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Selected Quarterly Financial Data
 12 Months Ended
Dec. 31, 2018
Quarterly Financial Information Disclosure [Abstract]  
Selected Quarterly Financial Data (Unaudited) Selected Quarterly Financial Data (Unaudited)
The following table summarizes unaudited quarterly financial data for the years ended December 31, 2018 and 2017 (in thousands, except per share data).
 
 
Three Months Ended (unaudited)
 
 
March 31,
 
June 30,
 
September 30,
 
December 31,
 
 
2018
 
2018
 
2018
 
2018
Grant and collaboration revenue
 
$
—

 
$
—

 
$
—

 
$
903

Operating expenses
 
$
15,813

 
$
18,769

 
$
15,941

 
$
15,402

Net loss
 
$
(15,888
)
 
$
(18,796
)
 
$
(16,001
)
 
$
(14,651
)
Net loss attributable to common stockholders
 
$
(15,888
)
 
$
(18,796
)
 
$
(16,001
)
 
$
(14,651
)
 
 
 
 
 
 
 
 
 
Net loss per share attributable to common stockholders, basic and diluted
 
$
(0.71
)
 
$
(0.84
)
 
$
(0.71
)
 
$
(0.65
)
 
 
 
 
 
 
 
 
 
 
 
Three Months Ended (unaudited)
 
 
March 31,
 
June 30,
 
September 30,
 
December 31,
 
 
2017
 
2017
 
2017
 
2017
Grant and collaboration revenue
 
$
137

 
$
26

 
$
27

 
$
17

Operating expenses
 
$
14,919

 
$
15,897

 
$
13,881

 
$
19,294

Net loss
 
$
(15,134
)
 
$
(15,967
)
 
$
(14,676
)
 
$
(19,544
)
Net loss attributable to common stockholders
 
$
(15,134
)
 
$
(15,967
)
 
$
(14,676
)
 
$
(19,544
)
 
 
 
 
 
 
 
 
 
Net loss per share attributable to common stockholders, basic and diluted
 
$
(0.82
)
 
$
(0.85
)
 
$
(0.66
)
 
$
(0.88
)