Collaboration, License and Research Agreements (Tables)

12 Months Ended

Dec. 31, 2021

Collaboration License And Research Agreements [Abstract]

Summary of Reconciliation of Deferred Revenue Related to Lilly Agreement

A reconciliation of the closing balance of deferred revenue related to the Lilly Agreement is as follows (in thousands):

Balance at December 31, 2019	$	18,940
Revenue recognized		(3,100	)
Balance at December 31, 2020		15,840
Revenue recognized		(4,732	)
Balance at December 31, 2021	$	11,108