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Common Stock and Stock-Based Compensation - Schedule of Stock-Based Compensation Expense Related to Founder's Shares (Details) - USD ($)
$ in Thousands
3 Months Ended 6 Months Ended
Jun. 30, 2015
Jun. 30, 2014
Jun. 30, 2015
Jun. 30, 2014
Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Line Items]        
Stock-based compensation expense $ 4,618 $ 1,076 $ 6,794 $ 1,567
Research and Development Expense        
Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Line Items]        
Stock-based compensation expense 2,280 468 3,456 690
General and Administrative Expense        
Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Line Items]        
Stock-based compensation expense 2,338 608 3,338 877
Restricted Common Stock (Founders Shares)        
Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Line Items]        
Stock-based compensation expense 3 134 9 238
Restricted Common Stock (Founders Shares) | Research and Development Expense        
Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Line Items]        
Stock-based compensation expense $ 3 133 $ 9 236
Restricted Common Stock (Founders Shares) | General and Administrative Expense        
Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Line Items]        
Stock-based compensation expense   $ 1   $ 2