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Research and Collaboration Arrangements - Summary of Collaboration and License Revenue (Details) - USD ($)
$ in Thousands
3 Months Ended 9 Months Ended
Sep. 30, 2021
Sep. 30, 2020
Sep. 30, 2021
Sep. 30, 2020
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]        
Total revenue $ 1,366 $ 7,421 $ 17,946 $ 14,589
Collaboration and License Revenue | uniQure        
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]        
Total revenue 64 255 724 504
Collaboration and License Revenue | Roche        
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]        
Total revenue 1,290 7,215 17,190 14,174
Collaboration and License Revenue | CFF        
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]        
Total revenue $ 12 $ (49) $ 32 $ (89)