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STOCK-BASED COMPENSATION (Tables)
 12 Months Ended
Dec. 31, 2023
Share-Based Payment Arrangement [Abstract]  
Summary of Fair Values of Stock Options Granted and Assumptions used in Black-Scholes Model
The estimated fair values of stock options granted, and the assumptions used for the Black-Scholes option pricing model were as follows:
 Year Ended December 31,
 20232022
Estimated fair values of stock options granted$— $42.83 
Assumptions:  
Risk-free interest rate
—%
1.9% - 3.5%
Expected volatility
—%
89.7% - 92.8%
Expected term (in years)
—
5.50 - 6.89
Summary of Stock Option Activity
Information on stock option activity is as follows:
 
Options
Exercisable
Weighted Average
Exercise
Price Per
Share
Options
Outstanding
Weighted Average
Exercise
Price Per
Share
Balance as of December 31, 202267,978$496.83 116,860$367.58 
Granted—— —— 
Vested42,333156.25 —— 
Exercised—— —— 
Forfeited(1,530)441.63 (7,339)371.04 
Balance as of December 31, 2023108,781$365.07 109,521$367.35 
Summary of Stock-based Compensation Expense
Stock-based compensation expense related to stock option awards is as follows:
 Year Ended December 31,
In Thousands20232022
Stock-based compensation expense$1,847 $2,031 


Stock-based compensation expense related to RSAs is as follows:
 Year Ended December 31,
In Thousands20232022
Stock-based compensation expense$11,497 $— 


Stock-based compensation expense related to RSUs is as follows:
 Year Ended December 31,
In Thousands20232022
Stock-based compensation expense$3,018 $1,318 


Stock-based compensation expense related to PSUs is as follows:
 Year Ended December 31,
In Thousands20232022
Stock-based compensation (benefit) expense$(270)$649 
Certain statement of operations amounts were as follows:
 Year Ended December 31,
In Thousands20232022
Stock-based compensation expense:
Research and development$7,554 $946 
Selling, general, and administrative8,538 3,052 
Total$16,092 $3,998 
Schedule of Nonvested Share Activity Information on Class A restricted stock award activity is as follows:
 
Restricted Stock
Awards
Weighted Average Grant
Date Fair Value
Unvested balance as of December 31, 2022—$— 
Granted1,019,28231.50 
Vested(349,907)31.50 
Forfeited(29,315)31.50 
Unvested balance as of December 31, 2023640,060$31.50 
Information on restricted stock unit activity is as follows:
 
Restricted Stock
Units
Weighted Average Grant
Date Fair Value
Unvested balance as of December 31, 202224,575$99.36 
Granted272,88519.02 
Vested(96,734)38.29 
Forfeited(53,504)23.29 
Unvested balance as of December 31, 2023147,222$18.21 
Summary of Grant Date Fair Value of Awards Vested
The total fair value of RSAs that vested is as follows:
 Year Ended December 31,
In Thousands20232022
Fair value of shares vested$5,361 $— 
The total fair value of RSUs that vested is as follows:
 Year Ended December 31,
In Thousands20232022
Fair value of shares vested$2,126 $175 
Summary of Stock-Based Compensation Granted As Deemed Dividends The Company recorded deemed dividends as follows:
 Year Ended December 31,
In Thousands20232022
Deemed dividends from grants to Cellectis employees$— $90 
Share-based Payment Arrangement, Performance Shares, Outstanding Activity
Information on performance stock unit activity is as follows:
 Performance Stock Units
Weighted Average Grant
Date Fair Value
Outstanding as of December 31, 202222,600$81.73 
Issued24,80025.65 
Forfeited/canceled(9,333)95.80 
Vested(38,067)31.82 
Outstanding as of December 31, 2023—$— 
Share-Based Compensation Arrangements by Share-Based Payment Award, Performance-Based Units, Vested and Expected to Vest The total fair value of PSUs that vested is as follows:

Year Ended December 31,
In Thousands20232022
Fair value of shares vested$1,005 $—