Note 7 - Accrued Expenses - Accrued Expenses and Other Current Liabilities (Details) - USD ($) $ in Thousands	Sep. 30, 2022	Dec. 31, 2021
Research and development fees	$ 5,036	$ 5,682
Compensation expense	3,248	3,581
Accrued license obligations	131	1,541
Accrued accounts payable	1,560	2,980
Lease liabilities	826	1,049
Other current liabilities	581	393
Collaboration cost-sharing obligation	0	1,610
Total	$ 11,382	$ 16,836