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Net Loss Per Common Share (Details) - USD ($)
$ / shares in Units, $ in Thousands
3 Months Ended 12 Months Ended
Dec. 31, 2017
Sep. 30, 2017
Jun. 30, 2017
Mar. 31, 2017
Dec. 31, 2016
Sep. 30, 2016
Jun. 30, 2016
Mar. 31, 2016
Dec. 31, 2017
Dec. 31, 2016
Dec. 31, 2015
Basic and diluted net loss per common share calculation:                      
Net loss $ (14,720) $ (15,999) $ (20,432) $ (20,714) $ (35,982) $ (29,985) $ (19,295) $ (17,732) $ (71,865) $ (102,994) $ (50,528)
Net loss attributable to common stockholders                 $ (71,865) $ (102,994) $ (50,528)
Weighted average common shares outstanding (in shares) 62,290,002 60,113,327 58,381,868 56,894,672 53,850,166 52,205,156 52,174,569 51,350,365 59,436,649 52,398,521 43,794,276
Net loss per share of common stock—basic and diluted (in dollars per share) $ (0.24) $ (0.27) $ (0.35) $ (0.36) $ (0.67) $ (0.57) $ (0.37) $ (0.35) $ (1.21) $ (1.97) $ (1.15)
Outstanding securities excluded from computation of diluted weighted shares outstanding (in shares)                 8,747,314 6,431,428 4,692,873
Options outstanding                      
Basic and diluted net loss per common share calculation:                      
Outstanding securities excluded from computation of diluted weighted shares outstanding (in shares)                 8,624,223 6,370,578 4,630,073
Warrants                      
Basic and diluted net loss per common share calculation:                      
Outstanding securities excluded from computation of diluted weighted shares outstanding (in shares)                 123,091 60,850 62,800