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Stockholders' Equity and Stock-Based Compensation (Tables)
 6 Months Ended
Jun. 30, 2020
Share-based Payment Arrangement [Abstract]  
Summary of stock option activity
The following is a summary of stock option activity for the six months ended June 30, 2020 (in thousands, except per share data):
Shares
Weighted-
Average
Exercise
Price per Share
Outstanding at December 31, 20194,253  $29.59  
Granted
1,185  29.36  
Exercised
(231) 16.85  
Canceled
(145) 39.56  
Outstanding at June 30, 20205,062  $29.83  
Schedule of restricted stock unit activity
The following is a summary of restricted stock unit activity for the six months ended June 30, 2020 (in thousands, except per share data):
Shares
Weighted- Average Fair Value per Share at Grant Date
Outstanding at December 31, 2019439  $36.97  
Granted
218  27.34  
Vested
(194) 23.95  
Canceled
(28) 38.74  
Outstanding at June 30, 2020435  $38.09  
Stock-based compensation expense
The following table summarizes the components of total stock-based compensation expense included in the condensed consolidated statements of operations (in thousands):
Three Months Ended June 30,Six Months Ended June 30,
2020201920202019
Research and development$3,488  $2,074  $6,217  $3,509  
Selling, general and administrative4,815  3,284  8,480  6,072  
Total$8,303  $5,358  $14,697  $9,581