Organization and Operations - Reverse Stock Split, Agreement and Plan of Merger (Details) - USD ($) $ / shares in Units, $ in Millions		9 Months Ended
	Jan. 28, 2021	Sep. 30, 2021	Jun. 02, 2021	Dec. 31, 2020
Agreement and Plan of Merger
Reverse stock split ratio		0.1667
Common stock, par value (in dollars per share)		$ 0.001		$ 0.001
Series X redeemable convertible preferred stock
Agreement and Plan of Merger
Issuance of preferred stock upon acquisition of Quellis (in shares)		50,504
Common stock
Agreement and Plan of Merger
Shares issued on acquisition (in shares)		555,444
Warrants to purchase number of shares			467,500
Common stock | Series X redeemable convertible preferred stock
Agreement and Plan of Merger
Exercise Price (in dollars per share)			$ 2.10
Quellis Biosciences, Inc | Series X redeemable convertible preferred stock
Agreement and Plan of Merger
Issuance of preferred stock upon acquisition of Quellis (in shares)	50,504
Conversion value on closing date	$ 122.7
Warrants to purchase number of shares	2,805
Exercise Price (in dollars per share)	$ 341.70
Quellis Biosciences, Inc | Common stock
Agreement and Plan of Merger
Shares issued on acquisition (in shares)	555,444
Common stock, par value (in dollars per share)	$ 0.001
Number of options assumed in Quellis Acquisition	55,414
Warrants to purchase number of shares	30,856
Exercise Price (in dollars per share)	$ 2.10		$ 2.10