Exhibit 107.1

CALCULATION OF FILING FEE TABLE

424(b)(5)

(Form Type)

4D Molecular Therapeutics, Inc.

(Exact Name of Registrant as Specified in its Charter)

Table 1: Newly Registered Securities

	Security  Type	Security Class Title	Fee Calculation Rule	Amount Registered	Proposed Maximum  Offering Price Per  Unit	Maximum Aggregate Offering Price	Fee Rate	Amount of Registration Fee
Newly Registered Securities
Fees to Be Paid	Equity	Common  Stock, par value  $0.0001 per share	Rule 457(o) (1)	—	—	$250,000,000	0.00014760	$36,900.00
Fee Previously Paid	—	—	—	—	—	—	—	—
Carry Forward Securities
N/A
	Total Offering Amounts					$250,000,000		$36,900.00
	Total Fees Previously Paid							—
	Total Fee Offsets							—
	Net Fee Due							$36,900.00

⁽¹⁾ The registration fee is calculated in accordance with Rule 457(o) under the Securities Act of 1933, as amended (the “Securities Act”), based on the proposed maximum aggregate offering price, and Rule 457(r) under the Securities Act. In accordance with Rules 456(b) and 457(r) under the Securities Act, the registrant initially deferred payment of all of the registration fee for Registration Statement on Form S-3 (File No. 333-276872), except with respect to unsold securities that have been previously registered.