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Related Party Transactions (Details) - USD ($)
1 Months Ended 3 Months Ended
Oct. 31, 2017
Nov. 30, 2015
Mar. 31, 2018
Mar. 31, 2017
Dec. 31, 2017
Related Party Transaction [Line Items]          
Shares issued as payment for services     $ 2,941,000    
Fair value of financial assets measured at fair value on a recurring basis     180,881,000   $ 183,173,000
Other nonoperating income     (659,000) $ 595,000  
Series A preferred shares          
Related Party Transaction [Line Items]          
Convertible preferred shares, conversion price, number of days used to calculate volume-weighted average market price 20 days        
Affiliate of Third Security, LLC | Series A preferred shares          
Related Party Transaction [Line Items]          
Value of convertible preferred shares authorized for issuance $ 100,000,000        
Convertible preferred shares, dividend rate 8.00%        
Third Security, LLC          
Related Party Transaction [Line Items]          
Expense for services     $ 14,000 $ 66,000  
Initial term of services agreement   1 year      
Shares issued as payment for services (in shares)     160,626 103,930  
Shares issued as payment for services     $ 2,041,000 $ 2,039,000  
Sublease rental income     21,000 11,000  
Third Security, LLC | Common Stock          
Related Party Transaction [Line Items]          
Expense for services   $ 800,000      
Harvest Intrexon Enterprise Fund I, LP          
Related Party Transaction [Line Items]          
Other nonoperating income       $ 603,000  
Convertible Note and Warrants | Other Assets | Fibrocell Science, Inc.          
Related Party Transaction [Line Items]          
Fair value of financial assets measured at fair value on a recurring basis     $ 494,000   $ 575,000
Third Security, LLC | Chief Executive Officer          
Related Party Transaction [Line Items]          
Ownership interest     100.00%