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Property and Equipment, net (Details) - USD ($)
$ in Thousands
12 Months Ended
Dec. 31, 2016
Dec. 31, 2015
Dec. 31, 2014
Property, Plant and Equipment [Line Items]      
Less - accumulated depreciation $ (12,457) $ (14,630)  
Property and equipment, net 3,320 2,616  
Depreciation 601 568 $ 681
Laboratory equipment      
Property, Plant and Equipment [Line Items]      
Total property and equipment 9,019 9,963  
Office and computer equipment      
Property, Plant and Equipment [Line Items]      
Total property and equipment 2,519 2,968  
Furniture, fixtures and fittings      
Property, Plant and Equipment [Line Items]      
Total property and equipment $ 4,239 $ 4,315