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Leases
 12 Months Ended
Dec. 31, 2022
Leases [Abstract]  
Leases Leases
The Company currently has operating leases for office and research laboratory space, equipment, and vehicles under agreements expiring at various dates through 2044, which include renewal options on leases which the Company is reasonably certain to exercise.
For the years ended December 31, 2022, and 2021, operating lease expense was $9.8 million and $10.0 million and variable lease expense was $1.7 million and $2.1 million, respectively. For the years ended December 31, 2022, and 2021, the Company paid $8.3 million and $8.4 million, respectively, for amounts included in the measurement of operating lease liabilities and recorded $8.9 million and $0.3 million, respectively, of right-of-use assets. For the year ended December 31, 2022, there were no tenant improvements paid through lease incentives in exchange for new operating lease liabilities. For the year ended December 31, 2021, there were $0.3 million of tenant improvements paid through lease incentives in exchange for new operating lease liabilities.
Commitments under finance leases are not significant for the year ended December 31, 2022.
Supplemental balance sheet information related to operating leases were as follows:
(in thousands, except year and discount rate amounts)December 31, 2022December 31, 2021
Operating lease ROU assets, net$29,534 $20,586 
Current portion of the operating lease liabilities$8,552 $7,409 
Non-current portion of the operating lease liabilities51,578 43,363 
Total operating lease liability$60,130 $50,772 
Weighted-average remaining lease terms (years)17.019.6
Weighted-average discount rate12.2 %13.1 %
At December 31, 2022, the future minimum operating lease payments were as follows:
(in thousands)Operating Lease
2023$9,466 
20248,624 
20258,419 
20268,571 
20278,754 
Thereafter124,979 
Total lease payments168,813 
Less lease incentives(22,299)
Less imputed interest(86,384)
Total operating lease liability$60,130