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OTHER BALANCE SHEET DETAILS
 12 Months Ended
Dec. 31, 2021
Balance Sheet Related Disclosures [Abstract]  
OTHER BALANCE SHEET DETAILS OTHER BALANCE SHEET DETAILS
Property and Equipment, Net
Property and equipment, net consist of the following (in thousands):
December 31,
20212020
Laboratory equipment$31,988 $24,737 
Leasehold improvements21,970 15,953 
Furniture and fixtures9,080 4,870 
Manufacturing equipment8,781 1,089 
Construction in progress4,729 12,091 
76,548 58,740 
Less: accumulated depreciation and amortization(25,025)(17,416)
Property and equipment, net$51,523 $41,324 
Depreciation and amortization expense was $9.4 million in 2021, $5.7 million in 2020 and $3.9 million in 2019.
Intangible Assets
The changes in intangible assets were as follows (in thousands):
December 31,
20212020
Balance at beginning of year$58,128 $53,156 
Foreign currency translation adjustment(4,368)4,972 
Balance at end of year$53,760 $58,128 
Goodwill
The changes in goodwill were as follows (in thousands):
December 31,
20212020
Balance at beginning of year$42,798 $39,273 
Foreign currency translation adjustment(3,096)3,525 
Balance at end of year$39,702 $42,798 
Other Accrued Liabilities
Other accrued liabilities consist of the following (in thousands):
December 31,
20212020
Accrued research and development expenses$4,878 $4,257 
Operating lease liabilities – current
4,026 3,690 
Accrued professional fees869 1,532 
Customer advance— 5,000 
Other1,804 4,133 
Total other accrued liabilities$11,577 $18,612