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Stock-Based Compensation and Warrants
3 Months Ended
Mar. 31, 2022
Share-based Payment Arrangement [Abstract]  
Stock-Based Compensation and Warrants Stock-Based Compensation and Warrants
The Company currently grants stock options to employees and directors under the 2020 Stock Incentive Plan (the "2020 Plan") and formerly, the Company granted stock options under the 2016 Long Term Incentive Plan (the "2016 Plan"). The 2020 Plan is administered by the Compensation Committee of the Company's Board of Directors. The 2020 Plan is intended to attract and retain employees and directors and provide an incentive for these individuals to assist the Company to achieve long-range performance goals and to enable these individuals to participate in the long-term growth of the Company.

The following table presents the stock option activity for both the 2016 Plan and the 2020 Plan as of March 31, 2022:
Three Months Ended
March 31, 2022
Weighted average exercise price
Outstanding at December 31, 2021
13,797,556 $5.55 
   Granted 336,381$2.82 
   Forfeited (979,873)$3.95 
   Exercised(11,995)$1.86 
Outstanding at March 31, 2022
13,142,069 $5.61 
Exercisable at March 31, 2022
2,135,201 $4.19 

The total intrinsic value of all outstanding and exercisable stock options at March 31, 2022 was $424 and $312, respectively.

The total stock-based compensation expense included in the Company's condensed consolidated statements of comprehensive loss was as follows:

Three Months Ended
March 31,
20222021
Research and development$1,874 $323 
General and administrative2,122 546 
   Total stock-based compensation expense$3,996 $869 

Warrants

The Company had outstanding and exercisable warrants of 5,821,137 with a weighted average exercise price of $1.56 as of March 31, 2022 and December 31, 2021.