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The Business and Summary of Significant Accounting Policies - Assets and Liabilities Measured at Fair Value on a Recurring Basis (Details) - USD ($)
$ in Thousands
3 Months Ended 9 Months Ended
Sep. 30, 2019
Sep. 30, 2018
Sep. 30, 2019
Sep. 30, 2018
Dec. 31, 2018
Fair Value, Separate Account Investment [Line Items]          
Unrealized loss on equity investment     $ 0 $ 311  
Short-term investments $ 97,366   97,366   $ 105,424
Fair value, measurements, recurring          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents 16,983   16,983   75,893
Short-term investments 97,366   97,366   105,424
Total assets 114,349   114,349   181,317
Fair value, measurements, recurring | Quoted Prices in Active Markets for Identical Assets (Level 1)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents 16,983   16,983   42,208
Short-term investments 23,213   23,213   12,627
Total assets 40,196   40,196   54,835
Fair value, measurements, recurring | Significant Other Observable Inputs (Level 2)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents 0   0   33,685
Short-term investments 74,153   74,153   92,797
Total assets 74,153   74,153   126,482
Fair value, measurements, recurring | Significant Unobservable Inputs (Level 3)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents 0   0   0
Short-term investments 0   0   0
Total assets 0   0   0
Money market funds | Fair value, measurements, recurring          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents 16,983   16,983   30,726
Money market funds | Fair value, measurements, recurring | Quoted Prices in Active Markets for Identical Assets (Level 1)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents 16,983   16,983   30,726
Money market funds | Fair value, measurements, recurring | Significant Other Observable Inputs (Level 2)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents 0   0   0
Money market funds | Fair value, measurements, recurring | Significant Unobservable Inputs (Level 3)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents 0   0   0
U.S. treasury securities | Fair value, measurements, recurring          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         11,482
Short-term investments 23,213   23,213   12,589
U.S. treasury securities | Fair value, measurements, recurring | Quoted Prices in Active Markets for Identical Assets (Level 1)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         11,482
Short-term investments 23,213   23,213   12,589
U.S. treasury securities | Fair value, measurements, recurring | Significant Other Observable Inputs (Level 2)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         0
Short-term investments 0   0   0
U.S. treasury securities | Fair value, measurements, recurring | Significant Unobservable Inputs (Level 3)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         0
Short-term investments 0   0   0
Common stock of U.S. corporation | Fair value, measurements, recurring          
Fair Value, Separate Account Investment [Line Items]          
Short-term investments         38
Common stock of U.S. corporation | Fair value, measurements, recurring | Quoted Prices in Active Markets for Identical Assets (Level 1)          
Fair Value, Separate Account Investment [Line Items]          
Short-term investments         38
Common stock of U.S. corporation | Fair value, measurements, recurring | Significant Other Observable Inputs (Level 2)          
Fair Value, Separate Account Investment [Line Items]          
Short-term investments         0
Common stock of U.S. corporation | Fair value, measurements, recurring | Significant Unobservable Inputs (Level 3)          
Fair Value, Separate Account Investment [Line Items]          
Short-term investments         0
Commercial paper | Fair value, measurements, recurring          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         29,677
Short-term investments 34,853   34,853   60,114
Commercial paper | Fair value, measurements, recurring | Quoted Prices in Active Markets for Identical Assets (Level 1)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         0
Short-term investments 0   0   0
Commercial paper | Fair value, measurements, recurring | Significant Other Observable Inputs (Level 2)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         29,677
Short-term investments 34,853   34,853   60,114
Commercial paper | Fair value, measurements, recurring | Significant Unobservable Inputs (Level 3)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         0
Short-term investments 0   0   0
Corporate bonds | Fair value, measurements, recurring          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         4,008
Short-term investments 39,300   39,300   32,683
Corporate bonds | Fair value, measurements, recurring | Quoted Prices in Active Markets for Identical Assets (Level 1)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         0
Short-term investments 0   0   0
Corporate bonds | Fair value, measurements, recurring | Significant Other Observable Inputs (Level 2)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         4,008
Short-term investments 39,300   39,300   32,683
Corporate bonds | Fair value, measurements, recurring | Significant Unobservable Inputs (Level 3)          
Fair Value, Separate Account Investment [Line Items]          
Cash equivalents         0
Short-term investments 0   0   $ 0
ContraVir investment | Common stock of U.S. corporation          
Fair Value, Separate Account Investment [Line Items]          
Unrealized loss on equity investment $ 3 $ 99 $ 33 $ 311