Exhibit 99.1

JOINT FILING AGREEMENT

The undersigned acknowledge and agree that the foregoing statement on Schedule 13D is filed on behalf of each of the undersigned and that all subsequent amendments to this foregoing shall be filed on behalf of each of the undersigned without the necessity of filing additional joint filing agreements.  The undersigned acknowledge that each shall be responsible for the timely filing of such amendments, and for the completeness and accuracy of the information concerning it contained therein, but shall not be responsible for the completeness and accuracy of the information concerning the others, except to the extent that it knows or has reason to believe that such information is inaccurate.

This Agreement may be executed in counterparts and each of such counterparts taken together shall constitute one and the same instrument.

Dated: April 12, 2018

Bpifrance Participations S.A.
By:	/s/ Sophie Paquin
Name:	Sophie Paquin
Title:	Director of Legal Affairs
Innobio FPCI
By:	/s/ Sophie Paquin
Name:	Sophie Paquin
Title:	Director of Legal Affairs (Bpifrance Investissement S.A.S.)
Bpifrance Investissement S.A.S.
By:	/s/ Sophie Paquin
Name:	Sophie Paquin
Title:	Director of Legal Affairs
Caisse des Dépôts
By:	/s/ Eric Flamarion
Name:	Eric Flamarion
Title:	Head of Financial Investment Division
Epic Bpifrance
By:	/s/ Sophie Paquin
Name:	Sophie Paquin
Title:	Director of Legal Affairs
Bpifrance S.A.
By:	/s/ Boubakar Dione
Name:	Boubakar Dione
Title:	Group Director of Legal Affairs