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Organization and Operations - Stock Purchase Agreement and Series X Preferred Stock (Details)
$ in Thousands
1 Months Ended 3 Months Ended 6 Months Ended
Jan. 28, 2021
USD ($)
shares
Feb. 28, 2021
shares
Jun. 30, 2021
shares
Mar. 31, 2021
USD ($)
shares
Jun. 30, 2022
shares
Jun. 30, 2021
USD ($)
Dec. 31, 2021
shares
Jun. 02, 2021
Stock Purchase Agreement and Series X Preferred Stock                
Net proceeds from issuance of private placement | $           $ 104,261    
Beneficial conversion feature | $       $ 19,565        
Common Stock Issuable Upon Conversion at Transaction Date   14,813,954            
Minimum                
Stock Purchase Agreement and Series X Preferred Stock                
Common stock, percentage of beneficial ownership initially, after conversion         4.99%      
Common stock, percentage of beneficial ownership thereafter, after conversion         4.99%      
Maximum                
Stock Purchase Agreement and Series X Preferred Stock                
Common stock, percentage of beneficial ownership initially, after conversion         9.99%      
Common stock, percentage of beneficial ownership thereafter, after conversion         19.99%      
Shares issued in merger                
Stock Purchase Agreement and Series X Preferred Stock                
Common Stock Issuable Upon Conversion at Transaction Date   8,417,502            
Shares issued in February 2021 Financing                
Stock Purchase Agreement and Series X Preferred Stock                
Common Stock Issuable Upon Conversion at Transaction Date   5,928,952            
Warrants assumed in merger                
Stock Purchase Agreement and Series X Preferred Stock                
Common Stock Issuable Upon Conversion at Transaction Date   467,500            
Series X redeemable convertible preferred stock                
Stock Purchase Agreement and Series X Preferred Stock                
Issuance of preferred stock in a private offering of public equity, net of issuance costs (in shares)       35,573        
Number of shares of common stock into which each share of Series X Preferred Stock may be converted         166.67      
Series X Preferred Stock at Transaction Date   88,882            
Common Stock Issuable Upon Conversion at Transaction Date         5,242,501      
Number of preferred shares converted during period         54,622      
Preferred stock, outstanding (in shares)         31,455   31,455  
Series X redeemable convertible preferred stock | Shares issued in merger                
Stock Purchase Agreement and Series X Preferred Stock                
Series X Preferred Stock at Transaction Date   50,504            
Series X redeemable convertible preferred stock | Shares issued in February 2021 Financing                
Stock Purchase Agreement and Series X Preferred Stock                
Series X Preferred Stock at Transaction Date   35,573            
Series X redeemable convertible preferred stock | Warrants assumed in merger                
Stock Purchase Agreement and Series X Preferred Stock                
Series X Preferred Stock at Transaction Date   2,805            
Common stock                
Stock Purchase Agreement and Series X Preferred Stock                
Number of shares of common stock into which each share of Series X Preferred Stock may be converted         166.67     166.67
Common Stock Issuable Upon Conversion at Transaction Date         5,242,501      
Number of common shares issued from conversion of Series X preferred shares     8,921,966   9,103,664      
Quellis Biosciences, Inc | Stock Purchase Agreement                
Stock Purchase Agreement and Series X Preferred Stock                
Beneficial conversion feature | $ $ 19,600              
Quellis Biosciences, Inc | Series X redeemable convertible preferred stock                
Stock Purchase Agreement and Series X Preferred Stock                
Number of shares of common stock into which each share of Series X Preferred Stock may be converted 166.67              
Quellis Biosciences, Inc | Series X redeemable convertible preferred stock | Stock Purchase Agreement                
Stock Purchase Agreement and Series X Preferred Stock                
Issuance of preferred stock in a private offering of public equity, net of issuance costs (in shares) 35,573              
Gross proceeds from issuance of preferred stock in a private offering | $ $ 110,000              
Net proceeds from issuance of private placement | $ 104,300              
Issuance costs | $ $ 5,700              
Quellis Biosciences, Inc | Common stock | Series X redeemable convertible preferred stock                
Stock Purchase Agreement and Series X Preferred Stock                
Number of shares of common stock into which each share of Series X Preferred Stock may be converted 166.67