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Stock-based Compensation (Tables)
3 Months Ended
Mar. 31, 2016
Disclosure of Compensation Related Costs, Share-based Payments [Abstract]  
Schedule of Share-Based Compensation Expense
Employee share-based compensation expense recognized related to stock options and the ESPP is as follows (in thousands):
 
 
 
Three Months Ended March 31,
 
 
2016
 
2015
Research and development expense
 
$
1,579

 
$
1,025

General and administrative expense
 
2,336

 
1,377

          Total share-based compensation expense
 
$
3,915

 
$
2,402