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STATEMENTS OF OPERATIONS - USD ($)
$ in Thousands
3 Months Ended
Mar. 31, 2017
Mar. 31, 2016
Income Statement [Abstract]    
Collaboration revenue $ 1,597 $ 1,969
Operating expenses:    
Research and development 11,967 10,199
General and administrative 3,595 3,810
Total operating expenses 15,562 14,009
Loss from operations (13,965) (12,040)
Other income (expense), net 38 21
Change in fair value of derivative liabilities (1,560)  
Net loss (15,487) (12,019)
Preferred stock dividends (4,171)  
Net loss applicable to common stockholders $ (19,658) $ (12,019)
Basic and diluted net loss per share $ (0.15) $ (0.09)
Weighted average common shares outstanding used to compute basic and diluted net loss per share 130,696,400 130,157,927