateral on securities loaned, at value

10,764,675

Total liabilities

76,014,183

Net Assets

$ 305,045,079

Net Assets consist of:

Paid in capital

$ 319,016,951

Accumulated undistributed net realized gain (loss) on investments and foreign currency transactions

(14,150,236)

Net unrealized appreciation (depreciation) on investments and assets and liabilities in foreign currencies

178,364

Net Assets

$ 305,045,079

Statement of Assets and Liabilities - continued

January 31, 2008

Calculation of Maximum Offering Price

Class A:
Net Asset Value
and redemption price per share ($1,936,249 ÷ 158,895 shares)

$ 12.19

Maximum offering price per share (100/94.25 of $12.19)

$ 12.93

Class T:
Net Asset Value
and redemption price per share ($591,188 ÷ 48,688 shares)

$ 12.14

Maximum offering price per share (100/96.50 of $12.14)

$ 12.58

Class B:
Net Asset Value
and offering price per share ($413,537 ÷ 34,216 shares)A

$ 12.09

Class C:
Net Asset Value
and offering price per share ($697,298 ÷ 57,672 shares)A

$ 12.09

Mid Cap Growth:
Net Asset Value
, offering price and redemption price per share ($301,224,741 ÷ 24,671,114 shares)

$ 12.21

Institutional Class:
Net Asset Value
, offering price and redemption price per share ($182,066 ÷ 14,896 shares)

$ 12.22

A Redemption price per share is equal to net asset value less any applicable contingent deferred sales charge.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Fidelity Mid Cap Growth Fund
Financial Statements - continued

Statement of Operations

Year ended January 31, 2008

Investment Income

Dividends

$ 2,094,532

Interest

386

Income from Fidelity Central Funds (including $313,679 from security lending)

756,204

Total income

2,851,122

Expenses

Management fee
Basic fee

$ 2,332,314

Performance adjustment

(430,070)

Transfer agent fees

1,202,314

Distribution fees

10,363

Accounting and security lending fees

170,995

Custodian fees and expenses

18,214

Independent trustees' compensation

1,531

Registration fees

100,229

Audit

56,662

Legal

2,290

Miscellaneous

25,324

Total expenses before reductions

3,490,166

Expense reductions

(113,488)

3,376,678

Net investment income (loss)

(525,556)

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

13,581,162

Foreign currency transactions

16,150

Total net realized gain (loss)

13,597,312

Change in net unrealized appreciation (depreciation) on:

Investment securities

(50,635,596)

Assets and liabilities in foreign currencies

1,909

Total change in net unrealized appreciation (depreciation)

(50,633,687)

Net gain (loss)

(37,036,375)

Net increase (decrease) in net assets resulting from operations

$ (37,561,931)

Statement of Changes in Net Assets

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ (525,556)

$ (1,458,396)

Net realized gain (loss)

13,597,312

(4,757,238)

Change in net unrealized appreciation (depreciation)

(50,633,687)

5,161,873

Net increase (decrease) in net assets resulting from operations

(37,561,931)

(1,053,761)

Distributions to shareholders from net realized gain

(21,909,497)

(5,089,264)

Share transactions - net increase (decrease)

(76,821,412)

97,426,141

Redemption fees

26,012

46,437

Total increase (decrease) in net assets

(136,266,828)

91,329,553

Net Assets

Beginning of period

441,311,907

349,982,354

End of period

$ 305,045,079

$ 441,311,907

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class A

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.05)

Net realized and unrealized gain (loss)

(1.30)

Total from investment operations

(1.35)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.19

Total Return B,C,D

(9.95)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.10% A

Expenses net of fee waivers, if any

1.10% A

Expenses net of all reductions

1.10% A

Net investment income (loss)

(.41)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 1,936

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class T

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.09)

Net realized and unrealized gain (loss)

(1.31)

Total from investment operations

(1.40)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.14

Total Return B,C,D

(10.30)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.36% A

Expenses net of fee waivers, if any

1.36% A

Expenses net of all reductions

1.36% A

Net investment income (loss)

(.68)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 591

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the sales charges.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Class B

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.16)

Net realized and unrealized gain (loss)

(1.29)

Total from investment operations

(1.45)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.09

Total Return B,C,D

(10.65)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.85% A

Expenses net of fee waivers, if any

1.85% A

Expenses net of all reductions

1.85% A

Net investment income (loss)

(1.16)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 414

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

Financial Highlights - Class C

Year ended January 31,

2008 H

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net investment income (loss) E

(.15)

Net realized and unrealized gain (loss)

(1.30)

Total from investment operations

(1.45)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.09

Total Return B,C,D

(10.65)%

Ratios to Average Net Assets F,I

Expenses before reductions

1.85% A

Expenses net of fee waivers, if any

1.85% A

Expenses net of all reductions

1.85% A

Net investment income (loss)

(1.16)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 697

Portfolio turnover rate G

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Total returns do not include the effect of the contingent deferred sales charge.

E Calculated based on average shares outstanding during the period.

F Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

G Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

H For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

I Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights - Mid Cap Growth

Year ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 14.31

$ 14.38

$ 11.58

$ 10.63

$ 7.32

Income from Investment Operations

Net investment income (loss) B

(.02)

(.04)

.01 E

(.03) F

(.07)

Net realized and unrealized gain (loss)

(1.29)

.15

3.09

1.15

3.38

Total from investment operations

(1.31)

.11

3.10

1.12

3.31

Distributions from net realized gain

(.79)

(.18)

(.30)

(.17)

-

Redemption fees added to paid in capital B,H

-

-

-

-

-

Net asset value, end of period

$ 12.21

$ 14.31

$ 14.38

$ 11.58

$ 10.63

Total Return A

(9.68)%

.80%

27.15%

10.55%

45.22%

Ratios to Average Net Assets C,G

Expenses before reductions

.83%

1.02%

1.04%

1.02%

1.25%

Expenses net of fee waivers, if any

.81%

1.00%

1.00%

1.02%

1.20%

Expenses net of all reductions

.81%

.99%

.95%

.99%

1.16%

Net investment income (loss)

(.12)%

(.33)%

.07% E

(.31)% F

(.77)%

Supplemental Data

Net assets, end of period (000 omitted)

$ 301,225

$ 441,312

$ 349,982

$ 77,658

$ 60,660

Portfolio turnover rate D

245%

178%

173%

220%

94%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.03 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.18)%.

F Investment income per share reflects a special dividend which amounted to $.01 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been (.44)%.

G Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

H Amount represents less than $.01 per share.

Financial Highlights - Institutional Class

Year ended January 31,

2008 G

Selected Per-Share Data

Net asset value, beginning of period

$ 14.33

Income from Investment Operations

Net realized and unrealized gain (loss) D

(1.32)

Distributions from net realized gain

(.79)

Net asset value, end of period

$ 12.22

Total Return B,C

(9.74)%

Ratios to Average Net Assets E,H

Expenses before reductions

.72% A

Expenses net of fee waivers, if any

.72% A

Expenses net of all reductions

.72% A

Net investment income (loss)

(.03)% A

Supplemental Data

Net assets, end of period (000 omitted)

$ 182

Portfolio turnover rate F

245%

A Annualized

B Total returns for periods of less than one year are not annualized.

C Total returns would have been lower had certain expenses not been reduced during the periods shown.

D Calculated based on average shares outstanding during the period.

E Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

F Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

G For the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

H Expense ratios reflect operating expenses of the class. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the class during periods when reimbursements or reductions occur. Expense ratios before reductions for start-up periods may not be representative of longer-term operating periods. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the class.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Notes to Financial Statements

For the period ended January 31, 2008

1. Organization.

Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund(the Funds) are funds of Fidelity Devonshire Trust(the trust) and are authorized to issue an unlimited number of shares. The trust is registered under the Investment Company Act of 1940, as amended(the 1940 Act), as an open-end management company organized as a Massachusetts business trust. The Funds offer Class A, Class T, Class B, Class C, Institutional Class and Large Cap Value, Mid Cap Value, Large Cap Growth and Mid Cap Growth shares, respectively, each of which has equal rights as to assets and voting privileges. Each class has exclusive voting rights with respect to matters that affect that class. The Funds commenced sale of Class A, Class T, Class B, Class C and Institutional Class shares and the existing class of each Fund was designated Large Cap Value, Mid Cap Value, Large Cap Growth and Mid Cap Growth, respectively, on February 13, 2007. Class B shares will automatically convert to Class A shares after a holding period of seven years from the initial date of purchase. Investment income, realized and unrealized capital gains and losses, the common expenses of the Fund, and certain fund-level expense reductions, if any, are allocated on a pro rata basis to each class based on the relative net assets of each class to the total net assets of the Fund. Each class differs with respect to transfer agent and distribution and service plan fees incurred. Certain expense reductions also differ by class. Certain Funds investments in emerging markets can be subject to social, economic, regulatory, and political uncertainties and can be extremely volatile.

2. Investments in Fidelity Central Funds.

The Funds may invest in Fidelity Central Funds, which are open-end investment companies available only to other investment companies and accounts managed by Fidelity Management & Research Company(FMR) and its affiliates. The Funds' Schedules of Investments list each of the Fidelity Central Funds as an investment of each Fund, but does not include the underlying holdings of each Fidelity Central Fund. As an Investing Fund, the Funds indirectly bear their proportionate share of the expenses of the underlying Fidelity Central Funds.

The Money Market Central Funds seek preservation of capital and current income and are managed by Fidelity Investments Money Management, Inc.(FIMM), an affiliate of FMR.

A complete list of holdings for each Fidelity Central Fund is available upon request or at the SEC's web site at www.sec.gov. In addition, the financial statements of the Fidelity Central Funds, which are not covered by the Funds' Report of Independent Registered Public Accounting Firm, are available on the SEC's web site or upon request.

3. Significant Accounting Policies.

The financial statements have been prepared in conformity with accounting principles generally accepted in the United States of America, which require management to make certain estimates and assumptions at the date of the financial statements. Actual results could differ from those estimates. The following summarizes the significant accounting policies of the Funds:

Security Valuation. Investments are valued and net asset value(NAV) per share is calculated(NAV calculation) as of the close of business of the New York Stock Exchange(NYSE), normally 4:00 p.m. Eastern time. Wherever possible, each Fund uses independent pricing services approved by the Board of Trustees to value their investments.

Equity securities, including restricted securities, for which market quotations are readily available, are valued at the last reported sale price or official closing price as reported by an independent pricing service on the primary market or exchange on which they are traded. In the event there were no sales during the day or closing prices are not available, securities are valued at the last quoted bid price. Investments in open-end mutual funds, including the Fidelity Central Funds, are valued at their closing net asset value each business day. Short-term securities with remaining maturities of sixty days or less for which quotations are not readily available are valued at amortized cost, which approximates value.

When current market prices or quotations are not readily available or do not accurately reflect fair value, valuations may be determined in accordance with procedures adopted by the Board of Trustees. For example, when developments occur between the close of a market and the close of the NYSE that may materially affect the value of some or all of the securities, or when trading in a security is halted, those securities may be fair valued. Factors used in the determination of fair value may include monitoring news to identify significant market or security specific events such as changes in the value of U.S. securities markets, reviewing developments in foreign markets and evaluating the performance of ADRs, futures contracts and exchange-traded funds. Because each Fund's utilization of fair value pricing depends on market activity, the frequency with which fair value pricing is used cannot be predicted and may be utilized to a significant extent. The value of securities used for NAV calculation under fair value pricing may differ from published prices for the same securities.

Foreign Currency. Certain Funds may use foreign currency contracts to facilitate transactions in foreign-denominated securities. Losses from these transactions may arise from changes in the value of the foreign currency or if the counterparties do not perform under the contracts' terms.

Annual Report

Notes to Financial Statements - continued

3. Significant Accounting Policies - continued

Foreign Currency - continued

Foreign-denominated assets, including investment securities, and liabilities are translated into U.S. dollars at the exchange rate at period end. Purchases and sales of investment securities, income and dividends received and expenses denominated in foreign currencies are translated into U.S. dollars at the exchange rate in effect on the transaction date.

The effects of exchange rate fluctuations on investments are included with the net realized and unrealized gain(loss) on investment securities. Other foreign currency transactions resulting in realized and unrealized gain(loss) are disclosed separately.

Investment Transactions and Income. For financial reporting purposes, the Funds' investment holdings and NAV include trades executed through the end of the last business day of the period. The NAV for processing shareholder transactions includes trades executed through the end of the prior business day. Gains and losses on securities sold are determined on the basis of identified cost and may include proceeds received from litigation. Dividend income is recorded on the ex-dividend date, except for certain dividends from foreign securities where the ex-dividend date may have passed, which are recorded as soon as the Funds are informed of the ex-dividend date. Non-cash dividends included in dividend income, if any, are recorded at the fair market value of the securities received. Distributions received on securities that represent a return of capital or capital gain are recorded as a reduction of cost of investments and/or as a realized gain. The Funds estimate the components of distributions received that may be considered return of capital distributions or capital gain distributions. Interest income and distributions from the Fidelity Central Funds are accrued as earned. Interest income includes coupon interest and amortization of premium and accretion of discount on debt securities.

Expenses. Most expenses of the trust can be directly attributed to a fund. Expenses which cannot be directly attributed are apportioned among each Fund in the trust. Expense estimates are accrued in the period to which they relate and adjustments are made when actual amounts are known.

Income Tax Information and Distributions to Shareholders. Each year, each Fund intends to qualify as a regulated investment company by distributing substantially all of its taxable income and realized gains under Subchapter M of the Internal Revenue Code and filing its U.S. federal tax return. As a result, no provision for income taxes is required. Each Fund adopted the provisions of FASB Interpretation No. 48, Accounting for Uncertainties in Income Taxes(FIN 48), on June 29, 2007. FIN 48 sets forth a minimum threshold for financial statement recognition of the benefit of a tax position taken or expected to be taken in a tax return. The implementation of FIN 48 did not result in any unrecognized tax benefits in the accompanying financial statements. Each of the Funds' federal tax returns for the prior three fiscal years remains subject to examination by the Internal Revenue Service. Foreign taxes are provided for based on each Fund's understanding of the tax rules and rates that exist in the foreign markets in which it invests.

Distributions are recorded on the ex-dividend date. Income dividends and capital gain distributions are declared separately for each class. Income and capital gain distributions are determined in accordance with income tax regulations, which may differ from generally accepted accounting principles. In addition, certain Funds will claim a portion of the payment made to redeeming shareholders as a distribution for income tax purposes.

Capital accounts within the financial statements are adjusted for permanent book-tax differences. These adjustments have no impact on net assets or the results of operations. Temporary book-tax differences will reverse in a subsequent period.

Book-tax differences are primarily due to foreign currency transactions, passive foreign investment companies (PFIC), partnerships, capital loss carryforwards and losses deferred due to wash sales and excise tax regulations.

The tax-basis components of distributable earnings and the federal tax cost as of period end were as follows for each Fund:

Cost for Federal
Income Tax
Purposes

Unrealized
Appreciation

Unrealized
Depreciation

Net Unrealized
Appreciation/
(Depreciation)

Fidelity Large Cap Value Fund

$ 1,571,171,429

$ 98,475,075

$ (92,166,547)

$ 6,308,528

Fidelity Mid Cap Value Fund

844,773,929

42,574,804

(57,370,158)

(14,795,354)

Fidelity Large Cap Growth Fund

166,610,940

4,786,892

(10,368,163)

(5,581,271)

Fidelity Mid Cap Growth Fund

319,883,846

21,520,033

(22,775,536)

(1,255,503)

Annual Report

3. Significant Accounting Policies - continued

Income Tax Information and Distributions to Shareholders - continued

The tax character of distributions paid was as follows:

January 31, 2008

Ordinary
Income

Long-term
Capital Gains

Total

Fidelity Large Cap Value Fund

$ 37,304,898

$ 67,715,379

$ 105,020,277

Fidelity Mid Cap Value Fund

10,958,373

29,219,096

40,177,469

Fidelity Large Cap Growth Fund

6,193,151

12,379,253

18,572,404

Fidelity Mid Cap Growth Fund

-

21,909,497

21,909,497

January 31, 2007

Ordinary Income

Long-term
Capital Gains

Total

Fidelity Large Cap Value Fund

$ 17,006,124

$ 7,416,999

$ 24,423,123

Fidelity Mid Cap Value Fund

7,722,049

9,225,041

16,947,090

Fidelity Large Cap Growth Fund

1,920,898

1,860,806

3,781,704

Fidelity Mid Cap Growth Fund

2,544,632

2,544,632

5,089,264

Short-Term Trading (Redemption) Fees. Shares held in the Fidelity Mid Cap Value Fund and Fidelity Mid Cap Growth Fund less than 30 days are subject to a redemption fee equal to 0.75% of the proceeds of the redeemed shares. All redemption fees, including any estimated redemption fees paid by FMR, are retained by the Funds and accounted for as an addition to paid in capital.

New Accounting Pronouncement. In September 2006, Statement of Financial Accounting Standards No. 157, Fair Value Measurements(SFAS 157), was issued and is effective for fiscal years beginning after November 15, 2007. SFAS 157 defines fair value, establishes a framework for measuring fair value and expands disclosures about fair value measurements. Management is currently evaluating the impact the adoption of SFAS 157 will have on the Funds' financial statement disclosures.

4. Operating Policies.

Repurchase Agreements. FMR has received an Exemptive Order from the Securities and Exchange Commission(the SEC) which permits certain Funds and other affiliated entities of FMR to transfer uninvested cash balances into joint trading accounts which are then invested in repurchase agreements. Certain Funds may also invest directly with institutions in repurchase agreements. Repurchase agreements are collateralized by government or non-government securities. Upon settlement date, collateral is held in segregated accounts with custodian banks and may be obtained in the event of a default of the counterparty. Each applicable Fund monitors, on a daily basis, the value of the collateral to ensure it is at least equal to the principal amount of the repurchase agreement(including accrued interest). In the event of a default by the counterparty, realization of the collateral proceeds could be delayed, during which time the value of the collateral may decline.

5. Purchases and Sales of Investments.

Purchases and sales of securities, other than short-term securities, are noted in the table below.

Purchases($)

Sales($)

Fidelity Large Cap Value Fund

3,485,833,978

3,246,788,832

Fidelity Mid Cap Value Fund

2,520,812,788

2,356,094,981

Fidelity Large Cap Growth Fund

745,952,270

764,990,241

Fidelity Mid Cap Growth Fund

995,464,036

1,090,139,044

6. Fees and Other Transactions with Affiliates.

Management Fee. FMR and its affiliates provide the Funds with investment management related services for which the Funds pay a monthly management fee. The management fee is the sum of an individual fund fee rate and a group fee rate. The individual fund fee rate is applied to each Fund's average net assets. The group fee rate is based upon the average net assets of all the mutual funds advised by FMR. The group fee rate decreases as assets under management increase and increases as assets under management decrease. In addition, the management fee for the funds is subject to a performance adjustment(up to a maximum ± .20% of each applicable Fund's average net assets over a 36 month performance period). The upward or downward adjustment to the management fee is based on each applicable Fund's relative investment performance of the asset-weighted return of all

Annual Report

Notes to Financial Statements - continued

6. Fees and Other Transactions with Affiliates - continued

Management Fee - continued

classes as compared to an appropriate benchmark index. For the period, each Fund's annual management fee rate expressed as a percentage of each Fund's average net assets, including the performance adjustment, if applicable was as follows:

Individual Rate

Group Rate

Total

Fidelity Large Cap Value Fund

.30%

.26%

.58%

Fidelity Mid Cap Value Fund

.30%

.26%

.52%

Fidelity Large Cap Growth Fund

.30%

.26%

.58%

Fidelity Mid Cap Growth Fund

.30%

.26%

.46%

Distribution and Service Plan. In accordance with Rule 12b-1 of the 1940 Act, the Fund has adopted separate Distribution and Service Plans for each class of shares. Certain classes pay Fidelity Distributors Corporation(FDC), an affiliate of FMR, separate Distribution and Service Fees, each of which is based on an annual percentage of each class' average net assets. In addition, FDC may pay financial intermediaries for selling shares of the Fund and providing shareholder support services. For the period, the Distribution and Service Fee rates and the total amounts paid to and retained by FDC were as follows:

Fidelity Large Cap Value Fund

Distribution
Fee

Service
Fee

Paid to
FDC

Retained
by FDC

Class A

0%

.25%

$ 14,716

$ 3,419

Class T

.25%

.25%

29,520

209

Class B

.75%

.25%

20,952

15,912

Class C

.75%

.25%

8,540

4,769

$ 73,728

$ 24,309

Fidelity Mid Cap Value Fund

Class A

0%

.25%

$ 9,176

$ 3,141

Class T

.25%

.25%

10,994

355

Class B

.75%

.25%

17,435

13,285

Class C

.75%

.25%

16,086

12,118

$ 53,691

$ 28,899

Fidelity Large Cap Growth Fund

Class A

0%

.25%

$ 1,502

$ 246

Class T

.25%

.25%

2,248

460

Class B

.75%

.25%

2,797

2,331

Class C

.75%

.25%

4,188

3,019

$ 10,735

$ 6,056

Fidelity Mid Cap Growth Fund

Class A

0%

.25%

$ 1,756

$ 245

Class T

.25%

.25%

1,796

463

Class B

.75%

.25%

2,959

2,452

Class C

.75%

.25%

3,852

3,025

$ 10,363

$ 6,185

Sales Load. FDC receives a front-end sales charge of up to 5.75% for selling Class A shares, and 3.50% for selling Class T shares, some of which is paid to financial intermediaries for selling shares of the Fund. FDC receives the proceeds of contingent deferred sales charges levied on Class A, Class T, Class B, and Class C redemptions. These charges depend on the holding period. The deferred sales charges range from 5% to 1% for Class B, 1% for Class C, 1.00% to .50% for certain purchases of Class A shares and .25% for certain purchases of Class T shares.

For the period, sales charge amounts retained by FDC were as follows:

Fidelity Large Cap Value Fund

Retained
by FDC

Class A

$ 18,265

Class T

4,391

Class B *

1,096

Class C *

11

$ 22,763

Annual Report

6. Fees and Other Transactions with Affiliates - continued

Sales Load - continued

For the period, sales charge amounts retained by FDC were as follows: - continued

Fidelity Mid Cap Value Fund

Retained
by FDC

Class A

$ 22,621

Class T

5,951

Class B*

2,448

Class C*

137

$ 31,157

Fidelity Large Cap Growth Fund

Class A

$ 4,515

Class T

1,123

Class B*

509

Class C*

62

$ 6,209

Fidelity Mid Cap Growth Fund

Class A

$ 6,603

Class T

1,049

Class B*

962

Class C*

165

$ 8,779

* When Class B and Class C shares are initially sold, FDC pays commissions from its own resources to financial intermediaries through which the sales are made.

Transfer Agent Fees. Fidelity Investments Institutional Operations Company, Inc.(FIIOC), an affiliate of FMR, is the transfer, dividend disbursing and shareholder servicing agent for each class of the Funds. FIIOC receives account fees and asset-based fees that vary according to the account size and type of account of the shareholders of the respective classes of each Fund. FIIOC pays for typesetting, printing and mailing of shareholder reports, except proxy statements. Prior to January 1, 2008, Fidelity Service Company, Inc. (FSC), also an affiliate of FMR, was the Fund's transfer agent. For the period, the total transfer agent fees paid by each class to FIIOC or FSC, were as follows:

Fidelity Large Cap Value Fund

Amount

% of
Average
Net Assets
*

Class A

$ 20,036

.34

Class T

20,697

.35

Class B

7,804

.37

Class C

2,694

.31

Large Cap Value

3,716,520

.23

Institutional Class

944

.23

$ 3,768,695

Fidelity Mid Cap Value Fund

Class A

$ 10,910

.30

Class T

6,730

.31

Class B

5,278

.30

Class C

5,017

.31

Mid Cap Value

2,206,272

.24

Institutional Class

1,803

.30

$ 2,236,010

Fidelity Large Cap Growth Fund

Class A

$ 1,787

.30

Class T

1,353

.31

Class B

833

.30

Class C

1,224

.29

Large Cap Growth

529,155

.31

Institutional Class

416

.20

$ 534,768

Annual Report

Notes to Financial Statements - continued

6. Fees and Other Transactions with Affiliates - continued

Transfer Agent Fees - continued

Fidelity Mid Cap Growth Fund

Amount

% of
Average
Net Assets
*

Class A

$ 2,094

.30

Class T

1,136

.32

Class B

899

.30

Class C

1,151

.30

Mid Cap Growth

1,196,809

.29

Institutional Class

225

.17

$ 1,202,314

* Annualized

Accounting and Security Lending Fees. Fidelity Service Company, Inc.(FSC), an affiliate of FMR, maintains each Fund's accounting records. The accounting fee is based on the level of average net assets for the month. Under a separate contract, FSC administers the security lending program. The security lending fee is based on the number and duration of lending transactions.

Brokerage Commissions. Certain Funds placed a portion of their portfolio transactions with brokerage firms which are affiliates of the investment adviser. The commissions paid to these affiliated firms were as follows:

Amount

Fidelity Large Cap Value Fund

$ 18,780

Fidelity Mid Cap Value Fund

12,752

Fidelity Large Cap Growth Fund

13,318

Fidelity Mid Cap Growth Fund

10,529

Interfund Lending Program. Pursuant to an Exemptive Order issued by the SEC, the Funds, along with other registered investment companies having management contracts with FMR, may participate in an interfund lending program. This program provides an alternative credit facility allowing the funds to borrow from, or lend money to, other participating affiliated funds. At period end, there were no interfund loans outstanding. Each applicable fund's activity in this program during the period for which loans were outstanding was as follows:

Borrower or
Lender

Average Daily Loan Balance

Weighted
Average Interest Rate

Interest
Expense

Fidelity Large Cap Value Fund

Borrower

$ 6,183,857

4.35%

$ 5,226

Fidelity Mid Cap Value Fund

Borrower

$ 5,637,000

5.27%

$ 4,946

7. Committed Line of Credit.

Certain Funds participate with other funds managed by FMR in a $4.2 billion credit facility(the "line of credit") to be utilized for temporary or emergency purposes to fund shareholder redemptions or for other short-term liquidity purposes. The participating funds have agreed to pay commitment fees on their pro rata portion of the line of credit, which is reflected in Miscellaneous Expense on the Statement of Operations, and is as follows:

Fidelity Large Cap Value Fund

$ 3,040

Fidelity Mid Cap Value Fund

1,696

Fidelity Large Cap Growth Fund

343

Fidelity Mid Cap Growth Fund

824

During the period, there were no borrowings on this line of credit.

8. Security Lending.

Certain Funds lend portfolio securities from time to time in order to earn additional income. On the settlement date of the loan, each applicable Fund receives collateral(in the form of U.S. Treasury obligations, letters of credit and/or cash) against the loaned securities and maintains collateral in an amount not less than 100% of the market value of the loaned securities during the period of the loan. The market value of the loaned securities is determined at the close of business of the Funds and any additional required collateral is delivered to the Funds on the next business day. If the borrower defaults on its obligation to return the securities loaned because of insolvency or other reasons, a fund could experience delays and costs in recovering the securities loaned or in gaining access to the collateral. Any cash collateral received is invested in the Fidelity Securities Lending Cash

Annual Report

8. Security Lending - continued

Central Fund. The value of loaned securities and cash collateral at period end are disclosed on each applicable Fund's Statement of Assets and Liabilities. Security lending income represents the income earned on investing cash collateral, less fees and expenses associated with the loan, plus any premium payments that may be received on the loan of certain types of securities. Security lending income is presented on each applicable Fund's Statement of Operations as a component of income from Fidelity Central Funds.

9. Expense Reductions.

FMR voluntarily agreed to reimburse funds to the extent annual operating expenses exceeded certain levels of average net assets as noted in the table below. Some expenses, for example interest expense, including commitment fees, are excluded from this reimbursement.

The following classes of each applicable Fund were in reimbursement during the period:

Expense
Limitations

Reimbursement
from adviser

Fidelity Large Cap Growth Fund

Large Cap Growth

1.00%

$ 58,552

In addition, FMR voluntarily agreed to reimburse a portion of certain fund's existing class' operating expenses. During the period, this reimbursement reduced certain fund's existing class' expenses by the following amounts:

Reimbursement
from adviser

Fidelity Large Cap Value Fund

Large Cap Value

$ 85,974

Fidelity Mid Cap Value Fund

Mid Cap Value

$ 85,078

Fidelity Large Cap Growth Fund

Large Cap Growth

$ 25,900

Fidelity Mid Cap Growth Fund

Mid Cap Growth

$ 84,552

In addition, through arrangements with each applicable Fund's custodian and each class' transfer agent, credits realized as a result of uninvested cash balances were used to reduce the Fund's expenses. All of the applicable expense reductions are noted in the table below.

Brokerage Service Arrangements

Transfer
Agent
expense
reduction

Fidelity Large Cap Value Fund

$ -

$ -

Large Cap Value

-

50,587

Institutional Class

-

50

Fidelity Mid Cap Value Fund

-

-

Class A

-

31

Class B

-

9

Mid Cap Value

-

19,810

Institutional Class

-

70

Fidelity Large Cap Growth Fund

-

-

Large Cap Growth

-

13,456

Fidelity Mid Cap Growth Fund

1,543

-

Mid Cap Growth

-

27,293

10. Other.

The Funds' organizational documents provide former and current trustees and officers with a limited indemnification against liabilities arising in connection with the performance of their duties to the Funds. In the normal course of business, the Funds may also enter into contracts that provide general indemnifications. The Funds' maximum exposure under these arrangements is unknown as this would be dependent on future claims that may be made against the Funds. The risk of material loss from such claims is considered remote.

Annual Report

Notes to Financial Statements - continued

10. Other - continued

In December 2006, the Independent Trustees, with the assistance of independent counsel, completed an investigation regarding gifts, gratuities and business entertainment provided by certain brokers to certain individuals who were employed on FMR's domestic equity trading desk during the period 2002 to 2004. The Independent Trustees and FMR agreed that, despite the absence of proof that the Fidelity mutual funds experienced diminished execution quality as a result of the improper receipt of gifts and business entertainment, the conduct at issue was serious and was worthy of redress. Accordingly, the Independent Trustees requested, and FMR agreed to make a payment of $42 million plus accrued interest, which equaled approximately $7.3 million, to certain Fidelity mutual funds. FMR also reimbursed the related legal expenses, which are recorded in the accompanying Statement of Operations as an expense reduction.

Subsequent to period end, the Trustees approved a method for allocating this payment among the funds and, in total, FMR paid each fund the following amounts:

Fidelity Large Cap Value Fund

$ 12,127

Fidelity Mid Cap Value Fund

$ 23,107

Fidelity Large Cap Growth Fund

$ 14,058

Fidelity Mid Cap Growth Fund

$ 16,394

In a related administrative order dated March 5, 2008, the U.S. Securities and Exchange Commission ("SEC") announced a settlement with FMR and FMR Co., Inc. (an affiliate of FMR) involving the SEC's regulatory rules for investment advisers and the improper receipt of gifts, gratuities and business entertainment. Without admitting or denying the SEC's findings, FMR agreed to pay an $8 million civil penalty to the United States Treasury.

11. Distributions to Shareholders.

Distributions to shareholders of each class were as follows:

Years ended January 31,

2008 A

2007

Fidelity Large Cap Value Fund

From net investment income

Class A

$ 75,483

$ -

Class T

44,031

-

Class B

824

-

Class C

4,115

-

Large Cap Value

14,152,017

10,401,399

Institutional Class

10,129

-

Total

$ 14,286,599

$ 10,401,399

From net realized gain

Class A

$ 372,920

$ -

Class T

317,190

-

Class B

97,442

-

Class C

51,607

-

Large Cap Value

89,854,195

14,021,724

Institutional Class

40,324

-

Total

$ 90,733,678

$ 14,021,724

Fidelity Mid Cap Value Fund

From net investment income

Class A

$ 18,691

$ -

Class T

5,836

-

Class B

259

-

Class C

66

-

Mid Cap Value

3,005,000

2,949,768

Institutional Class

7,163

-

Total

$ 3,037,015

$ 2,949,768

From net realized gain

Class A

$ 176,774

$ -

Class T

106,263

-

Class B

46,244

-

Class C

67,330

-

Mid Cap Value

36,700,729

13,997,322

Institutional Class

43,114

-

Total

$ 37,140,454

$ 13,997,322

Annual Report

11. Distributions to Shareholders - continued

Distributions to shareholders of each class were as follows: - continued

Years ended January 31,

2008 A

2007

Fidelity Large Cap Growth Fund

From net investment income

Large Cap Growth

$ -

$ 137,207

From net realized gain

Class A

$ 123,133

$ -

Class T

101,048

-

Class B

40,430

-

Class C

75,734

-

Large Cap Growth

18,196,705

3,644,497

Institutional Class

35,354

-

Total

$ 18,572,404

$ 3,644,497

Fidelity Mid Cap Growth Fund

From net realized gain

Class A

$ 100,774

$ -

Class T

34,957

-

Class B

25,358

-

Class C

40,118

-

Mid Cap Growth

21,697,261

5,089,264

Institutional Class

11,029

-

Total

$ 21,909,497

$ 5,089,264

A Distributions for Class A,T,B,C and Institutional are for the period February 13, 2007(commencement of sale of shares) to January 31, 2008.

12. Share Transactions.

Transactions for each class of shares were as follows:

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Fidelity Large Cap Value Fund

Class A

Shares sold

1,045,290

-

$ 16,232,942

$ -

Reinvestment of distributions

29,393

-

423,574

-

Shares redeemed

(352,749)

-

(5,365,826)

-

Net increase (decrease)

721,934

-

$ 11,290,690

$ -

Class T

Shares sold

935,752

-

$ 14,740,653

$ -

Reinvestment of distributions

24,935

-

359,076

-

Shares redeemed

(518,994)

-

(7,932,251)

-

Net increase (decrease)

441,693

-

$ 7,167,478

$ -

Class B

Shares sold

315,386

-

$ 4,951,665

$ -

Reinvestment of distributions

6,739

-

97,191

-

Shares redeemed

(184,721)

-

(2,835,906)

-

Net increase (decrease)

137,404

-

$ 2,212,950

$ -

Class C

Shares sold

138,185

-

$ 2,136,503

$ -

Reinvestment of distributions

3,506

-

50,496

-

Shares redeemed

(52,323)

-

(788,391)

-

Net increase (decrease)

89,368

-

$ 1,398,608

$ -

Annual Report

Notes to Financial Statements - continued

12. Share Transactions - continued

Transactions for each class of shares were as follows: - continued

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Fidelity Large Cap Value Fund - continued

Large Cap Value

Shares sold

52,779,386

72,550,724

$ 813,954,360

$ 1,029,577,278

Reinvestment of distributions

7,056,478

1,672,345

102,030,422

23,955,243

Shares redeemed

(40,898,915)

(25,616,795)

(619,081,485)

(368,491,524)

Net increase (decrease)

18,936,949

48,606,274

$ 296,903,297

$ 685,040,997

Institutional Class

Shares sold

92,735

-

$ 1,397,015

$ -

Reinvestment of distributions

3,477

-

50,127

-

Shares redeemed

(17,951)

-

(257,652)

-

Net increase (decrease)

78,261

-

$ 1,189,490

$ -

Fidelity Mid Cap Value Fund

Class A

Shares sold

615,995

-

$ 10,540,942

$ -

Reinvestment of distributions

11,024

-

178,558

-

Shares redeemed

(132,358)

-

(2,195,988)

-

Net increase (decrease)

494,661

-

$ 8,523,512

$ -

Class T

Shares sold

325,063

-

$ 5,643,056

$ -

Reinvestment of distributions

6,694

-

108,412

-

Shares redeemed

(84,728)

-

(1,422,111)

-

Net increase (decrease)

247,029

-

$ 4,329,357

$ -

Class B

Shares sold

204,574

-

$ 3,623,364

$ -

Reinvestment of distributions

2,791

-

45,364

-

Shares redeemed

(120,358)

-

(2,068,797)

-

Net increase (decrease)

87,007

-

$ 1,599,931

$ -

Class C

Shares sold

168,571

-

$ 2,987,730

$ -

Reinvestment of distributions

3,136

-

50,659

-

Shares redeemed

(61,027)

-

(991,710)

-

Net increase (decrease)

110,680

-

$ 2,046,679

$ -

Mid Cap Value

Shares sold

34,123,012

26,022,900

$ 603,742,926

$ 415,931,492

Reinvestment of distributions

2,338,406

1,003,867

38,409,538

16,337,499

Shares redeemed

(27,133,004)

(10,884,934)

(462,055,715)

(173,050,014)

Net increase (decrease)

9,328,414

16,141,833

$ 180,096,749

$ 259,218,977

Institutional Class

Shares sold

102,472

-

$ 1,790,525

$ -

Reinvestment of distributions

3,101

-

50,277

-

Shares redeemed

(9,162)

-

(148,581)

-

Net increase (decrease)

96,411

-

$ 1,692,221

$ -

Fidelity Large Cap Growth Fund

Class A

Shares sold

165,474

-

$ 1,900,337

$ -

Reinvestment of distributions

10,979

-

116,895

-

Shares redeemed

(44,281)

-

(486,246)

-

Net increase (decrease)

132,172

-

$ 1,530,986

$ -

Class T

Shares sold

167,227

-

$ 1,920,600

$ -

Reinvestment of distributions

9,490

-

101,048

-

Shares redeemed

(65,269)

-

(790,917)

-

Net increase (decrease)

111,448

-

$ 1,230,731

$ -

Annual Report

12. Share Transactions - continued

Transactions for each class of shares were as follows: - continued

Shares

Dollars

Years ended January 31,

2008 A

2007

2008 A

2007

Class B

Shares sold

55,847

-

$ 636,287

$ -

Reinvestment of distributions

3,795

-

40,430

-

Shares redeemed

(4,405)

-

(49,003)

-

Net increase (decrease)

55,237

-

$ 627,714

$ -

Class C

Shares sold

106,310

-

$ 1,230,146

$ -

Reinvestment of distributions

6,439

-

68,402

-

Shares redeemed

(16,516)

-

(191,737)

-

Net increase (decrease)

96,233

-

$ 1,106,811

$ -

Large Cap Growth

Shares sold

5,794,783

10,973,541

$ 66,845,422

$ 124,315,764

Reinvestment of distributions

1,665,396

318,299

17,914,862

3,701,456

Shares redeemed

(7,903,937)

(9,222,108)

(90,877,571)

(102,538,999)

Net increase (decrease)

(443,758)

2,069,732

$ (6,117,287)

$ 25,478,221

Institutional Class

Shares sold

38,550

-

$ 453,557

$ -

Reinvestment of distributions

3,006

-

32,121

-

Shares redeemed

(2,488)

-

(28,969)

-

Net increase (decrease)

39,068

-

$ 456,709

$ -

Fidelity Mid Cap Growth Fund

Class A

Shares sold

165,234

-

$ 2,326,735

$ -

Reinvestment of distributions

7,218

-

97,305

-

Shares redeemed

(13,557)

-

(186,849)

-

Net increase (decrease)

158,895

-

$ 2,237,191

$ -

Class T

Shares sold

54,813

-

$ 796,065

$ -

Reinvestment of distributions

2,601

-

34,957

-

Shares redeemed

(8,726)

-

(121,838)

-

Net increase (decrease)

48,688

-

$ 709,184

$ -

Class B

Shares sold

35,917

-

$ 516,488

$ -

Reinvestment of distributions

1,882

-

25,182

-

Shares redeemed

(3,583)

-

(47,283)

-

Net increase (decrease)

34,216

-

$ 494,387

$ -

Class C

Shares sold

59,795

-

$ 849,458

$ -

Reinvestment of distributions

2,811

-

37,643

-

Shares redeemed

(4,934)

-

(66,440)

-

Net increase (decrease)

57,672

-

$ 820,661

$ -

Mid Cap Growth

Shares sold

7,994,956

24,363,804

$ 114,388,509

$ 336,430,868

Reinvestment of distributions

1,580,341

357,726

21,334,599

4,986,704

Shares redeemed

(15,751,578)

(18,209,467)

(217,016,915)

(243,991,431)

Net increase (decrease)

(6,176,281)

6,512,063

$ (81,293,807)

$ 97,426,141

Institutional Class

Shares sold

14,080

-

$ 199,943

$ -

Reinvestment of distributions

816

-

11,029

-

Net increase (decrease)

14,896

-

$ 210,972

$ -

A Share transactions for class A,T,B,C and Institutional are for the period February 13, 2007 (commencement of sale of shares) to January 31, 2008.

Annual Report

Report of Independent Registered Public Accounting Firm

To the Trustees of Fidelity Devonshire Trust and the Shareholders of Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund:

In our opinion, the accompanying statements of assets and liabilities, including the schedules of investments, and the related statements of operations and of changes in net assets and the financial highlights present fairly, in all material respects, the financial position of Fidelity Large Cap Value Fund, Fidelity Mid Cap Value Fund, Fidelity Large Cap Growth Fund and Fidelity Mid Cap Growth Fund (funds of Fidelity Devonshire Trust) at January 31, 2008, the results of each of their operations for the year then ended, the changes in each of their net assets for each of the two years in the period then ended and the financial highlights for each of the periods indicated, in conformity with accounting principles generally accepted in the United States of America. These financial statements and financial highlights (hereafter referred to as "financial statements") are the responsibility of the Fidelity Devonshire Trust's management. Our responsibility is to express an opinion on these financial statements based on our audits. We conducted our audits of these financial statements in accordance with the standards of the Public Company Accounting Oversight Board (United States). Those standards require that we plan and perform the audit to obtain reasonable assurance about whether the financial statements are free of material misstatement. An audit includes examining, on a test basis, evidence supporting the amounts and disclosures in the financial statements, assessing the accounting principles used and significant estimates made by management, and evaluating the overall financial statement presentation. We believe that our audits, which included confirmation of securities at January 31, 2008 by correspondence with the custodian and brokers, provide a reasonable basis for our opinion.

/s/ PricewaterhouseCoopers LLP

PricewaterhouseCoopers LLP

Boston, Massachusetts
March 24, 2008

Annual Report

Trustees and Officers

The Trustees, Members of the Advisory Board, and executive officers of the trust and funds, as applicable, are listed below. The Board of Trustees governs each fund and is responsible for protecting the interests of shareholders. The Trustees are experienced executives who meet periodically throughout the year to oversee each fund's activities, review contractual arrangements with companies that provide services to each fund, and review each fund's performance. Except for James C. Curvey, each of the Trustees oversees 374 funds advised by FMR or an affiliate. Mr. Curvey oversees 369 funds advised by FMR or an affiliate.

The Trustees hold office without limit in time except that (a) any Trustee may resign; (b) any Trustee may be removed by written instrument, signed by at least two-thirds of the number of Trustees prior to such removal; (c) any Trustee who requests to be retired or who has become incapacitated by illness or injury may be retired by written instrument signed by a majority of the other Trustees; and (d) any Trustee may be removed at any special meeting of shareholders by a two-thirds vote of the outstanding voting securities of the trust. Each Trustee who is not an interested person (as defined in the 1940 Act) (Independent Trustee), shall retire not later than the last day of the calendar year in which his or her 72nd birthday occurs. The Independent Trustees may waive this mandatory retirement age policy with respect to individual Trustees. The executive officers and Advisory Board Members hold office without limit in time, except that any officer and Advisory Board Member may resign or may be removed by a vote of a majority of the Trustees at any regular meeting or any special meeting of the Trustees. Except as indicated, each individual has held the office shown or other offices in the same company for the past five years.

The funds' Statement of Additional Information (SAI) includes more information about the Trustees. To request a free copy, call Fidelity at 1-800-544-8544.

Interested Trustees*:

Correspondence intended for each Trustee who is an interested person may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Edward C. Johnson 3d (77)

Year of Election or Appointment: 1985

Mr. Johnson is Chairman of the Board of Trustees. Mr. Johnson serves as President (2006-present), Chief Executive Officer, Chairman, and a Director of FMR LLC; Chairman and a Director of FMR; Chairman and a Director of Fidelity Research & Analysis Company (FRAC); Chairman and a Director of Fidelity Investments Money Management, Inc.; and Chairman (2001-present) and a Director of FMR Co., Inc. In addition, Mr. Johnson serves as Chairman and Director of FIL Limited. Mr. Edward C. Johnson 3d and Mr. Arthur E. Johnson are not related.

James C. Curvey (72)

Year of Election or Appointment: 2007

Mr. Curvey also serves as Trustee (2007-present) or Member of the Advisory Board (2007-present) of other investment companies advised by FMR. Mr. Curvey is a Director of FMR and FMR Co., Inc. (2007-present). Mr. Curvey is also Vice Chairman (2006-present) and Director of FMR LLC. Mr. Curvey joined Fidelity in 1982 and served in numerous senior management positions, including President and Chief Operating Officer of FMR LLC (1997-2000) and President of Fidelity Strategic Investments (2000-2002). In addition, he serves as a member of the Board of Directors of Geerlings & Wade, Inc. (wine distribution).

* Trustees have been determined to be "Interested Trustees" by virtue of, among other things, their affiliation with the trust or various entities under common control with FMR. FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Independent Trustees:

Correspondence intended for each Independent Trustee (that is, the Trustees other than the Interested Trustees) may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235.

Name, Age; Principal Occupation

Dennis J. Dirks (59)

Year of Election or Appointment: 2005

Prior to his retirement in May 2003, Mr. Dirks was Chief Operating Officer and a member of the Board of The Depository Trust & Clearing Corporation (DTCC) (1999-2003). He also served as President, Chief Operating Officer, and Board member of The Depository Trust Company (DTC) (1999-2003) and President and Board member of the National Securities Clearing Corporation (NSCC) (1999-2003). In addition, Mr. Dirks served as Chief Executive Officer and Board member of the Government Securities Clearing Corporation (2001-2003) and Chief Executive Officer and Board member of the Mortgage-Backed Securities Clearing Corporation (2001-2003). Mr. Dirks also serves as a Trustee and a member of the Finance Committee of Manhattan College (2005-present) and a Trustee and a member of the Finance Committee of AHRC of Nassau County (2006-present).

Albert R. Gamper, Jr. (65)

Year of Election or Appointment: 2006

Prior to his retirement in December 2004, Mr. Gamper served as Chairman of the Board of CIT Group Inc. (commercial finance). During his tenure with CIT Group Inc. Mr. Gamper served in numerous senior management positions, including Chairman (1987-1989; 1999-2001; 2002-2004), Chief Executive Officer (1987-2004), and President (1989-2002). He currently serves as a member of the Board of Directors of Public Service Enterprise Group (utilities, 2001-present), Chairman of the Board of Governors, Rutgers University (2004-present), and Chairman of the Board of Saint Barnabas Health Care System.

George H. Heilmeier (71)

Year of Election or Appointment: 2004

Dr. Heilmeier is Chairman Emeritus of Telcordia Technologies (communication software and systems), where prior to his retirement, he served as company Chairman and Chief Executive Officer. He currently serves on the Boards of Directors of The Mitre Corporation (systems engineering and information technology support for the government), and HRL Laboratories (private research and development, 2004-present). He is Chairman of the General Motors Science & Technology Advisory Board and a Life Fellow of the Institute of Electrical and Electronics Engineers (IEEE). Dr. Heilmeier is a member of the Defense Science Board and the National Security Agency Advisory Board. He is also a member of the National Academy of Engineering, the American Academy of Arts and Sciences, and the Board of Overseers of the School of Engineering and Applied Science of the University of Pennsylvania. Previously, Dr. Heilmeier served as a Director of TRW Inc. (automotive, space, defense, and information technology, 1992-2002), Compaq (1994-2002), Automatic Data Processing, Inc. (ADP) (technology-based business outsourcing, 1995-2002), INET Technologies Inc. (telecommunications network surveillance, 2001-2004), and Teletech Holdings (customer management services). He is the recipient of the 2005 Kyoto Prize in Advanced Technology for his invention of the liquid crystal display, and a member of the Consumer Electronics Hall of Fame.

James H. Keyes (67)

Year of Election or Appointment: 2007

Prior to his retirement in 2003, Mr. Keyes was Chairman, President, and Chief Executive Officer of Johnson Controls, Inc. (automotive supplier, 1993-2003). He currently serves as a member of the boards of LSI Logic Corporation (semiconductor technologies), Navistar International Corporation (manufacture and sale of trucks, buses, and diesel engines, 2002-present), and Pitney Bowes, Inc. (integrated mail, messaging, and document management solutions).

Marie L. Knowles (61)

Year of Election or Appointment: 2001

Prior to Ms. Knowles' retirement in June 2000, she served as Executive Vice President and Chief Financial Officer of Atlantic Richfield Company (ARCO) (diversified energy, 1996-2000). From 1993 to 1996, she was a Senior Vice President of ARCO and President of ARCO Transportation Company. She served as a Director of ARCO from 1996 to 1998. She currently serves as a Director of Phelps Dodge Corporation (copper mining and manufacturing) and McKesson Corporation (healthcare service, 2002-present). Ms. Knowles is a Trustee of the Brookings Institution and the Catalina Island Conservancy and also serves as a member of the Advisory Board for the School of Engineering of the University of Southern California.

Ned C. Lautenbach (63)

Year of Election or Appointment: 2000

Mr. Lautenbach is Chairman of the Independent Trustees (2006-present). Mr. Lautenbach has been a partner of Clayton, Dubilier & Rice, Inc. (private equity investment firm) since September 1998. Previously, Mr. Lautenbach was with the International Business Machines Corporation (IBM) from 1968 until his retirement in 1998. Mr. Lautenbach serves as a Director of Sony Corporation (2006-present) and Eaton Corporation (diversified industrial) as well as the Philharmonic Center for the Arts in Naples, Florida. He also is a member of the Board of Trustees of Fairfield University (2005-present), as well as a member of the Council on Foreign Relations.

Cornelia M. Small (63)

Year of Election or Appointment: 2005

Ms. Small is a member (2000-present) and Chairperson (2002-present) of the Investment Committee, and a member (2002-present) of the Board of Trustees of Smith College. Previously, she served as Chief Investment Officer (1999-2000), Director of Global Equity Investments (1996-1999), and a member of the Board of Directors of Scudder, Stevens & Clark (1990-1997) and Scudder Kemper Investments (1997-1999). In addition, Ms. Small served as Co-Chair (2000-2003) of the Annual Fund for the Fletcher School of Law and Diplomacy.

William S. Stavropoulos (68)

Year of Election or Appointment: 2002

Mr. Stavropoulos is Chairman Emeritus of the Board of Directors of The Dow Chemical Company. Since joining The Dow Chemical Company in 1967, Mr. Stavropoulos served in numerous senior management positions, including President (1993-2000; 2002-2003), CEO (1995-2000; 2002-2004), and Chairman of the Executive Committee (2000-2004). Currently, he is a Director of NCR Corporation (data warehousing and technology solutions), Chemical Financial Corporation, Maersk Inc. (industrial conglomerate, 2002-present), Tyco International, Inc. (multinational manufacturing and services, 2007-present), and a member of the Advisory Board for Metalmark Capital (private equity investment firm, 2005-present). He is a special advisor to Clayton, Dubilier & Rice, Inc., a private equity investment firm. He also serves as a member of the Board of Trustees of the American Enterprise Institute for Public Policy Research. In addition, Mr. Stavropoulos is a member of The Business Council, J.P. Morgan International Council and the University of Notre Dame Advisory Council for the College of Science.

Kenneth L. Wolfe (68)

Year of Election or Appointment: 2005

Mr. Wolfe is Chairman and a Director of Hershey Foods Corporation (2007-present), where prior to his retirement in 2001, he was Chairman and Chief Executive Officer. Mr. Wolfe currently serves as a member of the board of Revlon Inc. (2004-present). Previously, Mr. Wolfe served as a member of the boards of Adelphia Communications Corporation (2003-2006) and Bausch & Lomb, Inc. (1993-2007).

Advisory Board Members and Executive Officers**:

Correspondence intended for Mr. Mauriello, Mr. Thomas, Mr. Wiley, Mr. Lacy, and Mr. Arthur Johnson may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235. Correspondence intended for each executive officer and Mr. Lynch may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Arthur E. Johnson (61)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Johnson serves as Senior Vice President of Corporate Strategic Development of Lockheed Martin Corporation (defense contractor). In addition, Mr. Johnson serves as a member of the Board of Directors of AGL Resources, Inc. (holding company, 2002-present), and IKON Office Solutions, Inc. (document management systems and services). Mr. Arthur E. Johnson and Mr. Edward C. Johnson 3d are not related.

Alan J. Lacy (54)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lacy serves as Senior Adviser (2007-present) of Oak Hill Capital Partners, L.P. (a private equity firm). Mr. Lacy also served as Vice Chairman and Chief Executive Officer of Sears Holdings Corporation and Sears, Roebuck and Co. (retail, 2005-2006; 2000-2005). In addition, Mr. Lacy serves as a member of the Board of Directors of The Western Union Company (global money transfer, 2006-present) and Bristol-Myers Squibb (global pharmaceuticals, 2007-present). Mr. Lacy is a Trustee of the National Parks Conservation Association and The Field Museum of Natural History.

Peter S. Lynch (64)

Year of Election or Appointment: 2003

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lynch is Vice Chairman and a Director of FMR, and Vice Chairman (2001-present) and a Director of FMR Co., Inc. Previously, Mr. Lynch served as a Trustee of the Fidelity funds (1990-2003). In addition, he serves as a Trustee of Boston College and as the Chairman of the Inner-City Scholarship Fund.

Joseph Mauriello (63)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Prior to his retirement in January 2006, Mr. Mauriello served in numerous senior management positions including Deputy Chairman and Chief Operating Officer (2004-2005), and Vice Chairman of Financial Services (2002-2004) of KPMG LLP US (professional services firm, 1965-2005). Mr. Mauriello currently serves as a member of the Board of Directors of XL Capital Ltd. (global insurance and re-insurance company, 2006-present) and of Arcadia Resources Inc. (health care services and products, 2007-present). He also served as a Director of the Hamilton Funds of the Bank of New York (2006-2007).

David M. Thomas (58)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Previously, Mr. Thomas served as Executive Chairman (2005-2006) and Chairman and Chief Executive Officer (2000-2005) of IMS Health, Inc. (pharmaceutical and healthcare information solutions). In addition, Mr. Thomas serves as a member of the Board of Directors of Fortune Brands, Inc. (consumer products holding company), and Interpublic Group of Companies, Inc. (marketing communication, 2004-present).

Michael E. Wiley (57)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Wiley also serves as a member of the Board of Trustees of the University of Tulsa (2000-2006; 2007-present). He serves as a Director of Tesoro Corporation (independent oil refiner and marketer, 2005-present), and a Director of Bill Barrett Corporation (exploration and production company, 2005-present). In addition, he also serves as a Director of Post Oak Bank (privately-held bank, 2004-present). Previously, Mr. Wiley served as a Sr. Energy Advisor of Katzenbach Partners, LLC (consulting firm, 2006-2007), as an Advisory Director of Riverstone Holdings (private investment firm), Chairman, President, and CEO of Baker Hughes, Inc. (oilfield services company, 2000-2004), and as Director of Spinnaker Exploration Company (exploration and production company, 2001-2005).

Kimberley H. Monasterio (44)

Year of Election or Appointment: 2007

President and Treasurer of the funds. Ms. Monasterio also serves as President and Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2004-present). Previously, Ms. Monasterio served as Deputy Treasurer of the Fidelity funds (2004-2006). Before joining Fidelity Investments, Ms. Monasterio served as Treasurer (2000-2004) and Chief Financial Officer (2002-2004) of the Franklin Templeton Funds and Senior Vice President of Franklin Templeton Services, LLC (2000-2004).

Walter C. Donovan (45)

Year of Election or Appointment: 2007

Vice President of the funds. Mr. Donovan also serves as Vice President of Fidelity's Equity Funds (2007-present). Mr. Donovan also serves as Executive Vice President of FMR (2005-present) and FMR Co., Inc. (2005-present). Previously, Mr. Donovan served as Vice President of Fidelity's High Income Funds (2005-2007), Fixed-Income Funds (2005-2006), certain Asset Allocation Funds (2005-2006), certain Balanced Funds (2005-2006), and as Vice President and Director of Fidelity's International Equity Trading group (1998-2005).

Bruce T. Herring (42)

Year of Election or Appointment: 2006

Vice President of the funds. Mr. Herring also serves as Vice President of certain Equity Funds (2006-present). Mr. Herring is Senior Vice President of FMR (2006-present) and Vice President of FMR Co., Inc. (2001-present). Previously, Mr. Herring served as a portfolio manager for Fidelity U.S. Equity Funds (2001-2005).

Eric D. Roiter (59)

Year of Election or Appointment: 2001

Secretary of the funds. He also serves as Secretary of other Fidelity funds; Vice President, General Counsel, and Secretary of FMR Co., Inc. (2001-present) and FMR; Assistant Secretary of Fidelity Management & Research (U.K.) Inc. (2001-present), Fidelity Research & Analysis Company (2001-present), and Fidelity Investments Money Management, Inc. (2001-present). Mr. Roiter is an Adjunct Member, Faculty of Law, at Boston College Law School (2003-present). Previously, Mr. Roiter served as Vice President and Secretary of Fidelity Distributors Corporation (FDC) (1998-2005).

John B. McGinty, Jr. (45)

Year of Election or Appointment: 2008

Assistant Secretary of the funds. Mr. McGinty also serves as Assistant Secretary of other Fidelity funds (2008-present) and is an employee of FMR LLC (2004-present). Mr. McGinty also serves as Senior Vice President, Secretary, and Chief Legal Officer of FDC (2007-present). Before joining Fidelity Investments, Mr. McGinty practiced law at Ropes & Gray, LLP.

R. Stephen Ganis (41)

Year of Election or Appointment: 2006

Anti-Money Laundering (AML) officer of the funds. Mr. Ganis also serves as AML officer of other Fidelity funds (2006-present) and FMR LLC (2003-present). Before joining Fidelity Investments, Mr. Ganis practiced law at Goodwin Procter, LLP (2000-2002).

Joseph B. Hollis (59)

Year of Election or Appointment: 2006

Chief Financial Officer of the funds. Mr. Hollis also serves as Chief Financial Officer of other Fidelity funds. Mr. Hollis is President of Fidelity Pricing and Cash Management Services (FPCMS) (2005-present). Mr. Hollis also serves as President and Director of Fidelity Service Company, Inc. (2006-present). Previously, Mr. Hollis served as Senior Vice President of Cash Management Services (1999-2002) and Investment Management Operations (2002-2005).

Kenneth A. Rathgeber (60)

Year of Election or Appointment: 2004

Chief Compliance Officer of the funds. Mr. Rathgeber also serves as Chief Compliance Officer of other Fidelity funds (2004-present) and Executive Vice President of Risk Oversight for Fidelity Investments (2002-present). He is Chief Compliance Officer of FMR (2005-present), FMR Co., Inc. (2005-present), Fidelity Management & Research (U.K.) Inc. (2005-present), Fidelity Research & Analysis Company (2005-present), Fidelity Investments Money Management, Inc. (2005-present), and Strategic Advisers, Inc. (2005-present). Previously, Mr. Rathgeber served as Executive Vice President and Chief Operating Officer for Fidelity Investments Institutional Services Company, Inc. (1998-2002).

Bryan A. Mehrmann (46)

Year of Election or Appointment: 2005

Deputy Treasurer of the funds. Mr. Mehrmann also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR. Previously, Mr. Mehrmann served as Vice President of Fidelity Investments Institutional Services Group (FIIS)/Fidelity Investments Institutional Operations Corporation, Inc. (FIIOC) Client Services (1998-2004).

Kenneth B. Robins (38)

Year of Election or Appointment: 2005

Deputy Treasurer of the funds. Mr. Robins also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2004-present). Before joining Fidelity Investments, Mr. Robins worked at KPMG LLP, where he was a partner in KPMG's department of professional practice (2002-2004) and a Senior Manager (1999-2000). In addition, Mr. Robins served as Assistant Chief Accountant, United States Securities and Exchange Commission (2000-2002).

Robert G. Byrnes (41)

Year of Election or Appointment: 2005

Assistant Treasurer of the funds. Mr. Byrnes also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Byrnes served as Vice President of FPCMS (2003-2005). Before joining Fidelity Investments, Mr. Byrnes worked at Deutsche Asset Management where he served as Vice President of the Investment Operations Group (2000-2003).

Peter L. Lydecker (54)

Year of Election or Appointment: 2004

Assistant Treasurer of the funds. Mr. Lydecker also serves as Assistant Treasurer of other Fidelity funds (2004) and is an employee of FMR.

Paul M. Murphy (60)

Year of Election or Appointment: 2007

Assistant Treasurer of the funds. Mr. Murphy also serves as Assistant Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2007-present). Previously, Mr. Murphy served as Chief Financial Officer of the Fidelity Funds (2005-2006), Vice President and Associate General Counsel of FMR (2007), and Senior Vice President of Fidelity Pricing and Cash Management Services Group (FPCMS) (1994-2007).

Gary W. Ryan (49)

Year of Election or Appointment: 2005

Assistant Treasurer of the funds. Mr. Ryan also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Ryan served as Vice President of Fund Reporting in FPCMS (1999-2005).

** FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Annual Report

Distributions

The funds hereby designate as capital gain dividend the amounts noted below for the taxable year ended January 31, 2008, or, if subsequently determined to be different, the net capital gain of such year.

Fidelity Large Cap Value Fund

$53,592,103

Fidelity Mid Cap Value Fund

$30,917,128

Fidelity Large Cap Growth Fund

$9,534,777

Fidelity Mid Cap Growth Fund

$25,157,607

A percentage of the dividends distributed during the fiscal year for the following funds qualifies for the dividends-received deduction for corporate shareholders:

March,
2007

December,
2007

Fidelity Large Cap Value Fund

Institutional Class

45%

100%

Fidelity Mid Cap Value Fund

Institutional Class

26%

100%

Fidelity Large Cap Growth Fund

Institutional Class

91%

17%

A percentage of the dividends distributed during the fiscal year for the following funds may be taken into account as a dividend for purposes of the maximum rate under section 1(h)(11) of the Internal Revenue Code.

March,
2007

December,
2007

Fidelity Large Cap Value Fund

Institutional Class

45%

100%

Fidelity Mid Cap Value Fund

Institutional Class

26%

100%

Fidelity Large Cap Growth Fund

Institutional Class

92%

19%

The fund will notify shareholders in January 2009 of amounts for use in preparing 2008 income tax returns.

Annual Report

Annual Report

Annual Report

Annual Report

Investment Adviser

Fidelity Management & Research Company
Boston, MA

Investment Sub-Adviser

FMR Co., Inc.

Fidelity Management & Research
(U.K.) Inc.

Fidelity Research & Analysis Company

Fidelity Investments Japan Limited

Fidelity International Investment Advisors

Fidelity International Investment Advisors
(U.K.) Limited

General Distributor

Fidelity Distributors Corporation
Boston, MA

Transfer and Service Agents

Fidelity Investments Institutional Operations
Company, Inc.
Boston, MA

Fidelity Service Company, Inc.
Boston, MA

Custodian

Citibank, N.A.
New York, NY

The Fidelity Telephone Connection

Mutual Fund 24-Hour Service

Exchanges/Redemptions
and Account Assistance 1-800-544-6666

Product Information 1-800-544-6666

Retirement Accounts 1-800-544-4774
(8 a.m. - 9 p.m.)

TDD Service 1-800-544-0118
for the deaf and hearing impaired
(9 a.m. - 9 p.m. Eastern time)

Fidelity Automated Service
Telephone (FAST®) (automated graphic)    1-800-544-5555

(automated graphic)    Automated line for quickest service

(Fidelity Investment logo)(registered trademark)
Corporate Headquarters
82 Devonshire St., Boston, MA 02109
www.fidelity.com

ALMCI-UANN-0308
1.863130.100

Fidelity®

Utilities

Fund

Annual Report

January 31, 2008

(2_fidelity_logos) (Registered_Trademark)

Contents

Chairman's Message

<Click Here>

Ned Johnson's message to shareholders.

Performance

<Click Here>

How the fund has done over time.

Management's Discussion

<Click Here>

The manager's review of fund performance, strategy and outlook.

Shareholder Expense Example

<Click Here>

An example of shareholder expenses.

Investment Changes

<Click Here>

A summary of major shifts in the fund's investments over the past six months.

Investments

<Click Here>

A complete list of the fund's investments with their market values.

Financial Statements

<Click Here>

Statements of assets and liabilities, operations, and changes in net assets,
as well as financial highlights.

Notes

<Click Here>

Notes to the financial statements.

Report of Independent Registered Public Accounting Firm

<Click Here>

Trustees and Officers

<Click Here>

Distributions

<Click Here>

To view a fund's proxy voting guidelines and proxy voting record for the 12-month period ended June 30, visit http://www.fidelity.com (search for "proxy voting guidelines") or visit the Securities and Exchange Commission's (SEC) web site at http://www.sec.gov. You may also call 1-800-544-8544 to request a free copy of the proxy voting guidelines.

Standard & Poor's, S&P and S&P 500 are registered service marks of The McGraw-Hill Companies, Inc. and have been licensed for use by Fidelity Distributors Corporation.

Other third party marks appearing herein are the property of their respective owners.

All other marks appearing herein are registered or unregistered trademarks or service marks of FMR LLC or an affiliated company.

Annual Report

This report and the financial statements contained herein are submitted for the general information of the shareholders of the fund. This report is not authorized for distribution to prospective investors in the fund unless preceded or accompanied by an effective prospectus.

A fund files its complete schedule of portfolio holdings with the SEC for the first and third quarters of each fiscal year on Form N-Q. Forms N-Q are available on the SEC's web site at http://www.sec.gov. A fund's Forms N-Q may be reviewed and copied at the SEC's Public Reference Room in Washington, DC. Information regarding the operation of the SEC's Public Reference Room may be obtained by calling 1-800-SEC-0330. For a complete list of a fund's portfolio holdings, view the most recent holdings listing, semiannual report, or annual report on Fidelity's web site at http://www.fidelity.com or http://www.advisor.fidelity.com, as applicable.

NOT FDIC INSURED · MAY LOSE VALUE · NO BANK GUARANTEE

Neither the fund nor Fidelity Distributors Corporation is a bank.

Annual Report

Chairman's Message

(photo_of_Edward_C_Johnson_3d)

Dear Shareholder:

Stocks got off to a poor start in 2008, while investment-grade bonds and money markets showed positive returns, once again underscoring the importance of a diversified portfolio. Financial markets are always unpredictable, but there are a number of time-tested principles that can put the historical odds in your favor.

One of the basic tenets is to invest for the long term. Over time, riding out the markets' inevitable ups and downs has proven much more effective than selling into panic or chasing the hottest trend. Even missing only a few of the markets' best days can significantly diminish investor returns. Patience also affords the benefits of compounding - of earning interest on additional income or reinvested dividends and capital gains. There are tax advantages and cost benefits to consider as well. The more you sell, the more taxes you pay, and the more you trade, the higher the costs. While staying the course doesn't eliminate risk, it can considerably lessen the effect of short-term declines.

You can further manage your investing risk through diversification. And today, more than ever, geographic diversification should be taken into account. Studies indicate that asset allocation is the single most important determinant of a portfolio's long-term success. The right mix of stocks, bonds and cash - aligned to your particular risk tolerance and investment objective - is very important. Age-appropriate rebalancing is also an essential aspect of asset allocation. For younger investors, an emphasis on equities - which historically have been the best-performing asset class over time - is encouraged. As investors near their specific goal, such as retirement or sending a child to college, consideration may be given to replacing volatile assets (e.g. common stocks) with more-stable fixed investments (bonds or savings plans).

A third investment principle - investing regularly - can help lower the average cost of your purchases. Investing a certain amount of money each month or quarter helps ensure you won't pay for all your shares at market highs. This strategy - known as dollar cost averaging - also reduces unconstructive "emotion" from investing, helping shareholders avoid selling weak performers just prior to an upswing, or chasing a hot performer just before a correction.

We invite you to contact us via the Internet, through our Investor Centers or over the phone. It is our privilege to provide you the information you need to make the investments that are right for you.

Sincerely,

/s/Edward C. Johnson 3d

Edward C. Johnson 3d

Annual Report

Performance: The Bottom Line

Average annual total return reflects the change in the value of an investment, assuming reinvestment of the fund's dividend income and capital gains (the profits earned upon the sale of securities that have grown in value, if any) and assuming a constant rate of performance each year. The $10,000 table and the fund's returns do not reflect the deduction of taxes that a shareholder would pay on fund distributions or the redemption of fund shares. During periods of reimbursement by Fidelity, a fund's total return will be greater than it would be had the reimbursement not occurred. How a fund did yesterday is no guarantee of how it will do tomorrow.

Average Annual Total Returns

Periods ended January 31, 2008

Past 1
year

Past 5
years

Past 10
years

Fidelity® Utilities Fund

0.24%

17.20%

5.58%

$10,000 Over 10 Years

Let's say hypothetically that $10,000 was invested in Fidelity® Utilities Fund on January 31, 1998. The chart shows how the value of your investment would have changed, and also shows how the Standard & Poor's 500SM Index (S&P 500®) performed over the same period.



Annual Report

Management's Discussion of Fund Performance

Comments from Douglas Simmons, Portfolio Manager of Fidelity® Utilities Fund

Many measures of the U.S. stock market retreated into negative territory for the 12 months ending January 31, 2008, felled by the subprime mortgage loan crisis, a housing market slowdown, rising inflation and a potential recession. Making matters worse was the subprime-related spillover into the credit markets, which caused liquidity to tighten significantly and led to downward revisions in economic growth expectations for 2008. The Federal Reserve Board sought to ease the credit crunch, cutting a key short-term interest rate five times during the past year. Despite the Fed's efforts, the Standard & Poor's 500SM Index fell 6.00% in January alone - its worst first month of the year since 1990 - and dropped 2.31% for the 12 months overall. The technology-heavy NASDAQ Composite® Index had its weakest January on record since its 1971 inception, and fell 2.35% during the past year. The bellwether Dow Jones Industrial AverageSM managed a modest increase of 2.56% for the period overall, but the small-cap-oriented Russell 2000® Index declined 9.79%.

For the 12 months that ended January 31, 2008, the fund returned 0.24%, outperforming the -1.03% return of the Russell 3000® Utilities Index. Overweighting the independent power producer/energy trading group had a positive influence on the fund's relative performance, as did strong stock selection among multi-utilities. Underweighting electric utilities detracted. Top contributions came from underweighting wireless telecommunication services provider Sprint Nextel, and from the positive performance of the fund's investments in Maryland-based independent power producer/energy trader Constellation Energy Group, multi-utility Public Service Enterprise Group, located in New Jersey, and Pennsylvania-domiciled electric utility PPL - and an out-of-benchmark position in independent power producer Reliant Energy, based in Houston and no longer held by the fund at period end. Detractors included underweighted positions in two stocks that performed well - Florida-based electric utility FPL Group and wireless telecom services provider Alltel, the latter of which was acquired. An investment in wireless telecom firm NII Holdings also dampened relative returns, as did an underweighting in electric utility Duke Energy - located in North Carolina and no longer held - during the second half of the period.

The views expressed above reflect those of the portfolio manager(s) only through the end of the period as stated on the cover of this report and do not necessarily represent the views of Fidelity or any other person in the Fidelity organization. Any such views are subject to change at any time based upon market or other conditions and Fidelity disclaims any responsibility to update such views. These views may not be relied on as investment advice and, because investment decisions for a Fidelity fund are based on numerous factors, may not be relied on as an indication of trading intent on behalf of any Fidelity fund.

Annual Report

Shareholder Expense Example

As a shareholder of the Fund, you incur two types of costs: (1) transaction costs, and (2) ongoing costs, including management fees and other Fund expenses. This Example is intended to help you understand your ongoing costs (in dollars) of investing in the Fund and to compare these costs with the ongoing costs of investing in other mutual funds.

The Example is based on an investment of $1,000 invested at the beginning of the period and held for the entire period (August 1, 2007 to January 31, 2008).

Actual Expenses

The first line of the accompanying table provides information about actual account values and actual expenses. You may use the information in this line, together with the amount you invested, to estimate the expenses that you paid over the period. Simply divide your account value by $1,000.00 (for example, an $8,600 account value divided by $1,000.00 = 8.6), then multiply the result by the number in the first line under the heading entitled "Expenses Paid During Period" to estimate the expenses you paid on your account during this period. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, the Fund, as a share-holder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Hypothetical Example for Comparison Purposes

The second line of the accompanying table provides information about hypothetical account values and hypothetical expenses based on the Fund's actual expense ratio and an assumed rate of return of 5% per year before expenses, which is not the Fund's actual return. The hypothetical account values and expenses may not be used to estimate the actual ending account balance or expenses you paid for the period. You may use this information to compare the ongoing costs of investing in the Fund and other funds. To do so, compare this 5% hypothetical example with the 5% hypothetical examples that appear in the shareholder reports of the other funds. A small balance maintenance fee of $12.00 that is charged once a year may apply for certain accounts with a value of less than $2,000. This fee is not included in the table below. If it was, the estimate of expenses you paid during the period would be higher, and your ending account value lower, by this amount. In addition, the Fund, as a shareholder in the underlying Fidelity Central Funds, will indirectly bear its pro rata share of the fees and expenses incurred by the underlying Fidelity Central Funds. These fees and expenses are not included in the Fund's annualized expense ratio used to calculate the expense estimate in the table below.

Annual Report

Investments - continued

Please note that the expenses shown in the table are meant to highlight your ongoing costs only and do not reflect any transaction costs. Therefore, the second line of the table is useful in comparing ongoing costs only, and will not help you determine the relative total costs of owning different funds.

Beginning
Account Value
August 1, 2007

Ending
Account Value
January 31, 2008

Expenses Paid
During Period
*
August 1, 2007 to January 31, 2008

Actual

$ 1,000.00

$ 943.50

$ 4.11

Hypothetical (5% return per year before expenses)

$ 1,000.00

$ 1,020.97

$ 4.28

* Expenses are equal to the Fund's annualized expense ratio of .84%; multiplied by the average account value over the period, multiplied by 184/365 (to reflect the one-half year period).

Annual Report

Investment Changes

Top Ten Stocks as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

AT&T, Inc.

23.5

24.3

Verizon Communications, Inc.

10.7

11.4

Exelon Corp.

6.2

4.5

AES Corp.

5.4

4.3

Comcast Corp. Class A

4.9

6.4

PPL Corp.

4.8

2.8

Constellation Energy Group, Inc.

4.6

3.2

Public Service Enterprise Group, Inc.

3.7

4.3

American Electric Power Co., Inc.

2.8

2.2

Entergy Corp.

2.8

2.8

69.4

Top Five Industries as of January 31, 2008

% of fund's
net assets

% of fund's net assets
6 months ago

Diversified Telecommunication Services

37.7

40.8

Electric Utilities

27.2

17.4

Independent Power Producers & Energy Traders

12.3

13.8

Multi-Utilities

9.0

9.8

Media

4.9

6.4

Asset Allocation (% of fund's net assets)

As of January 31, 2008 *

As of July 31, 2007 **

Stocks 98.1%

Stocks 99.9%

Short-Term
Investments and
Net Other Assets 1.9%

Short-Term
Investments and
Net Other Assets 0.1%

* Foreign investments

0.0%

** Foreign investments

0.3%

Annual Report

Investments January 31, 2008

Showing Percentage of Net Assets

Common Stocks - 98.1%

Shares

Value (000s)

CONSUMER DISCRETIONARY - 4.9%

Media - 4.9%

Comcast Corp. Class A (a)

3,338,598

$ 60,629

ENERGY - 1.4%

Oil, Gas & Consumable Fuels - 1.4%

Spectra Energy Corp.

765,600

17,486

FINANCIALS - 0.2%

Diversified Financial Services - 0.2%

Hicks Acquisition Co. I, Inc. unit

293,900

2,924

INFORMATION TECHNOLOGY - 0.2%

IT Services - 0.2%

NeuStar, Inc. Class A (a)

74,600

2,216

TELECOMMUNICATION SERVICES - 40.7%

Diversified Telecommunication Services - 37.7%

AT&T, Inc.

7,515,563

289,275

Embarq Corp.

181,700

8,231

Level 3 Communications, Inc. (a)

1,851,900

6,371

Qwest Communications International, Inc.

2,077,300

12,215

Time Warner Telecom, Inc. Class A (sub. vtg.) (a)

550,600

9,624

Verizon Communications, Inc.

3,395,241

131,871

Windstream Corp.

608,920

7,070

464,657

Wireless Telecommunication Services - 3.0%

Leap Wireless International, Inc. (a)

63,300

2,617

NII Holdings, Inc. (a)

209,300

8,929

Sprint Nextel Corp.

1,790,847

18,858

Telephone & Data Systems, Inc.

120,596

6,360

36,764

TOTAL TELECOMMUNICATION SERVICES

501,421

UTILITIES - 50.7%

Electric Utilities - 27.2%

Allegheny Energy, Inc.

463,500

25,395

American Electric Power Co., Inc.

807,900

34,602

Edison International

458,100

23,894

Entergy Corp.

312,302

33,785

Exelon Corp.

1,005,809

76,633

FirstEnergy Corp.

386,800

27,548

FPL Group, Inc.

495,100

31,924

Common Stocks - continued

Shares

Value (000s)

UTILITIES - continued

Electric Utilities - continued

Great Plains Energy, Inc.

115,500

$ 3,220

Northeast Utilities

187,300

5,192

Pepco Holdings, Inc.

232,000

5,907

PPL Corp.

1,216,727

59,522

Sierra Pacific Resources

288,200

4,314

Westar Energy, Inc.

117,600

2,865

334,801

Gas Utilities - 1.9%

Equitable Resources, Inc.

96,200

5,363

Questar Corp.

216,700

11,032

Southern Union Co.

254,700

6,923

23,318

Independent Power Producers & Energy Traders - 12.3%

AES Corp. (a)

3,457,384

65,967

Constellation Energy Group, Inc.

601,663

56,532

Mirant Corp. (a)

308,473

11,364

NRG Energy, Inc. (a)

448,600

17,311

151,174

Multi-Utilities - 9.0%

Ameren Corp.

63,700

2,854

CenterPoint Energy, Inc.

444,300

7,113

CMS Energy Corp.

1,339,300

20,987

Integrys Energy Group, Inc.

97,100

4,721

Public Service Enterprise Group, Inc.

471,372

45,252

Sempra Energy

319,300

17,849

Xcel Energy, Inc.

618,370

12,856

111,632

Water Utilities - 0.3%

Aqua America, Inc.

165,764

3,304

TOTAL UTILITIES

624,229

TOTAL COMMON STOCKS

(Cost $1,165,253)

1,208,905

Money Market Funds - 1.2%

Shares

Value (000s)

Fidelity Cash Central Fund, 3.79% (b)
(Cost $15,197)

15,197,095

$ 15,197

TOTAL INVESTMENT PORTFOLIO - 99.3%

(Cost $1,180,450)

1,224,102

NET OTHER ASSETS - 0.7%

8,413

NET ASSETS - 100%

$ 1,232,515

Legend

(a) Non-income producing

(b) Affiliated fund that is available only to investment companies and other accounts managed by Fidelity Investments. The rate quoted is the annualized seven-day yield of the fund at period end. A complete unaudited listing of the fund's holdings as of its most recent quarter end is available upon request.

Affiliated Central Funds

Information regarding fiscal year to date income earned by the Fund from investments in Fidelity Central Funds is as follows:

Fund

Income earned
(Amounts in thousands)

Fidelity Cash Central Fund

$ 1,220

Fidelity Securities Lending Cash Central Fund

155

Total

$ 1,375

Income Tax Information

At January 31, 2008, the fund had a capital loss carryforward of approximately $210,412,000 of which $199,347,000 and $11,065,000 will expire on January 31, 2011 and 2012, respectively.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Statements

Statement of Assets and Liabilities

Amounts in thousands (except per-share amount)

January 31, 2008

Assets

Investment in securities, at value - See accompanying schedule:

Unaffiliated issuers (cost $1,165,253)

$ 1,208,905

Fidelity Central Funds (cost $15,197)

15,197

Total Investments (cost $1,180,450)

$ 1,224,102

Receivable for investments sold

19,791

Receivable for fund shares sold

668

Dividends receivable

4,975

Distributions receivable from Fidelity Central Funds

15

Prepaid expenses

4

Other receivables

35

Total assets

1,249,590

Liabilities

Payable for investments purchased

$ 14,643

Payable for fund shares redeemed

1,389

Accrued management fee

655

Other affiliated payables

278

Other payables and accrued expenses

110

Total liabilities

17,075

Net Assets

$ 1,232,515

Net Assets consist of:

Paid in capital

$ 1,418,736

Undistributed net investment income

1,887

Accumulated undistributed net realized gain (loss) on investments and foreign currency transactions

(231,760)

Net unrealized appreciation (depreciation) on investments

43,652

Net Assets, for 64,858 shares outstanding

$ 1,232,515

Net Asset Value, offering price and redemption price per share ($1,232,515 ÷ 64,858 shares)

$ 19.00

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Statements - continued

Statement of Operations

Amounts in thousands

Year ended January 31, 2008

Investment Income

Dividends

$ 37,582

Interest

11

Income from Fidelity Central Funds

1,375

Total income

38,968

Expenses

Management fee
Basic fee

$ 7,519

Performance adjustment

1,812

Transfer agent fees

3,289

Accounting and security lending fees

516

Custodian fees and expenses

32

Independent trustees' compensation

6

Registration fees

66

Audit

63

Legal

12

Interest

76

Miscellaneous

55

Total expenses before reductions

13,446

Expense reductions

(52)

13,394

Net investment income (loss)

25,574

Realized and Unrealized Gain (Loss)

Net realized gain (loss) on:

Investment securities:

Unaffiliated issuers

158,456

Foreign currency transactions

(1)

Total net realized gain (loss)

158,455

Change in net unrealized appreciation (depreciation) on:

Investment securities

(143,736)

Net gain (loss)

14,719

Net increase (decrease) in net assets resulting from operations

$ 40,293

See accompanying notes which are an integral part of the financial statements.

Annual Report

Statement of Changes in Net Assets

Amounts in thousands

Year ended
January 31,
2008

Year ended
January 31,
2007

Increase (Decrease) in Net Assets

Operations

Net investment income (loss)

$ 25,574

$ 24,313

Net realized gain (loss)

158,455

110,048

Change in net unrealized appreciation (depreciation)

(143,736)

183,257

Net increase (decrease) in net assets resulting
from operations

40,293

317,618

Distributions to shareholders from net investment income

(28,120)

(19,332)

Share transactions
Proceeds from sales of shares

637,065

930,039

Reinvestment of distributions

25,789

17,712

Cost of shares redeemed

(1,077,520)

(639,834)

Net increase (decrease) in net assets resulting from share transactions

(414,666)

307,917

Total increase (decrease) in net assets

(402,493)

606,203

Net Assets

Beginning of period

1,635,008

1,028,805

End of period (including undistributed net investment income of $1,887 and undistributed net investment income of $4,355, respectively)

$ 1,232,515

$ 1,635,008

Other Information

Shares

Sold

30,728

53,456

Issued in reinvestment of distributions

1,227

1,027

Redeemed

(51,842)

(36,363)

Net increase (decrease)

(19,887)

18,120

See accompanying notes which are an integral part of the financial statements.

Annual Report

Financial Highlights

Years ended January 31,

2008

2007

2006

2005

2004

Selected Per-Share Data

Net asset value, beginning of period

$ 19.29

$ 15.44

$ 13.28

$ 11.73

$ 9.44

Income from Investment Operations

Net investment income (loss) B

.32

.33

.22

.30 E

.21

Net realized and unrealized gain (loss)

(.24) F

3.77

2.20

1.54

2.30

Total from investment operations

.08

4.10

2.42

1.84

2.51

Distributions from net investment income

(.37)

(.25)

(.26)

(.29)

(.22)

Net asset value, end of period

$ 19.00

$ 19.29

$ 15.44

$ 13.28

$ 11.73

Total Return A

.24%

26.77%

18.37%

15.85%

26.91%

Ratios to Average Net Assets C,G

Expenses before reductions

.82%

.85%

.87%

.89%

.75%

Expenses net of fee waivers, if any

.82%

.85%

.87%

.89%

.75%

Expenses net of all reductions

.82%

.84%

.84%

.85%

.73%

Net investment income (loss)

1.56%

1.92%

1.54%

2.49%E

2.01%

Supplemental Data

Net assets, end of period (in millions)

$ 1,233

$ 1,635

$ 1,029

$ 937

$ 864

Portfolio turnover rate D

56%

104%

66%

57%

21%

A Total returns would have been lower had certain expenses not been reduced during the periods shown.

B Calculated based on average shares outstanding during the period.

C Fees and expenses of the underlying Fidelity Central Funds are not included in the Fund's expense ratio. The Fund indirectly bears its proportionate share of the expenses of any underlying Fidelity Central Funds.

D Amount does not include the portfolio activity of any underlying Fidelity Central Funds.

E Investment income per share reflects a special dividend which amounted to $.12 per share. Excluding the special dividend, the ratio of net investment income (loss) to average net assets would have been 1.48%.

F The amount shown for a share outstanding does not correspond with the aggregate net gain (loss) on investments for the period due to the timing of sales and repurchases of shares in relation to fluctuating market values of the investments of the Fund.

G Expense ratios reflect operating expenses of the Fund. Expenses before reductions do not reflect amounts reimbursed by the investment adviser or reductions from brokerage service arrangements or other expense offset arrangements and do not represent the amount paid by the Fund during periods when reimbursements or reductions occur. Expenses net of fee waivers reflect expenses after reimbursement by the investment adviser but prior to reductions from brokerage service arrangements or other expense offset arrangements. Expenses net of all reductions represent the net expenses paid by the Fund.

See accompanying notes which are an integral part of the financial statements.

Annual Report

Notes to Financial Statements

For the period ended January 31, 2008

(Amounts in thousands except ratios)

1. Organization.

Fidelity Utilities Fund (the Fund) is a non-diversified fund of Fidelity Devonshire Trust (the trust) and is authorized to issue an unlimited number of shares. The trust is registered under the Investment Company Act of 1940, as amended (the 1940 Act), as an open-end management investment company organized as a Massachusetts business trust.

2. Investments in Fidelity Central Funds.

The Fund may invest in Fidelity Central Funds, which are open-end investment companies available only to other investment companies and accounts managed by Fidelity Management & Research Company (FMR) and its affiliates. The Fund's Schedule of Investments lists each of the Fidelity Central Funds as an investment of the Fund, but does not include the underlying holdings of each Fidelity Central Fund. As an Investing Fund, the Fund indirectly bears its proportionate share of the expenses of the underlying Fidelity Central Funds.

The Money Market Central Funds seek preservation of capital and current income and are managed by Fidelity Investments Money Management, Inc. (FIMM), an affiliate of FMR.

A complete list of holdings for each Fidelity Central Fund is available upon request or at the SEC's web site at www.sec.gov. In addition, the financial statements of the Fidelity Central Funds are available on the SEC's web site or upon request.

3. Significant Accounting Policies.

The financial statements have been prepared in conformity with accounting principles generally accepted in the United States of America, which require management to make certain estimates and assumptions at the date of the financial statements. Actual results could differ from those estimates. The following summarizes the significant accounting policies of the Fund:

Security Valuation. Investments are valued and net asset value (NAV) per share is calculated (NAV calculation) as of the close of business of the New York Stock Exchange (NYSE), normally 4:00 p.m. Eastern time. Wherever possible, the Fund uses independent pricing services approved by the Board of Trustees to value its investments.

Equity securities, including restricted securities, for which market quotations are readily available, are valued at the last reported sale price or official closing price as reported by an independent pricing service on the primary market or exchange on which they are traded. In the event there were no sales during the day or closing prices are not available, securities are valued at the last quoted bid price. Investments in open-end mutual funds, including the Fidelity Central Funds, are valued at their closing net asset value each

Annual Report

Notes to Financial Statements - continued

(Amounts in thousands except ratios)

3. Significant Accounting Policies - continued

Security Valuation - continued

business day. Short-term securities with remaining maturities of sixty days or less for which quotations are not readily available are valued at amortized cost, which approximates value.

When current market prices or quotations are not readily available or do not accurately reflect fair value, valuations may be determined in accordance with procedures adopted by the Board of Trustees. For example, when developments occur between the close of a market and the close of the NYSE that may materially affect the value of some or all of the securities, or when trading in a security is halted, those securities may be fair valued. Factors used in the determination of fair value may include monitoring news to identify significant market or security specific events such as changes in the value of U.S. securities markets, reviewing developments in foreign markets and evaluating the performance of ADRs, futures contracts and exchange-traded funds. Because the Fund's utilization of fair value pricing depends on market activity, the frequency with which fair value pricing is used cannot be predicted and may be utilized to a significant extent. The value of securities used for NAV calculation under fair value pricing may differ from published prices for the same securities.

Foreign Currency. The Fund uses foreign currency contracts to facilitate transactions in foreign-denominated securities. Losses from these transactions may arise from changes in the value of the foreign currency or if the counterparties do not perform under the contracts' terms.

Foreign-denominated assets, including investment securities, and liabilities are translated into U.S. dollars at the exchange rate at period end. Purchases and sales of investment securities, income and dividends received and expenses denominated in foreign currencies are translated into U.S. dollars at the exchange rate in effect on the transaction date.

The effects of exchange rate fluctuations on investments are included with the net realized and unrealized gain (loss) on investment securities. Other foreign currency transactions resulting in realized and unrealized gain (loss) are disclosed separately.

Investment Transactions and Income. For financial reporting purposes, the Fund's investment holdings and NAV include trades executed through the end of the last business day of the period. The NAV for processing shareholder transactions includes trades executed through the end of the prior business day. Gains and losses on securities sold are determined on the basis of identified cost and may include proceeds received from litigation. Dividend income is recorded on the ex-dividend date, except for certain dividends from foreign securities where the ex-dividend date may have passed, which are recorded as soon as the Fund is informed of the ex-dividend date. Non-cash dividends

Annual Report

3. Significant Accounting Policies - continued

Investment Transactions and Income - continued

included in dividend income, if any, are recorded at the fair market value of the securities received. Distributions received on securities that represent a return of capital or capital gain are recorded as a reduction of cost of investments and/or as a realized gain. The Fund estimates the components of distributions received that may be considered return of capital distributions or capital gain distributions. Interest income and distributions from the Fidelity Central Funds are accrued as earned. Interest income includes coupon interest and amortization of premium and accretion of discount on debt securities. Investment income is recorded net of foreign taxes withheld where recovery of such taxes is uncertain.

Expenses. Most expenses of the trust can be directly attributed to a fund. Expenses which cannot be directly attributed are apportioned among each Fund in the trust. Expense estimates are accrued in the period to which they relate and adjustments are made when actual amounts are known.

Deferred Trustee Compensation. Under a Deferred Compensation Plan (the Plan), Independent Trustees must defer receipt of a portion of, and may elect to defer receipt of an additional portion of, their annual compensation. Deferred amounts are invested in a cross-section of Fidelity funds, are marked-to-market and remain in the Fund until distributed in accordance with the Plan. The investment of deferred amounts and the offsetting payable to the Trustees are included in the accompanying Statement of Assets and Liabilities.

Income Tax Information and Distributions to Shareholders. Each year, the Fund intends to qualify as a regulated investment company by distributing substantially all of its taxable income and realized gains under Subchapter M of the Internal Revenue Code and filing its U.S. federal tax return. As a result, no provision for income taxes is required. The Fund adopted the provisions of FASB Interpretation No. 48, Accounting for Uncertainties in Income Taxes (FIN 48), on June 29, 2007. FIN 48 sets forth a minimum threshold for financial statement recognition of the benefit of a tax position taken or expected to be taken in a tax return. The implementation of FIN 48 did not result in any unrecognized tax benefits in the accompanying financial statements. Each of the Fund's federal tax returns for the prior three fiscal years remains subject to examination by the Internal Revenue Service. Foreign taxes are provided for based on the Fund's understanding of the tax rules and rates that exist in the foreign markets in which it invests.

Distributions are recorded on the ex-dividend date. Income and capital gain distributions are determined in accordance with income tax regulations, which may differ from generally accepted accounting principles.

Annual Report

Notes to Financial Statements - continued

(Amounts in thousands except ratios)

3. Significant Accounting Policies - continued

Income Tax Information and Distributions to Shareholders - continued

Capital accounts within the financial statements are adjusted for permanent book-tax differences. These adjustments have no impact on net assets or the results of operations. Temporary book-tax differences will reverse in a subsequent period.

Book-tax differences are primarily due to deferred trustees compensation, foreign currency transactions, capital loss carryforwards and losses deferred due to wash sales.

The tax-basis components of distributable earnings and the federal tax cost as of period end were as follows:

Unrealized appreciation

$ 136,238

Unrealized depreciation

(113,933)

Net unrealized appreciation (depreciation)

22,305

Undistributed ordinary income

1,926

Capital loss carryforward

(210,412)

Cost for federal income tax purposes

$ 1,201,797

The tax character of distributions paid was as follows:

January 31, 2008

January 31, 2007

Ordinary Income

$ 28,120

$ 19,332

New Accounting Pronouncement. In September 2006, Statement of Financial Accounting Standards No. 157, Fair Value Measurements (SFAS 157), was issued and is effective for fiscal years beginning after November 15, 2007. SFAS 157 defines fair value, establishes a framework for measuring fair value and expands disclosures about fair value measurements. Management is currently evaluating the impact the adoption of SFAS 157 will have on the Fund's financial statement disclosures.

4. Operating Policies.

Repurchase Agreements. FMR has received an Exemptive Order from the Securities and Exchange Commission (the SEC) which permits the Fund and other affiliated entities of FMR to transfer uninvested cash balances into joint trading accounts which are then invested in repurchase agreements. The Fund may also invest directly with institutions in repurchase agreements. Repurchase agreements are collateralized by government or non-government securities. Upon settlement date, collateral is held in segregated accounts with custodian banks and may be obtained in the event of a default of the counterparty. The Fund monitors, on a daily basis, the value of the collateral to ensure it is at least equal to the principal amount of the repurchase agreement (including accrued interest). In the event of a default by the counterparty, realization of the

Annual Report

4. Operating Policies - continued

Repurchase Agreements - continued

collateral proceeds could be delayed, during which time the value of the collateral may decline.

5. Purchases and Sales of Investments.

Purchases and sales of securities, other than short-term securities, aggregated $897,238 and $1,287,759, respectively.

6. Fees and Other Transactions with Affiliates.

Management Fee. FMR and its affiliates provide the Fund with investment management related services for which the Fund pays a monthly management fee. The management fee is the sum of an individual fund fee rate that is based on an annual rate of .20% of the Fund's average net assets and a group fee rate that averaged .26% during the period. The group fee rate is based upon the average net assets of all the mutual funds advised by FMR. The group fee rate decreases as assets under management increase and increases as assets under management decrease. In addition, the management fee is subject to a performance adjustment (up to a maximum of ± .15% of the Fund's average net assets over a 36 month performance period). The upward or downward adjustment to the management fee is based on the Fund's relative investment performance as compared to an appropriate benchmark index. For the period, the total annual management fee rate, including the performance adjustment, was .57% of the Fund's average net assets.

Transfer Agent Fees. Fidelity Investments Institutional Operations Company, Inc. (FIIOC), an affiliate of FMR, is the Fund's transfer, dividend disbursing and shareholder servicing agent. FIIOC receives account fees and asset-based fees that vary according to account size and type of account. FIIOC pays for typesetting, printing and mailing of shareholder reports, except proxy statements. For the period the transfer agent fees were equivalent to an annual rate of .20% of average net assets. Prior to January 1, 2008 Fidelity Service Company, Inc. (FSC), also an affiliate of FMR was the Fund's transfer agent.

Accounting and Security Lending Fees. FSC maintains the Fund's accounting records. The accounting fee is based on the level of average net assets for the month. Under a separate contract, FSC administers the security lending program. The security lending fee is based on the number and duration of lending transactions.

Brokerage Commissions. The Fund placed a portion of its portfolio transactions with brokerage firms which are affiliates of the investment adviser. The commissions paid to these affiliated firms were $11 for the period.

Annual Report

Notes to Financial Statements - continued

(Amounts in thousands except ratios)

6. Fees and Other Transactions with Affiliates - continued

Interfund Lending Program. Pursuant to an Exemptive Order issued by the SEC, the Fund, along with other registered investment companies having management contracts with FMR, may participate in an interfund lending program. This program provides an alternative credit facility allowing the funds to borrow from, or lend money to, other participating affiliated funds. At period end, there were no interfund loans outstanding. The Fund's activity in this program during the period for which loans were outstanding was as follows:

Borrower or Lender

Average Daily Loan Balance

Weighted Average Interest Rate

Interest
Expense

Borrower

$ 7,362

4.93%

$ 49

7. Committed Line of Credit.

The Fund participates with other funds managed by FMR in a $4.2 billion credit facility (the "line of credit") to be utilized for temporary or emergency purposes to fund shareholder redemptions or for other short-term liquidity purposes. The Fund has agreed to pay commitment fees on its pro rata portion of the line of credit, which amounted to $3 and is reflected in Miscellaneous Expense on the Statement of Operations. During the period, there were no borrowings on this line of credit.

8. Security Lending.

The Fund lends portfolio securities from time to time in order to earn additional income. On the settlement date of the loan, the Fund receives collateral (in the form of U.S. Treasury obligations, letters of credit and/or cash) against the loaned securities and maintains collateral in an amount not less than 100% of the market value of the loaned securities during the period of the loan. The market value of the loaned securities is determined at the close of business of the Fund and any additional required collateral is delivered to the Fund on the next business day. If the borrower defaults on its obligation to return the securities loaned because of insolvency or other reasons, a fund could experience delays and costs in recovering the securities loaned or in gaining access to the collateral. Any cash collateral received is invested in the Fidelity Securities Lending Cash Central Fund. At period end, there were no security loans outstanding. Security lending income represents the income earned on investing cash collateral, less fees and expenses associated with the loan, plus any premium payments that may be received on the loan of certain types of securities. Security lending income is presented in the Statement of Operations as a component of income from Fidelity Central Funds. Net income from lending portfolio securities during the period amounted to $155.

Annual Report

9. Bank Borrowings.

The Fund is permitted to have bank borrowings for temporary or emergency purposes to fund shareholder redemptions. The Fund has established borrowing arrangements with certain banks. The interest rate on the borrowings is the bank's base rate, as revised from time to time. The average daily loan balance during the period for which loans were outstanding amounted to $11,505. The weighted average interest rate was 5.54%. The interest expense amounted to $27 under the bank borrowing program. At period end, there were no bank borrowings outstanding.

10. Expense Reductions.

Many of the brokers with whom FMR places trades on behalf of the Fund provided services to the Fund in addition to trade execution. These services included payments of certain expenses on behalf of the Fund totaling $2 for the period. In addition, through arrangements with the Fund's transfer agent, credits realized as a result of uninvested cash balances were used to reduce the Fund's expenses. During the period, these credits reduced the Fund's expenses by $49, respectively.

11. Other.

The Fund's organizational documents provide former and current trustees and officers with a limited indemnification against liabilities arising in connection with the performance of their duties to the Fund. In the normal course of business, the Fund may also enter into contracts that provide general indemnifications. The Fund's maximum exposure under these arrangements is unknown as this would be dependent on future claims that may be made against the Fund. The risk of material loss from such claims is considered remote.

In December 2006, the Independent Trustees, with the assistance of independent counsel, completed an investigation regarding gifts, gratuities and business entertainment provided by certain brokers to certain individuals who were employed on FMR's domestic equity trading desk during the period 2002 to 2004. The Independent Trustees and FMR agreed that, despite the absence of proof that the Fidelity mutual funds experienced diminished execution quality as a result of the improper receipt of gifts and business entertainment, the conduct at issue was serious and was worthy of redress. Accordingly, the Independent Trustees requested, and FMR agreed to make a payment of $42 million plus accrued interest, which equaled approximately $7.3 million, to certain Fidelity mutual funds. FMR also reimbursed the related legal expenses, which are recorded in the accompanying Statement of Operations as an expense reduction.

Subsequent to period end, the Trustees approved a method for allocating this payment among the funds and, in total, FMR paid the fund $81.

In a related administrative order dated March 5, 2008, the U.S. Securities and Exchange Commission ("SEC") announced a settlement with FMR and FMR Co., Inc. (an affiliate

Annual Report

Notes to Financial Statements - continued

(Amounts in thousands except ratios)

11. Other - continued

of FMR) involving the SEC's regulatory rules for investment advisers and the improper receipt of gifts, gratuities and business entertainment. Without admitting or denying the SEC's findings, FMR agreed to pay an $8 million civil penalty to the United States Treasury.

Annual Report

Report of Independent Registered Public Accounting Firm

To the Trustees of Fidelity Devonshire Trust and the Shareholders of Fidelity Utilities Fund:

In our opinion, the accompanying statement of assets and liabilities, including the schedule of investments, and the related statements of operations and of changes in net assets and the financial highlights present fairly, in all material respects, the financial position of Fidelity Utilities Fund (a fund of Fidelity Devonshire Trust) at January 31, 2008, the results of its operations for the year then ended, the changes in its net assets for each of the two years in the period then ended and the financial highlights for each of the five years in the period then ended, in conformity with accounting principles generally accepted in the United States of America. These financial statements and financial highlights (hereafter referred to as "financial statements") are the responsibility of the Fidelity Utilities Fund's management. Our responsibility is to express an opinion on these financial statements based on our audits. We conducted our audits of these financial statements in accordance with the standards of the Public Company Accounting Oversight Board (United States). Those standards require that we plan and perform the audit to obtain reasonable assurance about whether the financial statements are free of material misstatement. An audit includes examining, on a test basis, evidence supporting the amounts and disclosures in the financial statements, assessing the accounting principles used and significant estimates made by management, and evaluating the overall financial statement presentation. We believe that our audits, which included confirmation of securities at January 31, 2008 by correspondence with the custodian and brokers, provide a reasonable basis for our opinion.

/s/ PricewaterhouseCoopers LLP

PricewaterhouseCoopers LLP

Boston, Massachusetts

March 24, 2008

Annual Report

Trustees and Officers

The Trustees, Members of the Advisory Board, and executive officers of the trust and fund, as applicable, are listed below. The Board of Trustees governs the fund and is responsible for protecting the interests of shareholders. The Trustees are experienced executives who meet periodically throughout the year to oversee the fund's activities, review contractual arrangements with companies that provide services to the fund, and review the fund's performance. Except for James C. Curvey, each of the Trustees oversees 374 funds advised by FMR or an affiliate. Mr. Curvey oversees 369 funds advised by FMR or an affiliate.

The Trustees hold office without limit in time except that (a) any Trustee may resign; (b) any Trustee may be removed by written instrument, signed by at least two-thirds of the number of Trustees prior to such removal; (c) any Trustee who requests to be retired or who has become incapacitated by illness or injury may be retired by written instrument signed by a majority of the other Trustees; and (d) any Trustee may be removed at any special meeting of shareholders by a two-thirds vote of the outstanding voting securities of the trust. Each Trustee who is not an interested person (as defined in the 1940 Act) (Independent Trustee), shall retire not later than the last day of the calendar year in which his or her 72nd birthday occurs. The Independent Trustees may waive this mandatory retirement age policy with respect to individual Trustees. The executive officers and Advisory Board Members hold office without limit in time, except that any officer and Advisory Board Member may resign or may be removed by a vote of a majority of the Trustees at any regular meeting or any special meeting of the Trustees. Except as indicated, each individual has held the office shown or other offices in the same company for the past five years.

The fund's Statement of Additional Information (SAI) includes more information about the Trustees. To request a free copy, call Fidelity at 1-800-544-8544.

Interested Trustees*:

Correspondence intended for each Trustee who is an interested person may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Edward C. Johnson 3d (77)

Year of Election or Appointment: 1985

Mr. Johnson is Chairman of the Board of Trustees. Mr. Johnson serves as President (2006-present), Chief Executive Officer, Chairman, and a Director of FMR LLC; Chairman and a Director of FMR; Chairman and a Director of Fidelity Research & Analysis Company (FRAC); Chairman and a Director of Fidelity Investments Money Management, Inc.; and Chairman (2001-present) and a Director of FMR Co., Inc. In addition, Mr. Johnson serves as Chairman and Director of FIL Limited. Mr. Edward C. Johnson 3d and Mr. Arthur E. Johnson are not related.

James C. Curvey (72)

Year of Election or Appointment: 2007

Mr. Curvey also serves as Trustee (2007-present) or Member of the Advisory Board (2007-present) of other investment companies advised by FMR. Mr. Curvey is a Director of FMR and FMR Co., Inc. (2007-
present). Mr. Curvey is also Vice Chairman (2006-present) and Director of FMR LLC. Mr. Curvey joined Fidelity in 1982 and served in numerous senior management positions, including President and Chief Operating Officer of FMR LLC (1997-2000) and President of Fidelity Strategic Investments (2000-2002). In addition, he serves as a member of the Board of Directors of Geerlings & Wade, Inc. (wine distribution).

* Trustees have been determined to be "Interested Trustees" by virtue of, among other things, their affiliation with the trust or various entities under common control with FMR. FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Independent Trustees:

Correspondence intended for each Independent Trustee (that is, the Trustees other than the Interested Trustees) may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235.

Name, Age; Principal Occupation

Dennis J. Dirks (59)

Year of Election or Appointment: 2005

Prior to his retirement in May 2003, Mr. Dirks was Chief Operating Officer and a member of the Board of The Depository Trust & Clearing Corporation (DTCC) (1999-2003). He also served as President, Chief Operating Officer, and Board member of The Depository Trust Company (DTC) (1999-2003) and President and Board member of the National Securities Clearing Corporation (NSCC) (1999-2003). In addition, Mr. Dirks served as Chief Executive Officer and Board member of the Government Securities Clearing Corporation (2001-2003) and Chief Executive Officer and Board member of the Mortgage-Backed Securities Clearing Corporation (2001-2003). Mr. Dirks also serves as a Trustee and a member of the Finance Committee of Manhattan College (2005-
present) and a Trustee and a member of the Finance Committee of AHRC of Nassau County (2006-present).

Albert R. Gamper, Jr. (65)

Year of Election or Appointment: 2006

Prior to his retirement in December 2004, Mr. Gamper served as Chairman of the Board of CIT Group Inc. (commercial finance). During his tenure with CIT Group Inc. Mr. Gamper served in numerous senior management positions, including Chairman (1987-1989; 1999-2001; 2002-2004), Chief Executive Officer (1987-2004), and President (1989-2002). He currently serves as a member of the Board of Directors of Public Service Enterprise Group (utilities, 2001-present), Chairman of the Board of Governors, Rutgers University (2004-present), and Chairman of the Board of Saint Barnabas Health Care System.

George H. Heilmeier (71)

Year of Election or Appointment: 2004

Dr. Heilmeier is Chairman Emeritus of Telcordia Technologies (communication software and systems), where prior to his retirement, he served as company Chairman and Chief Executive Officer. He currently serves on the Boards of Directors of The Mitre Corporation (systems engineering and information technology support for the government), and HRL Laboratories (private research and development, 2004-present). He is Chairman of the General Motors Science & Technology Advisory Board and a Life Fellow of the Institute of Electrical and Electronics Engineers (IEEE). Dr. Heilmeier is a member of the Defense Science Board and the National Security Agency Advisory Board. He is also a member of the National Academy of Engineering, the American Academy of Arts and Sciences, and the Board of Overseers of the School of Engineering and Applied Science of the University of Pennsylvania. Previously, Dr. Heilmeier served as a Director of TRW Inc. (automotive, space, defense, and information technology, 1992-2002), Compaq (1994-2002), Automatic Data Processing, Inc. (ADP) (technology-based business outsourcing, 1995-2002), INET Technologies Inc. (telecommunications network surveillance, 2001-2004), and Teletech Holdings (customer management services). He is the recipient of the 2005 Kyoto Prize in Advanced Technology for his invention of the liquid crystal display, and a member of the Consumer Electronics Hall of Fame.

James H. Keyes (67)

Year of Election or Appointment: 2007

Prior to his retirement in 2003, Mr. Keyes was Chairman, President, and Chief Executive Officer of Johnson Controls, Inc. (automotive supplier, 1993-2003). He currently serves as a member of the boards of LSI Logic Corporation (semiconductor technologies), Navistar International Corporation (manufacture and sale of trucks, buses, and diesel engines, 2002-present), and Pitney Bowes, Inc. (integrated mail, messaging, and document management solutions).

Marie L. Knowles (61)

Year of Election or Appointment: 2001

Prior to Ms. Knowles' retirement in June 2000, she served as Executive Vice President and Chief Financial Officer of Atlantic Richfield Company (ARCO) (diversified energy, 1996-2000). From 1993 to 1996, she was a Senior Vice President of ARCO and President of ARCO Transportation Company. She served as a Director of ARCO from 1996 to 1998. She currently serves as a Director of Phelps Dodge Corporation (copper mining and manufacturing) and McKesson Corporation (healthcare service, 2002-present). Ms. Knowles is a Trustee of the Brookings Institution and the Catalina Island Conservancy and also serves as a member of the Advisory Board for the School of Engineering of the University of Southern California.

Ned C. Lautenbach (63)

Year of Election or Appointment: 2000

Mr. Lautenbach is Chairman of the Independent Trustees (2006-present). Mr. Lautenbach has been a partner of Clayton, Dubilier & Rice, Inc. (private equity investment firm) since September 1998. Previously, Mr. Lautenbach was with the International Business Machines Corporation (IBM) from 1968 until his retirement in 1998. Mr. Lautenbach serves as a Director of Sony Corporation (2006-present) and Eaton Corporation (diversified industrial) as well as the Philharmonic Center for the Arts in Naples, Florida. He also is a member of the Board of Trustees of Fairfield University (2005-present), as well as a member of the Council on Foreign Relations.

Cornelia M. Small (63)

Year of Election or Appointment: 2005

Ms. Small is a member (2000-present) and Chairperson (2002-present) of the Investment Committee, and a member (2002-present) of the Board of Trustees of Smith College. Previously, she served as Chief Investment Officer (1999-2000), Director of Global Equity Investments (1996-1999), and a member of the Board of Directors of Scudder, Stevens & Clark (1990-1997) and Scudder Kemper Investments (1997-
1999). In addition, Ms. Small served as Co-Chair (2000-2003) of the Annual Fund for the Fletcher School of Law and Diplomacy.

William S. Stavropoulos (68)

Year of Election or Appointment: 2002

Mr. Stavropoulos is Chairman Emeritus of the Board of Directors of The Dow Chemical Company. Since joining The Dow Chemical Company in 1967, Mr. Stavropoulos served in numerous senior management positions, including President (1993-2000; 2002-2003), CEO (1995-2000; 2002-2004), and Chairman of the Executive Committee (2000-2004). Currently, he is a Director of NCR Corporation (data warehousing and technology solutions), Chemical Financial Corporation, Maersk Inc. (industrial conglomerate, 2002-present), Tyco International, Inc. (multinational manufacturing and services, 2007-present), and a member of the Advisory Board for Metalmark Capital (private equity investment firm, 2005-present). He is a special advisor to Clayton, Dubilier & Rice, Inc., a private equity investment firm. He also serves as a member of the Board of Trustees of the American Enterprise Institute for Public Policy Research. In addition, Mr. Stavropoulos is a member of The Business Council, J.P. Morgan International Council and the University of Notre Dame Advisory Council for the College of Science.

Kenneth L. Wolfe (68)

Year of Election or Appointment: 2005

Mr. Wolfe is Chairman and a Director of Hershey Foods Corporation (2007-present), where prior to his retirement in 2001, he was Chairman and Chief Executive Officer. Mr. Wolfe currently serves as a member of the board of Revlon Inc. (2004-present). Previously, Mr. Wolfe served as a member of the boards of Adelphia Communications Corporation (2003-2006) and Bausch & Lomb, Inc. (1993-2007).

Advisory Board Members and Executive Officers**:

Correspondence intended for Mr. Mauriello, Mr. Thomas, Mr. Wiley, Mr. Lacy, and Mr. Arthur Johnson may be sent to Fidelity Investments, P.O. Box 55235, Boston, Massachusetts 02205-5235. Correspondence intended for each executive officer and Mr. Lynch may be sent to Fidelity Investments, 82 Devonshire Street, Boston, Massachusetts 02109.

Name, Age; Principal Occupation

Arthur E. Johnson (61)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Johnson serves as Senior Vice President of Corporate Strategic Development of Lockheed Martin Corporation (defense contractor). In addition, Mr. Johnson serves as a member of the Board of Directors of AGL Resources, Inc. (holding company, 2002-present), and IKON Office Solutions, Inc. (document management systems and services). Mr.
Arthur E. Johnson and Mr. Edward C. Johnson 3d are not related.

Name, Age; Principal Occupation

Alan J. Lacy (54)

Year of Election or Appointment: 2008

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lacy serves as Senior Adviser (2007-present) of Oak Hill Capital Partners, L.P. (a private equity firm). Mr. Lacy also served as Vice Chairman and Chief Executive Officer of Sears Holdings Corporation and Sears, Roebuck and Co. (retail, 2005-2006; 2000-2005). In addition, Mr. Lacy serves as a member of the Board of Directors of The Western Union Company (global money transfer, 2006-present) and Bristol-Myers Squibb (global pharmaceuticals, 2007-present). Mr. Lacy is a Trustee of the National Parks Conservation Association and The Field Museum of Natural History.

Peter S. Lynch (64)

Year of Election or Appointment: 2003

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Lynch is Vice Chairman and a Director of FMR, and Vice Chairman (2001-
present) and a Director of FMR Co., Inc. Previously, Mr. Lynch served as a Trustee of the Fidelity funds (1990-2003). In addition, he serves as a Trustee of Boston College and as the Chairman of the Inner-City Scholarship Fund.

Joseph Mauriello (63)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Prior to his retirement in January 2006, Mr. Mauriello served in numerous senior management positions including Deputy Chairman and Chief Operating Officer (2004-2005), and Vice Chairman of Financial Services (2002-
2004) of KPMG LLP US (professional services firm, 1965-2005). Mr. Mauriello currently serves as a member of the Board of Directors of XL Capital Ltd. (global insurance and re-insurance company, 2006-present) and of Arcadia Resources Inc. (health care services and products, 2007-present). He also served as a Director of the Hamilton Funds of the Bank of New York (2006-2007).

David M. Thomas (58)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Previously, Mr. Thomas served as Executive Chairman (2005-2006) and Chairman and Chief Executive Officer (2000-2005) of IMS Health, Inc. (pharmaceutical and healthcare information solutions). In addition, Mr. Thomas serves as a member of the Board of Directors of Fortune Brands, Inc. (consumer products holding company), and Interpublic Group of Companies, Inc. (marketing communication, 2004-present).

Michael E. Wiley (57)

Year of Election or Appointment: 2007

Member of the Advisory Board of Fidelity Devonshire Trust. Mr. Wiley also serves as a member of the Board of Trustees of the University of Tulsa (2000-2006, 2007-present). He serves as a Director of Tesoro Corporation (independent oil refiner and marketer, 2005-present), and a Director of Bill Barrett Corporation (exploration and production company, 2005-present). In addition, he also serves as a Director of Post Oak Bank (privately-held bank, 2004-present). Previously, Mr. Wiley served as a Sr. Energy Advisor of Katzenbach Partners, LLC (consulting firm, 2006-2007), as an Advisory Director of Riverstone Holdings (private investment firm), Chairman, President, and CEO of Baker Hughes, Inc. (oilfield services company, 2000-2004), and as Director of Spinnaker Exploration Company (exploration and production company, 2001-2005).

Kimberley H. Monasterio (44)

Year of Election or Appointment: 2007

President and Treasurer of Utilities. Ms. Monasterio also serves as President and Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2004-present). Previously, Ms. Monasterio served as Deputy Treasurer of the Fidelity funds (2004-2006). Before joining Fidelity Investments, Ms. Monasterio served as Treasurer (2000-2004) and Chief Financial Officer (2002-2004) of the Franklin Templeton Funds and Senior Vice President of Franklin Templeton Services, LLC (2000-2004).

Brian B. Hogan (43)

Year of Election or Appointment: 2007

Vice President of Utilities. Mr. Hogan also serves as Vice President of Sector Funds (2007-present). Mr. Hogan is Senior Vice President of Equity Research (2006-present). Mr. Hogan also serves as Vice President of FMR and FMR Co., Inc. Previously, Mr. Hogan served as a portfolio manager.

Eric D. Roiter (59)

Year of Election or Appointment: 1998

Secretary of Utilities. He also serves as Secretary of other Fidelity funds; Vice President, General Counsel, and Secretary of FMR Co., Inc. (2001-
present) and FMR; Assistant Secretary of Fidelity Management & Research (U.K.) Inc. (2001-present), Fidelity Research & Analysis Company (2001-present), and Fidelity Investments Money Management, Inc. (2001-present). Mr. Roiter is an Adjunct Member, Faculty of Law, at Boston College Law School (2003-present). Previously, Mr. Roiter served as Vice President and Secretary of Fidelity Distributors Corporation (FDC) (1998-2005).

John B. McGinty, Jr. (45)

Year of Election or Appointment: 2008

Assistant Secretary of Utilities. Mr. McGinty also serves as Assistant Secretary of other Fidelity funds (2008-present) and is an employee of FMR LLC (2004-present). Mr. McGinty also serves as Senior Vice President, Secretary, and Chief Legal Officer of FDC (2007-present). Before joining Fidelity Investments, Mr. McGinty practiced law at Ropes & Gray, LLP.

R. Stephen Ganis (41)

Year of Election or Appointment: 2006

Anti-Money Laundering (AML) officer of Utilities. Mr. Ganis also serves as AML officer of the other Fidelity funds (2006-present) and FMR LLC (2003-present). Before joining Fidelity Investments, Mr. Ganis practiced law at Goodwin Procter, LLP (2000-2002).

Joseph B. Hollis (59)

Year of Election or Appointment: 2006

Chief Financial Officer of Utilities. Mr. Hollis also serves as Chief Financial Officer of other Fidelity funds. Mr. Hollis is President of Fidelity Pricing and Cash Management Services (FPCMS) (2005-present). Mr. Hollis also serves as President and Director of Fidelity Service Company, Inc. (2006-present). Previously, Mr. Hollis served as Senior Vice President of Cash Management Services (1999-2002) and Investment Management Operations (2002-2005).

Kenneth A. Rathgeber (60)

Year of Election or Appointment: 2004

Chief Compliance Officer of Utilities. Mr. Rathgeber also serves as Chief Compliance Officer of other Fidelity funds (2004-present) and Executive Vice President of Risk Oversight for Fidelity Investments (2002-present). He is Chief Compliance Officer of FMR (2005-present), FMR Co., Inc. (2005-present), Fidelity Management & Research (U.K.) Inc. (2005-
present), Fidelity Research & Analysis Company (2005-present), Fidelity Investments Money Management, Inc. (2005-present), and Strategic Advisers, Inc. (2005-present). Previously, Mr. Rathgeber served as Executive Vice President and Chief Operating Officer for Fidelity Investments Institutional Services Company, Inc. (1998-2002).

Bryan A. Mehrmann (46)

Year of Election or Appointment: 2005

Deputy Treasurer of Utilities. Mr. Mehrmann also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR. Previously, Mr. Mehrmann served as Vice President of Fidelity Investments Institutional Services Group (FIIS)/Fidelity Investments Institutional Operations Corporation, Inc. (FIIOC) Client Services (1998-2004).

Kenneth B. Robins (38)

Year of Election or Appointment: 2005

Deputy Treasurer of Utilities. Mr. Robins also serves as Deputy Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2004-present). Before joining Fidelity Investments, Mr. Robins worked at KPMG LLP, where he was a partner in KPMG's department of professional practice (2002-2004) and a Senior Manager (1999-2000). In addition, Mr. Robins served as Assistant Chief Accountant, United States Securities and Exchange Commission (2000-2002).

Robert G. Byrnes (41)

Year of Election or Appointment: 2005

Assistant Treasurer of Utilities. Mr. Byrnes also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Byrnes served as Vice President of FPCMS (2003-2005). Before joining Fidelity Investments, Mr. Byrnes worked at Deutsche Asset Management where he served as Vice President of the Investment Operations Group (2000-2003).

Peter L. Lydecker (54)

Year of Election or Appointment: 2004

Assistant Treasurer of Utilities. Mr. Lydecker also serves as Assistant Treasurer of other Fidelity funds (2004) and is an employee of FMR.

Paul M. Murphy (60)

Year of Election or Appointment: 2007

Assistant Treasurer of Utilities. Mr. Murphy also serves as Assistant Treasurer of other Fidelity funds (2007-present) and is an employee of FMR (2007-present). Previously, Mr. Murphy served as Chief Financial Officer of the Fidelity Funds (2005-2006), Vice President and Associate General Counsel of FMR (2007), and Senior Vice President of Fidelity Pricing and Cash Management Services Group (FPCMS) (1994-2007).

Gary W. Ryan (49)

Year of Election or Appointment: 2005

Assistant Treasurer of Utilities. Mr. Ryan also serves as Assistant Treasurer of other Fidelity funds (2005-present) and is an employee of FMR (2005-present). Previously, Mr. Ryan served as Vice President of Fund Reporting in FPCMS (1999-2005).

** FMR Corp. merged with and into FMR LLC on October 1, 2007. Any references to FMR LLC for prior periods are deemed to be references to the prior entity.

Annual Report

Distributions

A total of 0.18% of the dividends distributed during the fiscal year was derived from interest on U.S. Government securities which is generally exempt from state income tax.

The fund designates 100% of the dividends distributed during the fiscal year as qualifying for the dividends-received deduction for corporate shareholders.

The fund designates 100% of the dividend distributed in April, July, October, and December during the fiscal year as amounts which may be taken into account as a dividend for the purposes of the maximum rate under section 1(h)(11) of the Internal Revenue Code.

The fund will notify shareholders in January 2009 of amounts for use in preparing 2008 income tax returns.

Annual Report

Managing Your Investments

Fidelity offers several ways to conveniently manage your personal investments via your telephone or PC. You can access your account information, conduct trades and research your investments 24 hours a day.

By Phone

Fidelity Automated Service Telephone provides a single toll-free number to access account balances, positions, quotes and trading. It's easy to navigate the service, and on your first call, the system will help you create a personal identification number (PIN) for security.

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Fidelity Automated
Service Telephone (FAST
®)
1-800-544-5555

Press

1   For mutual fund and brokerage trading.

2   For quotes.*

3   For account balances and holdings.

4   To review orders and mutual
fund activity.

5   To change your PIN.

*0   To speak to a Fidelity representative.

By PC

Fidelity's web site on the Internet provides a wide range of information, including daily financial news, fund performance, interactive planning tools and news about Fidelity products and services.

(computer_graphic)

Fidelity's Web Site
www.fidelity.com

* When you call the quotes line, please remember that a fund's yield and return will vary and, except for money market funds, share price will also vary. This means that you may have a gain or loss when you sell your shares. There is no assurance that money market funds will be able to maintain a stable $1 share price; an investment in a money market fund is not insured or guaranteed by the U.S. government. Total returns are historical and include changes in share price, reinvestment of dividends and capital gains, and the effects of any sales charges.

Annual Report

To Write Fidelity

We'll give your correspondence immediate attention and send you written confirmation upon completion of your request.

(letter_graphic)

Making Changes
To Your Account

(such as changing name, address, bank, etc.)

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0002

(letter_graphic)

For Non-Retirement
Accounts

Buying shares

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0003

Overnight Express
Fidelity Investments
Attn: Distribution Services
100 Crosby Parkway - KC1H
Covington, KY 41015

Selling shares

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0035

Overnight Express
Fidelity Investments
Attn: Distribution Services
100 Crosby Parkway - KC1H
Covington, KY 41015

General Correspondence

Fidelity Investments
P.O. Box 500
Merrimack, NH 03054-0500

(letter_graphic)

For Retirement
Accounts

Buying shares

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0003

Selling shares

Fidelity Investments
P.O. Box 770001
Cincinnati, OH 45277-0035

Overnight Express
Fidelity Investments
Attn: Distribution Services
100 Crosby Parkway - KC1H
Covington, KY 41015

General Correspondence

Fidelity Investments
P.O. Box 500
Merrimack, NH 03054-0500

Annual Report

To Visit Fidelity

For directions and hours,
please call 1-800-544-9797.

Arizona

7001 West Ray Road
Chandler, AZ

15445 N. Scottsdale Road
Scottsdale, AZ

California

815 East Birch Street
Brea, CA

1411 Chapin Avenue
Burlingame, CA

851 East Hamilton Avenue
Campbell, CA

19200 Von Karman Avenue
Irvine, CA

601 Larkspur Landing Circle
Larkspur, CA

2000 Avenue of the Stars
Los Angeles, CA

27101 Puerta Real
Mission Viejo, CA

73-575 El Paseo
Palm Desert, CA

251 University Avenue
Palo Alto, CA

123 South Lake Avenue
Pasadena, CA

16656 Bernardo Ctr. Drive
Rancho Bernardo, CA

1220 Roseville Parkway
Roseville, CA

1740 Arden Way
Sacramento, CA

7676 Hazard Center Drive
San Diego, CA

11943 El Camino Real
San Diego, CA

8 Montgomery Street
San Francisco, CA

3793 State Street
Santa Barbara, CA

1200 Wilshire Boulevard
Santa Monica, CA

398 West El Camino Real
Sunnyvale, CA

111 South Westlake Blvd
Thousand Oaks, CA

21701 Hawthorne Boulevard
Torrance, CA

2001 North Main Street
Walnut Creek, CA

6326 Canoga Avenue
Woodland Hills, CA

Colorado

281 East Flatiron Circle
Broomfield, CO

1625 Broadway
Denver, CO

9185 Westview Road
Lone Tree, CO

Connecticut

48 West Putnam Avenue
Greenwich, CT

265 Church Street
New Haven, CT

300 Atlantic Street
Stamford, CT

29 South Main Street
West Hartford, CT

Delaware

400 Delaware Avenue
Wilmington, DE

Florida

175 East Altamonte Drive
Altamonte Springs, FL

4400 N. Federal Highway
Boca Raton, FL

121 Alhambra Plaza
Coral Gables, FL

2948 N. Federal Highway
Ft. Lauderdale, FL

4671 Town Center Parkway
Jacksonville, FL

8880 Tamiami Trail, North
Naples, FL

230 Royal Palm Way
Palm Beach, FL

3501 PGA Boulevard
Palm Beach Gardens, FL

3550 Tamiami Trail, South
Sarasota, FL

1502 N. Westshore Blvd.
Tampa, FL

2465 State Road 7
Wellington, FL

Georgia

3445 Peachtree Road, N.E.
Atlanta, GA

1000 Abernathy Road
Atlanta, GA

Illinois

One North LaSalle Street
Chicago, IL

401 North Michigan Avenue
Chicago, IL

One Skokie Valley Road
Highland Park, IL

1415 West 22nd Street
Oak Brook, IL

15105 S LaGrange Road
Orland Park, IL

1572 East Golf Road
Schaumburg, IL

Indiana

4729 East 82nd Street
Indianapolis, IN

8480 Keystone Crossing
Indianapolis, IN

Kansas

5400 College Boulevard
Overland Park, KS

Maine

Three Canal Plaza
Portland, ME

Maryland

7315 Wisconsin Avenue
Bethesda, MD

610 York Road
Towson, MD

Massachusetts

801 Boylston Street
Boston, MA

155 Congress Street
Boston, MA

300 Granite Street
Braintree, MA

44 Mall Road
Burlington, MA

238 Main Street
Cambridge, MA

200 Endicott Street
Danvers, MA

Fidelity Brokerage Services, Inc., 100 Summer St., Boston, MA 02110 Member NYSE/SIPC

Annual Report

405 Cochituate Road
Framingham, MA

551 Boston Turnpike
Shrewsbury, MA

Michigan

500 E. Eisenhower Pkwy.
Ann Arbor, MI

280 Old N. Woodward Ave.
Birmingham, MI

30200 Northwestern Hwy.
Farmington Hills, MI

43420 Grand River Avenue
Novi, MI

Minnesota

7740 France Avenue South
Edina, MN

8342 3rd Street North
Oakdale, MN

Missouri

1524 South Lindbergh Blvd.
St. Louis, MO

Nevada

2225 Village Walk Drive
Henderson, NV

New Jersey

501 Route 73 South
Marlton, NJ

150 Essex Street
Millburn, NJ

35 Morris Street
Morristown, NJ

396 Route 17, North
Paramus, NJ

3518 Route 1 North
Princeton, NJ

530 Broad Street
Shrewsbury, NJ

New Mexico

2261 Q Street NE
Albuquerque, NM

New York

1130 Franklin Avenue
Garden City, NY

37 West Jericho Turnpike
Huntington Station, NY

1271 Avenue of the Americas
New York, NY

980 Madison Avenue
New York, NY

61 Broadway
New York, NY

350 Park Avenue
New York, NY

200 Fifth Avenue
New York, NY

733 Third Avenue
New York, NY

11 Penn Plaza
New York, NY

2070 Broadway
New York, NY

1075 Northern Blvd.
Roslyn, NY

799 Central Park Avenue
Scarsdale, NY

North Carolina

4611 Sharon Road
Charlotte, NC

7011 Fayetteville Road
Durham, NC

Ohio

3805 Edwards Road
Cincinnati, OH

1324 Polaris Parkway
Columbus, OH

1800 Crocker Road
Westlake, OH

28699 Chagrin Boulevard
Woodmere Village, OH

Oregon

7493 SW Bridgeport Road
Tigard, OR

Pennsylvania

600 West DeKalb Pike
King of Prussia, PA

1735 Market Street
Philadelphia, PA

12001 Perry Highway
Wexford, PA

Rhode Island

10 Memorial Boulevard
Providence, RI

Tennessee

3018 Peoples Street
Johnson City, TN

7628 West Farmington Blvd.
Germantown, TN

2035 Mallory Lane
Franklin, TN

Texas

10000 Research Boulevard
Austin, TX

4001 Northwest Parkway
Dallas, TX

12532 Memorial Drive
Houston, TX

2701 Drexel Drive
Houston, TX

6560 Fannin Street
Houston, TX

1701 Lake Robbins Drive
The Woodlands, TX

6500 N. MacArthur Blvd.
Irving, TX

6005 West Park Boulevard
Plano, TX

14100 San Pedro
San Antonio, TX

1576 East Southlake Blvd.
Southlake, TX

Utah

279 West South Temple
Salt Lake City, UT

Virginia

1861 International Drive
McLean, VA

Washington

10500 NE 8th Street
Bellevue, WA

1518 6th Avenue
Seattle, WA

Washington, DC

1900 K Street, N.W.
Washington, DC

Wisconsin

16020 West Bluemound Road
Brookfield, WI

Annual Report

Investment Adviser

Fidelity Management & Research Company

Boston, MA

Investment Sub-Advisers

FMR Co., Inc.

Fidelity Research & Analysis Company

Fidelity Management & Research (U.K.) Inc.

Fidelity Investments Japan Limited

Fidelity International Investment Advisors

Fidelity International Investment Advisors (U.K.) Limited

General Distributor

Fidelity Distributors Corporation

Boston, MA

Transfer and Service Agents

Fidelity Investments Institutional Operations Company, Inc.

Boston, MA

Fidelity Service Company, Inc.

Boston, MA

Custodian

Brown Brothers Harriman & Co.

Boston, MA

The Fidelity Telephone Connection

Mutual Fund 24-Hour Service

Exchanges/Redemptions
and Account Assistance 1-800-544-6666

Product Information 1-800-544-6666

Retirement Accounts 1-800-544-4774
(8 a.m. - 9 p.m.)

TDD Service 1-800-544-0118
for the deaf and hearing impaired
(9 a.m. - 9 p.m. Eastern time)

Fidelity Automated Service
Telephone (FAST®) (automated graphic)    1-800-544-5555

(automated graphic)    Automated line for quickest service

(Fidelity Investment logo)(registered trademark)
Corporate Headquarters
82 Devonshire St., Boston, MA 02109
www.fidelity.com

UIF-UANN-0308
1.789258.105

Item 2. Code of Ethics

As of the end of the period, January 31, 2008, Fidelity Devonshire Trust (the trust) has adopted a code of ethics, as defined in Item 2 of Form N-CSR, that applies to its President and Treasurer and its Chief Financial Officer. A copy of the code of ethics is filed as an exhibit to this Form N-CSR.

Item 3. Audit Committee Financial Expert

The Board of Trustees of the trust has determined that Marie L. Knowles is an audit committee financial expert, as defined in Item 3 of Form N-CSR. Ms. Knowles is independent for purposes of Item 3 of Form N-CSR.

Item 4. Principal Accountant Fees and Services

(a) Audit Fees.

For the fiscal years ended January 31, 2008 and January 31, 2007, the aggregate Audit Fees billed by PricewaterhouseCoopers LLP (PwC) for professional services rendered for the audits of the financial statements, or services that are normally provided in connection with statutory and regulatory filings or engagements for those fiscal years, for the Fidelity Equity-Income Fund, Fidelity Large Cap Growth Fund, Fidelity Large Cap Value Fund, Fidelity Mid Cap Growth Fund, Fidelity Mid Cap Value Fund and Fidelity Utilities Fund (the funds) and for all funds in the Fidelity Group of Funds are shown in the table below.

Fund

2008A

2007A

Fidelity Equity-Income Fund

$221,000

$211,000

Fidelity Large Cap Growth Fund

$49,000

$48,000

Fidelity Large Cap Value Fund

$52,000

$50,000

Fidelity Mid Cap Growth Fund

$50,000

$49,000

Fidelity Mid Cap Value Fund

$51,000

$50,000

Fidelity Utilities Fund

$54,000

$56,000

All funds in the Fidelity Group of Funds audited by PwC

$14,300,000

$13,800,000

A

Aggregate amounts may reflect rounding.

(b) Audit-Related Fees.

In each of the fiscal years ended January 31, 2008 and January 31, 2007 the aggregate Audit-Related Fees billed by PwC for services rendered for assurance and related services to each fund that are reasonably related to the performance of the audit or review of the fund's financial statements, but not reported as Audit Fees, are shown in the table below.

Fund

2008A

2007A

Fidelity Equity-Income Fund

$0

$0

Fidelity Large Cap Growth Fund

$0

$0

Fidelity Large Cap Value Fund

$0

$0

Fidelity Mid Cap Growth Fund

$0

$0

Fidelity Mid Cap Value Fund

$0

$0

Fidelity Utilities Fund

$0

$0

A

Aggregate amounts may reflect rounding.

In each of the fiscal years ended January 31, 2008 and January 31, 2007, the aggregate Audit-Related Fees that were billed by PwC that were required to be approved by the Audit Committee for services rendered on behalf of Fidelity Management & Research Company (FMR) and entities controlling, controlled by, or under common control with FMR (not including any sub-adviser whose role is primarily portfolio management and is subcontracted with or overseen by another investment adviser) that provide ongoing services to the funds ("Fund Service Providers") for assurance and related services that relate directly to the operations and financial reporting of each fund that are reasonably related to the performance of the audit or review of the fund's financial statements, but not reported as Audit Fees, are shown in the table below.

Billed By

2008A

2007A

PwC

$0

$0

A

Aggregate amounts may reflect rounding.

Fees included in the audit-related category comprise assurance and related services (e.g., due diligence services) that are traditionally performed by the independent registered public accounting firm. These audit-related services include due diligence related to mergers and acquisitions, accounting consultations and audits in connection with acquisitions, internal control reviews, attest services that are not required by statute or regulation and consultation concerning financial accounting and reporting standards.

(c) Tax Fees.

In each of the fiscal years ended January 31, 2008 and January 31, 2007, the aggregate Tax Fees billed by PwC for professional services rendered for tax compliance, tax advice, and tax planning for each fund is shown in the table below.

Fund

2008A

2007A

Fidelity Equity-Income Fund

$3,900

$4,800

Fidelity Large Cap Growth Fund

$3,200

$2,900

Fidelity Large Cap Value Fund

$3,200

$2,900

Fidelity Mid Cap Growth Fund

$3,200

$2,900

Fidelity Mid Cap Value Fund

$3,200

$2,900

Fidelity Utilities Fund

$3,200

$3,800

A

Aggregate amounts may reflect rounding.

In each of the fiscal years ended January 31, 2008 and January 31, 2007, the aggregate Tax Fees billed by PwC that were required to be approved by the Audit Committee for professional services rendered on behalf of the Fund Service Providers for tax compliance, tax advice, and tax planning that relate directly to the operations and financial reporting of each fund is shown in the table below.

Billed By

2008A

2007A

PwC

$0

$0

A

Aggregate amounts may reflect rounding.

Fees included in the Tax Fees category comprise all services performed by professional staff in the independent registered public accounting firm's tax division except those services related to the audit. Typically, this category would include fees for tax compliance, tax planning, and tax advice. Tax compliance, tax advice, and tax planning services include preparation of original and amended tax returns, claims for refund and tax payment-planning services, assistance with tax audits and appeals, tax advice related to mergers and acquisitions and requests for rulings or technical advice from taxing authorities.

(d) All Other Fees.

In each of the fiscal years ended January 31, 2008 and January 31, 2007, the aggregate Other Fees billed by PwC for all other non-audit services rendered to the funds is shown in the table below.

Fund

2008A

2007A

Fidelity Equity-Income Fund

$18,800

$19,400

Fidelity Large Cap Growth Fund

$1,100

$1,200

Fidelity Large Cap Value Fund

$1,900

$1,700

Fidelity Mid Cap Growth Fund

$1,300

$1,400

Fidelity Mid Cap Value Fund

$1,500

$1,400

Fidelity Utilities Fund

$2,000

$1,900

A

Aggregate amounts may reflect rounding.

In each of the fiscal years ended January 31, 2008 and January 31, 2007, the aggregate Other Fees billed by PwC that were required to be approved by the Audit Committee for all other non-audit services rendered on behalf of the Fund Service Providers that relate directly to the operations and financial reporting of each fund is shown in the table below.

Billed By

2008A

2007A

PwC

$220,000

$125,000

A

Aggregate amounts may reflect rounding.

Fees included in the All Other Fees category include services related to internal control reviews, strategy and other consulting, financial information systems design and implementation, consulting on other information systems, and other tax services unrelated to the fund.

(e) (1)

Audit Committee Pre-Approval Policies and Procedures:

The trust's Audit Committee must pre-approve all audit and non-audit services provided by the independent registered public accounting firm relating to the operations or financial reporting of the funds. Prior to the commencement of any audit or non-audit services to a fund, the Audit Committee reviews the services to determine whether they are appropriate and permissible under applicable law.

The trust's Audit Committee has adopted policies and procedures to, among other purposes, provide a framework for the Committee's consideration of non-audit services by the audit firms that audit the Fidelity funds. The policies and procedures require that any non-audit service provided by a fund audit firm to a Fidelity Fund and any non-audit service provided by a fund auditor to a Fund Service Provider that relates directly to the operations and financial reporting of a Fidelity fund (Covered Service) are subject to approval by the Audit Committee before such service is provided. Non-audit services provided by a fund audit firm for a Fund Service Provider that do not relate directly to the operations and financial reporting of a Fidelity fund (Non-Covered Service) but that are expected to exceed $50,000 are also subject to pre-approval by the Audit Committee.

All Covered Services, as well as Non-Covered Services that are expected to exceed $50,000, must be approved in advance of provision of the service either: (i) by formal resolution of the Audit Committee, or (ii) by oral or written approval of the service by the Chair of the Audit Committee (or if the Chair is unavailable, such other member of the Audit Committee as may be designated by the Chair to act in the Chair's absence). The approval contemplated by (ii) above is permitted where the Treasurer determines that action on such an engagement is necessary before the next meeting of the Audit Committee. Neither pre-approval nor advance notice of Non-Covered Service engagements for which fees are not expected to exceed $50,000 is required; such engagements are to be reported to the Audit Committee monthly.

(e) (2)

Services approved pursuant to paragraph (c)(7)(i)(C) of Rule 2-01 of Regulation S-X:

Audit-Related Fees:

There were no amounts that were approved by the Audit Committee pursuant to the de minimis exception for the fiscal years ended January 31, 2008 and January 31, 2007 on behalf of each fund.

There were no amounts that were required to be approved by the Audit Committee pursuant to the de minimis exception for the fiscal years ended January 31, 2008 and January 31, 2007 on behalf of the Fund Service Providers that relate directly to the operations and financial reporting of each fund.

Tax Fees:

There were no amounts that were approved by the Audit Committee pursuant to the de minimis exception for the fiscal years ended January 31, 2008 and January 31, 2007 on behalf of each fund.

There were no amounts that were required to be approved by the Audit Committee pursuant to the de minimis exception for the fiscal years ended January 31, 2008 and January 31, 2007 on behalf of the Fund Service Providers that relate directly to the operations and financial reporting of each fund.

All Other Fees:

There were no amounts that were approved by the Audit Committee pursuant to the de minimis exception for the fiscal years ended January 31, 2008 and January 31, 2007 on behalf of each fund.

There were no amounts that were required to be approved by the Audit Committee pursuant to the de minimis exception for the fiscal years ended January 31, 2008 and January 31, 2007 on behalf of the Fund Service Providers that relate directly to the operations and financial reporting of each fund.

(f) Not applicable.

(g) For the fiscal years ended January 31, 2008 and January 31, 2007, the aggregate fees billed by PwC of $1,560,000A and $1,300,000A for non-audit services rendered on behalf of the funds, FMR (not including any sub-adviser whose role is primarily portfolio management and is subcontracted with or overseen by another investment adviser) and Fund Service Providers relating to Covered Services and Non-Covered Services are shown in the table below.

2008A

2007A

Covered Services

$265,000

$170,000

Non-Covered Services

$1,295,000

$1,130,000

A

Aggregate amounts may reflect rounding.

(h) The trust's Audit Committee has considered Non-Covered Services that were not pre-approved that were provided by PwC to Fund Service Providers to be compatible with maintaining the independence of PwC in its audit of the funds, taking into account representations from PwC, in accordance with Independence Standards Board Standard No.1, regarding its independence from the funds and their related entities.

Item 5. Audit Committee of Listed Registrants

Not applicable.

Item 6. Schedule of Investments

Not applicable.

Item 7. Disclosure of Proxy Voting Policies and Procedures for Closed-End Management Investment Companies

Not applicable.

Item 8. Portfolio Managers of Closed-End Management Investment Companies

Not applicable.

Item 9. Purchase of Equity Securities by Closed-End Management Investment Company and Affiliated Purchasers

Not applicable.

Item 10. Submission of Matters to a Vote of Security Holders

There were no material changes to the procedures by which shareholders may recommend nominees to the trust's Board of Trustees.

Item 11. Controls and Procedures

(a)(i) The President and Treasurer and the Chief Financial Officer have concluded that the trust's disclosure controls and procedures (as defined in Rule 30a-3(c) under the Investment Company Act) provide reasonable assurances that material information relating to the trust is made known to them by the appropriate persons, based on their evaluation of these controls and procedures as of a date within 90 days of the filing date of this report.

(a)(ii) There was no change in the trust's internal control over financial reporting (as defined in Rule 30a-3(d) under the Investment Company Act) that occurred during the second fiscal quarter of the period covered by this report that has materially affected, or is reasonably likely to materially affect, the trust's internal control over financial reporting.

Item 12. Exhibits

(a)

(1)

Code of Ethics pursuant to Item 2 of Form N-CSR is filed and attached hereto as EX-99.CODE ETH.

(a)

(2)

Certification pursuant to Rule 30a-2(a) under the Investment Company Act of 1940 (17 CFR 270.30a-2(a)) is filed and attached hereto as Exhibit 99.CERT.

(a)

(3)

Not applicable.

(b)

Certification pursuant to Rule 30a-2(b) under the Investment Company Act of 1940 (17 CFR 270.30a-2(b)) is furnished and attached hereto as Exhibit 99.906CERT.

SIGNATURES

Pursuant to the requirements of the Securities Exchange Act of 1934 and the Investment Company Act of 1940, the registrant has duly caused this report to be signed on its behalf by the undersigned, thereunto duly authorized.

Fidelity Devonshire Trust

By:

/s/Kimberley Monasterio

Kimberley Monasterio

President and Treasurer

Date:

March 28, 2008

Pursuant to the requirements of the Securities Exchange Act of 1934 and the Investment Company Act of 1940, this report has been signed below by the following persons on behalf of the registrant and in the capacities and on the dates indicated.

By:

/s/Kimberley Monasterio

Kimberley Monasterio

President and Treasurer

Date:

March 28, 2008

By:

/s/Joseph B. Hollis

Joseph B. Hollis

Chief Financial Officer

Date:

March 28, 2008