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Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($)
$ in Thousands
3 Months Ended
Mar. 31, 2019
Mar. 31, 2018
Income Statement [Abstract]    
Collaboration revenue $ 4,205 $ 641
Operating expenses:    
Research and development 37,403 20,819
General and administrative 9,310 5,570
Total operating expenses 46,713 26,389
Loss from operations (42,508) (25,748)
Interest and other income, net 3,516 2,070
Net loss (38,992) (23,678)
Other comprehensive income (loss):    
Net unrealized income (loss) on marketable securities, net of tax 981 (919)
Comprehensive loss $ (38,011) $ (24,597)
Net loss per share, basic and diluted (usd per share) $ (0.41) $ (0.26)
Weighted average number of shares outstanding, basic and diluted (in shares) 94,984,503 89,560,576