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Roche License Agreement (Details) - Roche [Member] - USD ($)
$ in Millions
1 Months Ended 3 Months Ended 6 Months Ended
May 31, 2014
Feb. 28, 2014
Jun. 30, 2016
Jun. 30, 2015
Jun. 30, 2016
Jun. 30, 2015
Collaborative Arrangement [Member]            
License Agreement [Line Items]            
Cost sharing payments recognized as research and development expense     $ 0.9 $ 0.8 $ 1.4 $ 1.3
Multiple element consideration, relative sales price method, amt allocable to license   $ 35.6        
Development reimbursement     0.7 1.5 1.9 3.0
Milestone payment received, clinical milestone $ 15.0          
Research Reimbursement [Member]            
License Agreement [Line Items]            
Collaboration service revenue, research services     0.3 0.3 0.6 0.6
Development Costs Reimbursement [Member]            
License Agreement [Line Items]            
Development reimbursement recognized as collaboration license revenue     0.0 0.0 0.0 0.2
Development reimbursement recognized as offset to research and development expense     $ 0.7 $ 1.5 $ 1.9 $ 2.7
License [Member]            
License Agreement [Line Items]            
Upfront payment pursuant to license agreement   $ 30.0