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Equity Incentive Awards
 3 Months Ended
Mar. 31, 2021
Share-based Payment Arrangement [Abstract]  
Equity Incentive Awards Equity Incentive Awards
Stock Options
The following table summarizes the Company’s option activity and related information:
Number of
Options
(in thousands)		Weighted-
Average
Exercise Price
Balance at December 31, 202010,432 $11.65 
Options granted3,301 12.92 
Options exercised(9)5.86 
Options forfeited(375)9.00 
Balance at March 31, 202113,349 $12.04 
Exercisable as of March 31, 20214,559 $8.41 
Restricted Stock Units (“RSUs”)
The following table summarizes the Company’s RSUs activity and related information:
Number of Units
(in thousands)		Weighted-
Average Grant-
Date Fair Value
Outstanding at December 31, 2020540 $6.41 
Granted382 12.25 
Vested and released(249)5.20 
Forfeited(1)14.10 
Outstanding at March 31, 2021672 $10.16 
Stock-Based Compensation Expense
The following table presents, by operating expense, the Company’s stock-based compensation expense:
Three Months Ended
March 31,
20212020
(In thousands)
Research and development$2,339 $1,248 
General and administrative4,885 2,161 
Total stock-based compensation expense$7,224 $3,409 
During the three months ended March 31, 2021, the Company recorded approximately $0.2 million of stock-based compensation expense as a result of the modification of the vesting and exercisability of stock awards associated with the departure of two of its executives.