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Condensed Consolidated Statements of Operations (Unaudited) - USD ($)
shares in Thousands, $ in Thousands
3 Months Ended 9 Months Ended
Sep. 30, 2016
Sep. 30, 2015
Sep. 30, 2016
Sep. 30, 2015
Revenues:        
Collaboration revenue   $ 829 $ 2,578 $ 2,230
Grant revenue $ 162 359 289 608
Service and license revenue     884 527
Total revenues 162 1,188 3,751 3,365
Operating expenses:        
Research and development 23,234 24,105 66,051 66,011
General and administrative 11,766 5,524 29,086 15,481
Total operating expenses 35,000 29,629 95,137 81,492
Loss from operations (34,838) (28,441) (91,386) (78,127)
Other income (expense):        
Interest income 170 33 615 78
Interest expense   (62)   (572)
Other (expense) income, net (26) 17 68 360
Loss on extinguishment of debt   (1,671)   (1,671)
Net loss $ (34,694) $ (30,124) $ (90,703) $ (79,932)
Basic and diluted net loss per share $ (0.90) $ (0.82) $ (2.36) $ (2.43)
Weighted average shares used to compute basic and diluted net loss per share 38,512 36,532 38,493 32,880