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Stockholders' Equity and Stock-Based Compensation (Tables)
 9 Months Ended
Sep. 30, 2020
Share-based Payment Arrangement [Abstract]  
Summary of stock option activity
The following is a summary of stock option activity for the nine months ended September 30, 2019 (in thousands, except per share data):
Shares
Weighted-
Average
Exercise
Price per Share
Outstanding at December 31, 20194,253 $29.59 
Granted
1,396 28.83 
Exercised
(239)16.64 
Canceled
(230)39.02 
Outstanding at September 30, 20205,180 $29.57 
Schedule of restricted stock unit activity
The following is a summary of restricted stock unit activity for the nine months ended September 30, 2019 (in thousands, except per share data):
Shares
Weighted- Average Fair Value per Share at Grant Date
Outstanding at December 31, 2019439 $36.97 
Granted
218 27.34 
Vested
(220)26.39 
Canceled
(42)38.79 
Outstanding at September 30, 2020395 $37.61 
Stock-based compensation expense
The following table summarizes the components of total stock-based compensation expense included in the condensed consolidated statements of operations (in thousands):
Three Months Ended September 30,Nine Months Ended September 30,
2020201920202019
Research and development$2,865 $2,012 $9,082 $5,521 
Selling, general and administrative4,276 3,423 12,756 9,495 
Compensation expense related to acquired IPR&D— 4,927 — 4,927 
Total$7,141 $10,362 $21,838 $19,943