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Condensed Consolidated Statements of Operations - USD ($)
$ in Thousands
3 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Condensed Consolidated Statements of Operations    
Collaboration and other research and development revenues $ 6,771 $ 6,499
Revenue type us-gaap:ServiceMember us-gaap:ServiceMember
Operating expenses:    
Research and development $ 37,976 $ 41,937
General and administrative 19,545 21,445
Total operating expenses 57,521 63,382
Operating loss (50,750) (56,883)
Other income, net:    
Other (expense) income, net (234) 21
Interest income, net 469 134
Total other income, net 235 155
Net loss $ (50,515) $ (56,728)
Net loss per share, basic (in dollars per share) $ (0.74) $ (0.86)
Net loss per share, diluted (in dollars per share) $ (0.74) $ (0.86)
Weighted-average common shares outstanding, basic (in shares) 68,484,978 65,992,395
Weighted-average common shares outstanding, diluted (in shares) 68,484,978 65,992,395