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Net Loss per Share - Computation of Basic and Diluted Net Loss per Share (Details) - USD ($)
$ / shares in Units, $ in Thousands
3 Months Ended 12 Months Ended
Dec. 31, 2017
Sep. 30, 2017
Jun. 30, 2017
Mar. 31, 2017
Dec. 31, 2016
Sep. 30, 2016
Jun. 30, 2016
Mar. 31, 2016
Dec. 31, 2017
Dec. 31, 2016
Dec. 31, 2015
Numerator:                      
Net loss $ (81,731) $ (79,227) $ (72,891) $ (68,290) $ (71,287) $ (64,907) $ (56,923) $ (52,757) $ (302,139) $ (245,874) $ (145,618)
Denominator:                      
Weighted-average shares used to compute net loss per share, basic and diluted                 42,453,135 39,586,908 36,782,603
Net loss per share, basic and diluted $ (1.89) $ (1.87) $ (1.72) $ (1.63) $ (1.75) $ (1.64) $ (1.46) $ (1.35) $ (7.12) $ (6.21) $ (3.96)