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Common Stock and Stock-Based Compensation (Tables)
9 Months Ended
Sep. 30, 2017
Employees and Nonemployees Stock Option Restricted Stock Units  
Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Line Items]  
Schedule of Stock-Based Compensation Expense

The stock-based compensation expense recorded related to options and restricted stock units granted to employees and nonemployees were as follows (in thousands):

 

 

 

Three Months Ended

 

 

Nine Months Ended

 

 

 

September 30,

 

 

September 30,

 

 

 

2017

 

 

2016

 

 

2017

 

 

2016

 

Research and development

 

$

3,659

 

 

$

4,399

 

 

$

11,163

 

 

$

9,842

 

General and administrative

 

 

4,524

 

 

 

3,871

 

 

 

13,902

 

 

 

9,894

 

 

 

$

8,183

 

 

$

8,270

 

 

$

25,065

 

 

$

19,736