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Condensed Consolidated Statements of Operations - USD ($)
shares in Thousands, $ in Thousands
3 Months Ended
Mar. 31, 2019
Mar. 31, 2018
Income Statement [Abstract]    
Collaboration revenue $ 186 $ 227
Total revenue 186 227
Operating expenses:    
Research and development 13,296 34,706
General and administrative 9,905 14,229
Restructuring charges (credits) (61) 0
Total operating expenses 23,140 48,935
Loss from operations (22,954) (48,708)
Other income (expense):    
Interest income, net 2,304 200
Other expense, net (17) (272)
Total other income (expense), net 2,287 (72)
Loss before income taxes (20,667) (48,780)
Provision for (benefit from) income taxes 198 (37)
Net loss $ (20,865) $ (48,743)
Basic and diluted net loss per share (in dollars per share) $ (0.52) $ (1.26)
Shares used to compute basic and diluted net loss per share (in shares) 39,864 38,684